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Chapter - III
The term ecosystem was coined by A.G. TansIey (1935) who stated the following words,
“………………… the more fundamental conception is………….. the whole system (in the sense of
physics), including not only the organism complex but also the whole complex of physical factors
forming what we call the environment of the biome – the habitat factors in the widest sense.
Though the organisms may claim out primary interest, when we are trying to think
fundamentally we cannot separate them from their special environment which they form one
physical system. It is the systems so formed which…………………..are basic units of nature on
the face of the earth……………….. The ecosystems as we may call them are of most various kinds
and sizes”. Thus, the ecosystem can be of any size.
It has been observed that the green plants and some autotrophic bacteria are able to prepare their
own food by utilizing solar radiant energy (light). They are capable of binding up the simple
molecules of carbon-dioxide, water and other elements, such as, nitrogen, phosphorous,
potassium, magnesium etc. by the help of light.
These substances are confined to the physical surroundings. Some animals (herbivores) eat
plants directly and others (carnivores) eat such animals, which feed on the plants. Another group
of organisms depends on the dead plants and animals to obtain their food from the decomposed
tissues.
The food is digested and assimilated to synthesize different organic compounds from which the
energy is derived for the life activities. The mechanism involves two distinct processes, firstly,
the energy used by the plants passes through various organisms and finally it is lost as heat. The
process has been referred as the ―flow of energy‖.
Secondly, along with energy, the substances also pass through the organisms but remain
available for use, again and again. This process is known as biogeochemical cycling. These two
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processes bind the organisms to their environment and the complex network is termed as the
ecosystem.
An ecosystem in a laboratory may be as small as an aquarium. It may range from the size of a
lake or forest to the size of the earth. Biosphere (W. Vernadsky 1929) or the Ecosphere (L.C.
Cole, 1958) is the usual term which applies to the ecosystem that covers the whole of the earth.
In fact, none of the ecosystems in independent, rather all are interdependent in some way or
other.
All ecosystems consist of two major components—biotic and abiotic. The biotic component is
comprised the living organisms, whereas the abiotic component includes the physical (non-
living) environment. But, both of these components interact very closely to exhibit a definite
structural organization. Sometimes, it is very difficult to separate the biotic components from the
abiotic components.
Biotic Component:
When we consider the biotic components, the organisms are divided into two categories, the
autotrophs and the heterotrophs. The autotrophs can produce their own food. They are the green
plants with chlorophyll and certain types of bacteria—chemosynthetic and photosynthetic. Since
these organisms produce food for all other organisms, they are also known as ‗Producer‘. The
heterotrophs depend directly or indirectly on the autotrophs for their food. This type of
organisms is further divided into two groups, such as, Photographs and Osmotrophs.
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The photographs take food from outside and digest it inside their bodies. They are called
consumers. All animals—herbivores (plant eating), carnivores (animal eating) or omnivores
(eating all kinds of food) fall in this group. The osmotrophs are those organisms who secrete
digestive enzymes to break down the food into simpler substances and then absorb the digested
food.
This group embraces the parasitic and saprophytic bacteria as well as the fungi. They may also
be called Decomposers because their role has been well documented in the decomposition of the
dead organic matter. But the most interesting point is that all of these parasites are not
decomposers, rather some of them are consumers (insects and such small animals) who help in
the decomposition by breaking down the dead organic matter into small bits. However, the
heterotrophs can also be divided into two broad groups as the bio phages (feeding on living
organisms) and the sarcophagus (feeding on dead organisms) Fig. 10.3.
The above mentioned divisions of organisms are principally based on the nature of food
which in turn gives rise to the trophic structure of the ecosystem. It signifies a step-wise
systematic relationships among the living organisms for food, which forms a chain,
known as the food chain.
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Such food chains can be traced back to the Producers and the position of the organism in
the food chain is indicated by the trophic levels. This trophic level is defined as the
number of links by which it is separated from the producer; the producers always belong
to the first trophic level or the base.
A single species can operate more than one trophic level in the ecosystem which means
that this species get its food from more than one source. Again, the same species may be
eaten by several other species of a higher trophic level; thus, one can find out several
food chains linked together in an intersecting manner to develop a network, known as
food web. Such a network provides a clear- cut idea about the functioning of the
ecosystem.
Abiotic Component:
Abiotic component of ecosystem refers to the physical environment and its several
interacting variables which can be divided into four folds:
(i) Lithosphere which means the solid mineral matter on the earth and the land form as
well;
(ii) Hydrosphere, i.e. the water in oceans, lakes, river, ice-caps, etc.;
The position and movement of the earth with its gravitational force are additional
components of the environment Fig. 10.4. However, these components create
invariability of magnitude and duration of other environmental factors.
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The energy interacts with rocks, water and gases to produce a complex environment with
a large number of identifiable variables such as heat, light, rain, wind, snow, fog, dust,
storm, fire, etc. Thus, by the interaction of variables, the environment is created and
maintained as a unit where any single component cannot be removed or altered without
disturbing the other components.
Functionally the ecosystem allows the flow of energy and cycling of materials which
ensures the stability of the system and continuity of life. The energy needed for all life
processes come from solar radiation Fig. 10.5.
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During photosynthesis, green plants convert light energy to chemical (potential) energy and
make it available to other organisms as food. Thus, a continuous flow of energy from sun
through organisms maintains the life on earth. Laws of Thermodynamics govern on the transfer
and transformation of energy.
It says, the energy can never be destroyed, but it is transformed into different forms. Therefore,
the part of solar radiant energy that is not used in photosynthesis is used in heating of air, water
and soil. So some change does occur in cyclic order of nature.
Ultimately the energy is reflected back to outer space as heat. In fact, a small fraction of
available light energy is utilized during photosynthesis and a very little part is stored in animal
tissues; the bulk is wasted as heat.
The next point is the ratio between the production and assimilation of energy. The small
organisms utilize a large part of the assimilated energy for growth while larger organisms
consume a larger part of the assimilated energy for maintenance of the organism (respiration).
However, all these mechanisms-the transformation of energy, the food chain, the assimilation
etc., are expressed as ecological efficiency.
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It is therefore clear that the two processes, namely the flow of energy and the cycling of
materials are equally important for the functioning of ecosystem. These two processes are
inseparable and run concurrently. As a consequence of interactions, a variety of organic
substances are produced. This production is called biological production, which is
essentially different from the chemical and industrial production. Biologists are interested
in this production as it a part of a perpetual process.
In dealing with ecosystem, the concept of community is very important which can be
defined as an assemblage of a number of organisms including man. The organisms
include several species, which occupy the same habitat.
Even the micro-organisms are the part of a community. Here, ‗community‘ is essentially
treated as a biotic component of the ecosystem. A population may also be defined
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Naturally the members live in a common area and like to use the same resources of the
particular ecosystem. In this manner, by the way of interactions between the organism
and its environment, each population develops a style of life, which is designated as the
niche or ecological niche of the population.
The niche denotes a specific portion of the habitat occupied by each species. In 1917, J.
Grinnell first brought this word. In 1927, C. Elton defined niche as an animal‘s place in
the biotic environ-population in the community. In a simple way, it can be said that the
habitat is the address of the species and the niche is its profession.
So, several populations with different functions may occupy the same habitat. The
ecological niche is referred to the totality of biotic and abiotic factors to which a given
population is uniquely adapted. The concept of niche was analyzed by R.H. Whittakar in
1970 as ―functional system of interacting niche differentiated species populations that
tend to complement one-another rather than directly competing, in their utilization of the
community‘s space, time, resources and possible kinds of interactions‖.
The growth as well as life forms of climax populations suggests the result of interactions
between the organisms and their environment. Though the climax species is a subject of
controversial discussion, still each climatic zone represents some particular species as
dominants.
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As a matter of fact, biome has been emphasized on the recognition and classification of
natural biotic communities, which can be correlated, with the abiotic environment of that
area. The other important concept, apart from Biome and Habitat, is the culture. Culture
indicates a man-made environment is contrast to natural environment.
Man tends to develop an environment around him by invention as well as learning. The
ecological niche appears by the interplay of the biome, habitat and culture. It keeps a
harmonic balance between the biotic and abiotic components in the biosphere of
ecosystem. (Fig. 10.7.)
As man is the most superior animal on the earth, he is able to manipulate his own
environment. Sometimes he exploits the nature so severely that many life forms,
beneficial to his survival are destroyed quite unknowingly. This disturbs the balance of
the natural ecosystem and man faces odd consequences. For example, modem techniques
of Agriculture and industrial development have brought adverse impact on ecosystem.
Ecological niche
An ecological niche is the role and position a species has in its environment; how it
meets its needs for food and shelter, how it survives, and how it reproduces. A
species' nicheincludes all of its interactions with the biotic and abiotic factors of its
environment.
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Every living thing on Earth has a role to play in its environment. In fact, you are
filling a niche right now as you read this lesson. Whether you are a student, have
a full time job, or are a mother or father, these are parts of the niche you fill.
Your niche also includes where and how you obtain food and all of the things
you do in order to survive.
If you closely look at a typical habitat in the environment, you will see many
organisms living and working together, fulfilling their ecological niches. For
example, imagine you are walking through the forest where there are leaves
scattered on the ground and an old rotting log sitting on the forest floor. If you
look closely, you could probably find earthworms just under the soil feeding on
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decaying organic matter. There could also be centipedes eating small beetles
and other organisms as well as a colony of ants that work and feed on dead
insects. You may even find a couple of millipedes strolling around feeding on
decaying leaves.
In this small section of the vast forest, all of these organisms are filling an
individual ecological niche. To some degree, their niches may overlap, but if you
look into all aspects of their lives, including where they live, how they survive,
and how they reproduce, you will see that they are each truly individual niches.
You could think of each ecological niche as parts of a puzzle that go together to
make the environment successful.
The study of the interactions between living things and their environments is
known as ecology. In ecology, everything is connected to everything else and
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The primary importance of the hydrosphere is that it contains water, which sustains
a variety of life forms and plays an important role in regulating the atmosphere and
surrounding ecosystems. The hydrosphere includes all water located on the surface of
the Earth. It contains freshwater, saltwater and frozen water as well, including
groundwater and water in the lower levels of the atmosphere.
The water in the hydrosphere varies in texture and consistency, but shares the important
function of sustaining human, plant, animal and bacterial life on Earth. Living organisms
contain approximately 75 percent water. Cells within living beings rely on water to carry
out important life functions. Water also allows cells to carry out critical chemical
reactions, which otherwise would not happen, and therefore cause life to cease.
In addition to existing within organisms, water exists in habitats where plants and animals
live. Water helps to regulate climates and atmospheric conditions and facilitates human
activities, such as irrigation. The hydrosphere contains bodies of water around the world.
Most of its composition derives from oceans, which have water trapped in layers of
sedimentary rock. Oceans, lakes, rivers and ice include the main geographical features in
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the hydrosphere, and exist in places around the world. Of these features, ice makes up the
smallest portion of the hydrosphere, contributing only one percent to the total.
The hydrosphere provides an important place for the human beings and animals to live.
There are also nutrients like the nitrate, nitrite, ammonium ions and other ions are
dissolved in water. These substances is really needed for life to exist in water. The
hydrosphere also helps the earth to regulate its temperature for the reason water takes a
long time to heat up and a long time to cool down making the temperature of the earth
stay within a range that is acceptable for life to exist. Humans and animals use water in a
number of ways. Humans and animals uses water for drinking, which is essential to
maintain life, and water is used for other purposes like washing, cleaning, and generating
electricity through hydropower. If the hydrosphere or the so called water sphere is
depleted or will totally vanish, there will be no more supply of water that is essential to
maintain life and there will be no more human beings or even animals that will live in the
planet earth. If the hydrosphere will vanish, the interaction between the hydrosphere and
othere spheres of the earth like the litosphere, atmosphere and biosphere will be out of
balance and will create disturbance or malfunction of the litosphere, atmosphere, and
biosphere.
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An ecosystem consists of the biological community that occurs in some locale, and the
physical and chemical factors that make up its non-living or abiotic environment. There
are many examples of ecosystems -- a pond, a forest, an estuary, a grassland. The
boundaries are not fixed in any objective way, although sometimes they seem obvious, as
with the shoreline of a small pond. Usually the boundaries of an ecosystem are chosen for
practical reasons having to do with the goals of the particular study.
The study of ecosystems mainly consists of the study of certain processes that link the
living, or biotic, components to the non-living, or abiotic, components. Energy
transformations andbiogeochemical cycling are the main processes that comprise the
field of ecosystem ecology. As we learned earlier, ecology generally is defined as the
interactions of organisms with one another and with the environment in which they occur.
We can study ecology at the level of the individual, the population, the community, and
the ecosystem.
In ecosystem ecology we put all of this together and, insofar as we can, we try to
understand how the system operates as a whole. This means that, rather than worrying
mainly about particular species, we try to focus on major functional aspects of the
system. These functional aspects include such things as the amount of energy that is
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produced by photosynthesis, how energy or materials flow along the many steps in a food
chain, or what controls the rate of decomposition of materials or the rate at which
nutrients are recycled in the system.
Components of an Ecosystem
You are already familiar with the parts of an ecosystem. You have learned about climate
and soils from past lectures. From this course and from general knowledge, you have a
basic understanding of the diversity of plants and animals, and how plants and animals
and microbes obtain water, nutrients, and food. We can clarify the parts of an ecosystem
by listing them under the headings "abiotic" and "biotic".
By and large, this set of environmental factors is important almost everywhere, in all
ecosystems.
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depend very much on who the actual players (species) happen to be, only on what
function they perform in the ecosystem.
Processes of Ecosystems
This figure with the plants, zebra, lion, and so forth illustrates the two main ideas about
how ecosystems function: ecosystems have energy flows and ecosystems cycle materials.
These two processes are linked, but they are not quite the same (see Figure 1).
Energy enters the biological system as light energy, or photons, is transformed into
chemical energy in organic molecules by cellular processes including photosynthesis and
respiration, and ultimately is converted to heat energy. This energy is dissipated, meaning
it is lost to the system as heat; once it is lost it cannot be recycled. Without the continued
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input of solar energy, biological systems would quickly shut down. Thus the earth is
an open system with respect to energy.
During decomposition these materials are not destroyed or lost, so the earth is a closed
system with respect to elements (with the exception of a meteorite entering the system
now and then). The elements are cycled endlessly between their biotic and abiotic states
within ecosystems. Those elements whose supply tends to limit biological activity are
callednutrients.
The transformations of energy in an ecosystem begin first with the input of energy from
the sun. Energy from the sun is captured by the process of photosynthesis. Carbon
dioxide is combined with hydrogen (derived from the splitting of water molecules) to
produce carbohydrates (CHO). Energy is stored in the high energy bonds of adenosine
triphosphate, or ATP (see lecture on photosynthesis).
The prophet Isaah said "all flesh is grass", earning him the title of first ecologist, because
virtually all energy available to organisms originates in plants. Because it is the first step
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For reasons that we will explore more fully in subsequent lectures, energy transfer
through the food chain is inefficient. This means that less energy is available at the
herbivore level than at the primary producer level, less yet at the carnivore level, and so
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on. The result is a pyramid of energy, with important implications for understanding the
quantity of life that can be supported.
Usually when we think of food chains we visualize green plants, herbivores, and so on.
These are referred to asgrazer food chains, because living plants are directly consumed.
In many circumstances the principal energy input is not green plants but dead organic
matter. These are called detritus food chains. Examples include the forest floor or a
woodland stream in a forested area, a salt marsh, and most obviously, the ocean floor in
very deep areas where all sunlight is extinguished 1000's of meters above. In subsequent
lectures we shall return to these important issues concerning energy flow.
Finally, although we have been talking about food chains, in reality the organization of
biological systems is much more complicated than can be represented by a simple
"chain". There are many food links and chains in an ecosystem, and we refer to all of
these linkages as a food web. Food webs can be very complicated, where it appears
that "everything is connected to everything else", and it is important to understand what
are the most important linkages in any particular food web.
Biogeochemistry
How can we study which of these linkages in a food web are most important? One
obvious way is to study the flow of energy or the cycling of elements. For example, the
cycling of elements is controlled in part by organisms, which store or transform elements,
and in part by the chemistry and geology of the natural world. The
term Biogeochemistry is defined as the study of how living systems influence, and are
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controlled by, the geology and chemistry of the earth. Thus biogeochemistry
encompasses many aspects of the abiotic and biotic world that we live in.
There are several main principles and tools that biogeochemists use to study earth
systems. Most of the major environmental problems that we face in our world toady can
be analyzed using biogeochemical principles and tools. These problems include global
warming, acid rain, environmental pollution, and increasing greenhouse gases. The
principles and tools that we use can be broken down into 3 major components: element
ratios, mass balance, and element cycling.
1. Element ratios
C : N : P : Fe = 106 : 16 : 1 : 0.01
Once we know these ratios, we can compare them to the ratios that we measure in a
sample of algae to determine if the algae are lacking in one of these limiting
nutrients.
2. Mass Balance
Another important tool that biogeochemists use is a simple mass balance equation to
describe the state of a system. The system could be a snake, a tree, a lake, or the entire
globe. Using a mass balance approach we can determine whether the system is changing
and how fast it is changing. The equation is:
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In this equation the net change in the system from one time period to another is
determined by what the inputs are, what the outputs are, and what the internal change in
the system was. The example given in class is of the acidification of a lake, considering
the inputs and outputs and internal change of acid in the lake.
3. Element Cycling
Element cycling describes where and how fast elements move in a system. There are two
general classes of systems that we can analyze, as mentioned above: closed and open
systems.
A closed system refers to a system where the inputs and outputs are negligible compared
to the internal changes. Examples of such systems would include a bottle, or our entire
globe. There are two ways we can describe the cycling of materials within this closed
system, either by looking at the rate of movement or at the pathways of movement.
In an open system there are inputs and outputs as well as the internal cycling. Thus we
can describe the rates of movement and the pathways, just as we did for the closed
system, but we can also define a new concept called the residence time. The residence
time indicates how long on average an element remains within the system before leaving
the system.
1. Rate
2. Pathways
3. Residence time, Rt
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Now that we have learned something about how ecosystems are put together and how
materials and energy flow through ecosystems, we can better address the question of
"what controls ecosystem function"? There are two dominant theories of the control of
ecosystems. The first, called bottom-up control, states that it is the nutrient supply to the
primary producers that ultimately controls how ecosystems function. If the nutrient
supply is increased, the resulting increase in production of autotrophs is propagated
through the food web and all of the other trophic levels will respond to the increased
availability of food (energy and materials will cycle faster).
The second theory, called top-down control, states that predation and grazing by higher
trophic levels on lower trophic levels ultimately controls ecosystem function. For
example, if you have an increase in predators, that increase will result in fewer grazers,
and that decrease in grazers will result in turn in more primary producers because fewer
of them are being eaten by the grazers. Thus the control of population numbers and
overall productivity "cascades" from the top levels of the food chain down to the bottom
trophic levels.
So, which theory is correct? Well, as is often the case when there is a clear dichotomy to
choose from, the answer lies somewhere in the middle. There is evidence from many
ecosystem studies that BOTH controls are operating to some degree, but that NEITHER
control is complete. For example, the "top-down" effect is often very strong at trophic
levels near to the top predators, but the control weakens as you move further down the
food chain. Similarly, the "bottom-up" effect of adding nutrients usually stimulates
primary production, but the stimulation of secondary production further up the food chain
is less strong or is absent.
Thus we find that both of these controls are operating in any system at any time, and we
must understand the relative importance of each control in order to help us to predict how
an ecosystem will behave or change under different circumstances, such as in the face of
a changing climate.
There are many different ecosystems: rain forests and tundra, coral reefs and ponds,
grasslands and deserts. Climate differences from place to place largely determine the
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The word "biome" is used to describe a major vegetation type such as tropical rain forest,
grassland, tundra, etc., extending over a large geographic area (Figure 3). It is never used
for aquatic systems, such as ponds or coral reefs. It always refers to a vegetation category
that is dominant over a very large geographic scale, and so is somewhat broader than an
ecosystem.
We can draw upon previous lectures to remember that temperature and rainfall patterns
for a region are distinctive. Every place on earth gets the same total number of hours of
sunlight each year, but not the same amount of heat. The sun's rays strike low latitudes
directly but high latitudes obliquely. This uneven distribution of heat sets up not just
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temperature differences, but global wind and ocean currents that in turn have a great deal
to do with where rainfall occurs. Add in the cooling effects of elevation and the effects of
land masses on temperature and rainfall, and we get a complicated global pattern of
climate.
A schematic view of the earth shows that, complicated though climate may be, many
aspects are predictable (Figure 4). High solar energy striking near the equator ensures
nearly constant high temperatures and high rates of evaporation and plant transpiration.
Warm air rises, cools, and sheds its moisture, creating just the conditions for a tropical
rain forest. Contrast the stable temperature but varying rainfall of a site in Panama with
the relatively constant precipitation but seasonally changing temperature of a site in New
York State. Every location has a rainfall- temperature graph that is typical of a broader
region.
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We can draw upon plant physiology to know that certain plants are distinctive of certain
climates, creating the vegetation appearance that we call biomes. Note how well the
distribution of biomes plots on the distribution of climates (Figure 5). Note also that some
climates are impossible, at least on our planet. High precipitation is not possible at low
temperatures -- there is not enough solar energy to power the water cycle, and most water
is frozen and thus biologically unavailable throughout the year. The high tundra is as
much a desert as is the Sahara.
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Types of Ecosystems
There are many types of ecosystems on earth. There are three major classes of ecosystems:
Pond Ecosystem: In a pond, frogs, fishes, other tiny insects, several bacteria, fungi, algae
with a variety of plants may be present. Some plants may float on water surface and some
completely submerged in the water. Some water birds may be swimming on water
surface. Fishes and frogs eat small insects. Small animals (insects etc) eat small plants.
The water plants need CO2 to prepare their food, this CO2 comes from the pond animals.
During food preparation process plants give out oxygen, this oxygen is utilized by the
pond animals. This form a pond ecosystem.
Forest Ecosystem: This includes inorganic and organic substances present in the
atmosphere and soil. Tree and shrubs are also present in forest. There are small animals
feeding on tree leaves these are ants, beetles, flies, spiders and grasshoppers. Different
kinds of big animals and birds, snakes, lizards are also present in the forest. They all are
inter-related with each other for food, energy, oxygen and other components which are
essential for their survival and life processes.
Sea Ecosystem: In sea, we find so many types of plants and animals. Some plants and
animals are very minute and some are very big. Green plants prepare food, these plants
are eaten by the small animals and small animals are eaten by the huge animals. Their
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main need is food, oxygen, carbon-dioxide and other life process components. They are
inter-related with each other for their essential components for life process.
Every ecosystem has several interrelated mechanisms that affect human life. These are
The Water Cycle, The Carbon Cycle, The Oxygen Cycle, The Nitrogen Cycle and The
Energy Cycle. Every ecosystem is controlled by these cycles and each ecosystem‘s
abiotic and biotic features are distinct from each other. All functions of the ecosystem
are in some way related to the growth and re-generation of its plant and animal species.
The water cycle depends on the rainfall, which is necessary for plants and animals to
live.
Carbon is found in components of ecosystem and it is building block of both plant and
animal tissues. In the atmosphere carbon occurs as CO2 .Plants help in regulating and
monitoring the percentage of oxygen and carbon dioxide in the Earth‘s atmosphere. Both
plants and animals release CO2 during respiration. They also return fixed carbon to the
soil in the waste they excrete. When plants and animals die, they return their carbon to
the soil. These processes complete the carbon cycle and the oxygen cycle.
In the nitrogen cycle, plants absorb nitrogen in the form of nutrients and use for their
growth. Nutrients are re-cycle back from animals to plants.
The energy cycle re-cycle nutrients into the soil on which plant life grows.
Our own lives are closely linked to the proper functioning of these cycles of life.
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These cycles are a part of global life processes and have specific features in each of the
ecosystems. They are also linked to adjacent ecosystems as well as plants and animals
communities in the region. Together, the cycles are responsible for maintaining life on
Earth. If human activities go on altering them beyond the limits that nature can sustain,
they will eventually break down and lead to a degraded Earth on which man will not be
able to survive.
Summary
Energy is continually input into an ecosystem in the form of light energy, and some
energy is lost with each transfer to a higher trophic level. Nutrients, on the other hand, are
recycled within an ecosystem, and their supply normally limits biological activity. So,
"energy flows, elements cycle".
Energy is moved through an ecosystem via a food web, which is made up of interlocking
food chains. Energy is first captured by photosynthesis (primary production). The amount
of primary production determines the amount of energy available to higher trophic levels.
A biome is a major vegetation type extending over a large area. Biome distributions are
determined largely by temperature and precipitation patterns on the Earth's surface.
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It is not easy to compare terrestrial and aquatic systems because there is such a large
variety of these environments. It is possible to recognize in the terrestrial part of the
biosphere a small number of units with distinctive vegetation and climate, each with a
complex of communities of large extent. These units are known as biomes and six major
biomes are usually recognized, namely the:
Tundra,
Taiga ( coniferous forests ),
Deciduous Forests,
Grasslands,
Tropical Rain Forests,
Deserts.
In this section we will focus on the similarities and differences between terrestrial and aquatic
ecosystems.
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because aquatic environments are so rich in nutrients they support more live than
equivalent terrestrial ecosystems. The small drifting photosynthetic organisms of
the oceans, referred to collectively as phytoplankton are regarded as the major
photosynthesizers, or primary producers, of the earth,
aquatic environments are much more stable than terrestrial environments, with
smaller fluctuations in temperature and other variables,
aquatic organisms are seldom exposed to desiccation while terrestrial organisms
are often exposed to desiccation and are usually relatively resistant to drying out,
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oxygen (because there is very much less present) is sometimes a limiting factor an
aquatic habitats but this is seldom the case in terrestrial habitats,
light can be a limiting factor in some aquatic habitats, but in most terrestrial
environments there is hardly ever a a shortage of light,
terrestrial animals are influenced far more by gravity, while water supports aquatic
organisms.
The low rate of energy transfer between trophic levels makes decomposers
generally more important than producers in terms of energy flow. Decomposers
process large amounts of organic material and return nutrients to the ecosystem in
inorganic form, which are then taken up again by primary producers. Energy is
not recycled during decomposition, but rather is released, mostly as heat
(this is what makes compost piles and fresh garden mulch warm). Figure 6
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shows the flow of energy (dark arrows) and nutrients (light arrows) through
ecosystems.
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In the oceans, light and nutrients are important controlling factors for
productivity. As noted in Unit 3, "Oceans," light penetrates only into the
uppermost level of the oceans, so photosynthesis occurs in surface and near-
surface waters. Marine primary productivity is high near coastlines and other
areas where upwelling brings nutrients to the surface, promoting plankton
blooms. Runoff from land is also a source of nutrients in estuaries and along
the continental shelves. Among aquatic ecosystems, algal beds and coral
reefs have the highest net primary production, while the lowest rates occur in
the open due to a lack of nutrients in the illuminated surface layers (Fig. 8).
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How many trophic levels can an ecosystem support? The answer depends on
several factors, including the amount of energy entering the ecosystem,
energy loss between trophic levels, and the form, structure, and physiology
of organisms at each level. At higher trophic levels, predators generally are
physically larger and are able to utilize a fraction of the energy that was
produced at the level beneath them, so they have to forage over increasingly
large areas to meet their caloric needs.
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Growth rates may also be a factor. Phytoplankton are extremely small but
grow very rapidly, so they support large populations of herbivores even
though there may be fewer algae than herbivores at any given moment. In
contrast, land plants may take years to reach maturity, so an average carbon
atom spends a longer residence time at the primary producer level on land
than it does in a marine ecosystem. In addition, locomotion costs are
generally higher for terrestrial organisms compared to those in aquatic
environments.
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The insecticide DDT, which was widely used in the United States from the
1940s through the 1960s, is a famous case of bioaccumulation. DDT built up
in eagles and other raptors to levels high enough to affect their reproduction,
causing the birds to lay thin-shelled eggs that broke in their nests.
Fortunately, populations have rebounded over several decades since the
pesticide was banned in the United States. However, problems persist in
some developing countries where toxic bioaccumulating pesticides are still
used.
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Along with energy, water and several other chemical elements cycle through
ecosystems and influence the rates at which organisms grow and reproduce. About
10 major nutrients and six trace nutrients are essential to all animals and plants,
while others play important roles for selected species (footnote 3). The most
important biogeochemical cycles affecting ecosystem health are the water, carbon,
nitrogen, and phosphorus cycles.
As noted earlier, most of the Earth's area that is covered by water is ocean. In terms
of volume, the oceans dominate further still: nearly all of Earth's water inventory is
contained in the oceans (about 97 percent) or in ice caps and glaciers (about 2
percent), with the rest divided among groundwater, lakes, rivers, streams, soils, and
the atmosphere. In addition, water moves very quickly through land ecosystems.
These two factors mean that water's residence time in land ecosystems is generally
short, on average one or two months as soil moisture, weeks or months in shallow
groundwater, or up to six months as snow cover.
But land ecosystems process a lot of water: almost two-thirds of the water that falls
on land as precipitation annually is transpired back into the atmosphere by plants,
with the rest flowing into rivers and then to the oceans. Because cycling of water is
central to the functioning of land ecosystems, changes that affect the hydrologic
cycle are likely to have significant impacts on land ecosystems. (Global water
cycling is discussed in more detail in Unit 8, "Water Resources.")
Both land and ocean ecosystems are important sinks for carbon, which is taken up
by plants and algae during photosynthesis and fixed as plant tissue. Table 2
compares the quantities of carbon stored in Earth's major reservoirs.
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Carbon cycles relatively quickly through land and surface-ocean ecosystems, but
may remain locked up in the deep oceans or in sediments for thousands of years.
The average residence time that a molecule of carbon spends in a terrestrial
ecosystem is about 17.5 years, although this varies widely depending on the type of
ecosystem: carbon can be held in old-growth forests for hundreds of years, but its
residence time in heavily grazed ecosystems where plants and soils are repeatedly
turned over may be as short as a few months.
Currently, it is not clear what mechanisms are responsible for high absorption of
carbon by land ecosystems. One hypothesis suggests that higher atmospheric CO2
concentrations have increased the rates at which plants carry out photosynthesis
(so-called CO2 fertilization), but this idea is controversial. Controlled experiments
have shown that elevated CO2 levels are only likely to produce short-term
increases in plant growth, because plants soon exhaust available supplies of
important nutrients such as nitrogen and phosphorus that also are essential for
growth.
Nitrogen and phosphorus are two of the most essential mineral nutrients for all
types of ecosystems and often limit growth if they are not available in sufficient
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quantities. (This is why the basic ingredients in plant fertilizer are nitrogen,
phosphorus, and potassium, commonly abbreviated as NPK.) A slightly expanded
version of the basic equation for photosynthesis shows how plants use energy from
the sun to turn nutrients and carbon into organic compounds:
Because atmospheric nitrogen (N2) is inert and cannot be used directly by most
organisms, microorganisms that convert it into usable forms of nitrogen play
central roles in the nitrogen cycle. So-called nitrogen-fixing bacteria take inert
nitrogen (N2) from the atmosphere and convert it to ammonia (NH4) nitrate (NO3)
and another nitrogen compounds, which in turn are taken up by plants. Some of
these bacteria live in mutualistic relationships on the roots of plants, mainly
legumes (peas and beans), and provide nitrogen directly to the plants; farmers often
plant these crops to restore nitrogen to depleted soils. At the back end of the cycle,
decomposers break down dead organisms and wastes, converting organic materials
to inorganic nutrients. Other bacteria carry out denitrification, breaking down
nitrate to gain oxygen and returning gaseous nitrogen to the atmosphere (Fig. 9).
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Phosphorus, the other major plant nutrient, does not have a gaseous phase like
carbon or nitrogen. As a result it cycles more slowly through the biosphere. Most
phosphorus in soils occurs in forms that organisms cannot use directly, such as
calcium and iron phosphate. Usable forms (mainly orthophosphate, or PO 4) are
produced mainly by decomposition of organic material, with a small contribution
from weathering of rocks (Fig. 11).
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The amount of phosphate available to plants depends on soil pH. At low pH,
phosphorus binds tightly to clay particles and is transformed into relatively
insoluble forms containing iron and aluminum. At high pH, it is lost to other
inaccessible forms containing calcium. As a result, the highest concentrations of
available phosphate occur at soil pH values between 6 and 7. Thus soil pH is an
important factor affecting soil fertility.
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Drastic changes at the top of the food web can trigger trophic cascades, or domino
effects that are felt through many lower trophic levels. The likelihood of a trophic
cascade depends on the number of trophic levels in the ecosystem and the extent to
which predators reduce the abundance of a trophic level to below their resource-
limited carrying capacity. Some species are so important to an entire ecosystem
that they are referred to as keystone species, connoting that they occupy an
ecological niche that influences many other species. Removing or seriously
impacting a keystone species produces major impacts throughout the ecosystem.
"Species, like riparian songbirds, insects, and in particular, rodents, have come
back into these preferred habitat types, and other species are starting to respond,"
says biologist Robert Crabtree of the Yellowstone Ecological Research Center.
"For example, fox and coyotes are moving into these areas because there's more
prey for them. There's been an erupting trophic cascade in some of these lush
riparian habitat sites."
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Ecological Niches
Each species in an ecosystem occupies a niche, which comprises the sum total of
its relationships with the biotic and abiotic elements of its environment—more
simply, what it needs to survive. In a 1957 address, zoologist George Evelyn
Hutchinson framed the view that most ecologists use today when he defined the
niche as the intersection of all of the ranges of tolerance under which an organism
can live (footnote 5). This approach makes ecological niches easier to quantify and
analyze because they can be described as specific ranges of variables like
temperature, latitude, and altitude. For example, the African Fish Eagle occupies a
very similar ecological niche to the American Bald Eagle (Fig. 13). In practice it is
hard to measure all of the variables that a species needs to survive, so descriptions
of an organism's niche tend to focus on the most important limiting factors .
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The full range of habitat types in which a species can exist and reproduce without
any competition from other species is called its fundamental niche. The presence of
other species means that few species live in such conditions. A species' realized
niche can be thought of as its niche in practice—the range of habitat types from
which it is not excluded by competing species. Realized niches are usually smaller
than fundamental niches, since competitive interactions exclude species from at
least some conditions under which they would otherwise grow. Species may
occupy different realized niches in various locations if some constraint, such as a
certain predator, is present in one area but not in another.
In a classic set of laboratory experiments, Russian biologist G.F. Gause showed the
difference between fundamental and realized niches. Gause compared how two
strains of Paramecium grew when they were cultured separately in the same type
of medium to their growth rates when cultured together. When cultured separately
both strains reproduced rapidly, which indicated that they were adapted to living
and reproducing under the same conditions. But when they were cultured together,
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one strain out-competed and eventually eliminated the other. From this work
Gause developed a fundamental concept in community ecology: the competitive
exclusion principle, which states that if two competitors try to occupy the same
realized niche, one species will eliminate the other (footnote 6).
Many key questions about how species function in ecosystems can be answered by
looking at their niches. Species with narrow niches tend to be specialists, relying
on comparatively few food sources. As a result, they are highly sensitive to
changes in key environmental conditions, such as water temperature in aquatic
ecosystems. For example, pandas, which only eat bamboo, have a highly
specialized diet. Many endangered species are threatened because they live or
forage in particular habitats that have been lost or converted to other uses. One
well-known case, the northern spotted owl lives in cavities of trees in old-growth
forests (forests with trees that are more than 200 years old and have not been cut,
pruned, or managed), but these forests have been heavily logged, reducing the
owl's habitat.
In contrast, species with broad niches are generalists that can adapt to wider ranges
of environmental conditions within their own lifetimes (i.e., not through evolution
over generations, but rather through changes in their behavior or physiologic
functioning) and survive on diverse types of prey. Coyotes once were found only
on the Great Plains and in the western United States, but have spread through the
eastern states in part because of their flexible lifestyle. They can kill and eat large,
medium, or small prey, from deer to house cats, as well as other foods such as
invertebrates and fruit, and can live in a range of habitats, from forests to open
landscapes, farmland, and suburban neighborhoods (footnote 7).
Overlap between the niches of two species (more precisely, overlap between their
resource use curves) causes the species to compete if resources are limited. One
might expect to see species constantly dying off as a result, but in many cases
competing species can coexist without either being eliminated. This happens
through niche partitioning (also referred to as resource partitioning), in which two
species divide a limiting resource such as light, food supply, or habitat.
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These common goals drive natural selection for a wide range of traits and
behaviors, including:
Mimicry by either predators or prey. A predator such as a praying mantis that blends in
with surrounding plants is better able to surprise its target. However, many prey species
also engage in mimicry, developing markings similar to those of unpalatable species so
that predators avoid them. For example, harmless viceroy butterflies have similar
coloration to monarch butterflies, which store toxins in their tissues, so predators avoid
viceroy butterflies.
Optimal foraging strategies enable predators to obtain a maximum amount of net
energy per unit of time spent foraging. Predators are more likely to survive and
reproduce if they restrict their diets to prey that provide the most energy per unit of
handling time and focus on areas that are rich with prey or that are close together. The
Ideal Free Distribution model suggests that organisms that are able to move will
distribute themselves according to the amount of food available, with higher
concentrations of organisms located near higher concentrations of food (footnote 8).
Many exceptions have been documented, but this theory is a good general predictor of
animal behavior.
Avoidance/escape features help prey elude predators. These attributes may be
behavioral patterns, such as animal herding or fish schooling to make individual
organisms harder to pick out. Markings can confuse and disorient predators: for
example, the automeris moth has false eye spots on its hind wings that misdirect
predators (Fig. 14).
Features that increase handling time help to discourage predators. Spines serve this
function for many plants and animals, and shells make crustaceans and mollusks harder
to eat. Behaviors can also make prey harder to handle: squid and octopus emit clouds of
ink that distract and confuse attackers, while hedgehogs and porcupines increase the
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Natural selection based on features that make predators and prey more likely to
survive can generate predator-prey "arms races," with improvements in prey
defenses triggering counter-improvements in predator attack tools and vice versa
over many generations. Many cases of predator-prey arms races have been
identified. One widely known case is bats' use of echolocation to find insects. Tiger
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moths respond by emitting high-frequency clicks to "jam" bats' signals, but some
bat species have overcome these measures through new techniques such as flying
erratically to confuse moths or sending echolocation chirps at frequencies that
moths cannot detect. This type of pattern involving two species that interact in
important ways and evolve in a series of reciprocal genetic steps is called
coevolution and represents an important factor in adaptation and the evolution of
new biological species.
Other types of relationship, such as competition, also affect evolution and the
characteristics of individual species. For example, if a species has an opportunity
to move into a vacant niche, the shift may facilitate evolutionary changes over
succeeding generations because the species plays a different ecological role in the
new niche. By the early 20th century, large predators such as wolves and puma had
been largely eliminated from the eastern United States. This has allowed coyotes,
who compete with wolves where they are found together, to spread throughout
urban, suburban, and rural habitats in the eastern states, including surprising
locations such as Cape Cod in Massachusetts and Central Park in New York City.
Research suggests that northeastern coyotes are slightly larger than their
counterparts in western states, although it is not yet clear whether this is because
the northeastern animals are hybridizing with wolves and domestic dogs or because
they have adapted genetically to preying on larger species such as white-tailed
deer.
Just as relationships between individual species are dynamic, so too is the overall
makeup of ecosystems. The process by which one natural community changes into
another over a time scale of years to centuries is called succession. Common
succession patterns include plant colonization of sand dunes and the regrowth of
forests on abandoned farmland (Fig. 15). While the general process is widely
recognized, ecologists have offered differing views of what drives succession and
how to define its end point. By analyzing the natural succession process, scientists
seek to measure how stable ecosystems are at different stages in their trajectory of
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In the early 20th century, plant biologist Frederic Clements described two types of
succession: primary (referring to colonization of a newly exposed landform, such
as sand dunes or lava flows after a volcanic eruption) and secondary (describing
the return of an area to its natural vegetation following a disturbance such as fire,
treefall, or forest harvesting). British ecologist Arthur Tansley distinguished
between autogenic succession—change driven by the inhabitants of an ecosystem,
such as forests regrowing on abandoned agricultural fields—and allogenic
succession, or change driven by new external geophysical conditions such as rising
average temperatures resulting from global climate change.
As discussed above, ecologists often group species depending on whether they are
better adapted for survival at low or high population densities (r-selected versus K-
selected). Succession represents a natural transition from r- to K-selected species.
Ecosystems that have recently experienced traumatic extinction events such as
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floods or fires are favorable environments for r-selected species because these
organisms, which are generalists and grow rapidly, can increase their populations
in the absence of competition immediately after the event. Over time, however,
they will be out-competed by K-selected species, which often derive a competitive
advantage from the habitat modification that takes place during early stages of
primary succession.
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Chapter References:
Paul A. Colinvaux, Why Big Fierce Animals Are Rare: An Ecologist's Perspective (Princeton, NJ:
Princeton University Press, 1979). A survey of major questions in ecology, including why every species
has its own niche.
Chris Reiter and Gina C. Gould, "Thirteen Ways of Looking at a Hedgehog," Natural History,
July/August 1998. Hedgehogs' spines are unique adaptations, but they have thrived in many regions for
millions of years because they are generalists in terms of climate zones and diet.
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Another way to visualize major land biomes is to compare them based on their
average temperature ranges and rainfall levels, which shows how these variables
combine to create a range of climates (Fig. 3).
Figure 3. Biome type in relation to temperature and rainfall Land biomes are typically named for their
characteristic types of vegetation, which in turn influence what kinds of animals will live there. Soil characteristics
also vary from one biome to another, depending on local climate and geology.
Temperate -30 to 30°C 75-150 cm/yr Fertile, high in 3-4 tree species/km2
nutrients
Boreal Very low 40-100 cm/year, mostly Thin, low in nutrients, Evergreens
(taiga) snow acidic
Aquatic biomes (marine and freshwater) cover three-quarters of the Earth's surface
and include rivers, lakes, coral reefs, estuaries, and open ocean (Fig. 4). Oceans
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account for almost all of this area. Large bodies of water (oceans and lakes) are
stratified into layers: surface waters are warmest and contain most of the available
light, but depend on mixing to bring up nutrients from deeper levels (for more
details, see Unit 3, "Oceans"). The distribution of temperature, light, and nutrients
set broad conditions for life in aquatic biomes in much the same way that climate
and soils do for land biomes.
Marine and freshwater biomes change daily or seasonally. For example, in the
intertidal zone where the oceans and land meet, areas are submerged and exposed
as the tide moves in and out. During the winter months lakes and ponds can freeze
over, and wetlands that are covered with water in late winter and spring can dry out
during the summer months.
There are important differences between marine and freshwater biomes. The
oceans occupy large continuous areas, while freshwater habitats vary in size from
small ponds to lakes covering thousands of square kilometers. As a result,
organisms that live in isolated and temporary freshwater environments must be
adapted to a wide range of conditions and able to disperse between habitats when
their conditions change or disappear.
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Since biomes represent consistent sets of conditions for life, they will support
similar kinds of organisms wherever they exist, although the species in the
communities in different places may not be taxonomically related. For example,
large areas of Africa, Australia, South America, and India are covered by savannas
(grasslands with scattered trees). The various grasses, shrubs, and trees that grow
on savannas all are generally adapted to hot climates with distinct rainy and dry
seasons and periodic fires, although they may also have characteristics that make
them well-suited to specific conditions in the areas where they appear.
Species are not uniformly spread among Earth's biomes. Tropical areas generally
have more plant and animal biodiversity than high latitudes, measured in species
richness (the total number of species present) (footnote 1). This pattern, known as
the latitudinal biodiversity gradient, exists in marine, freshwater, and terrestrial
ecosystems in both hemispheres. Figure 5 shows the gradient for plant species, but
it also holds true for animals.
Source: Barthlott, W., Biedinger, N., Braun, G., Feig, F., Kier, G., and Mutke, J. (1999):
Terminology and methodological aspects of the mapping and analysis of global diversity. Acta
Botanica Fennica 162, 103–110.
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Benefits of ecosystems
The interaction of living things depending on each other and relating to their
environments has immense benefits in terms of the health and spiritual wellbeing
of humans, the health of members of the ecosystem themselves, as well as the
environment. Living things do not exist in isolation. They depend on abiotic
factors too. The benefit of ecosystems therefore is not exclusive to living things.
So, what is the role of ecosystems?
A. Supportive
Ecosystems provide a supporting role for all its members. In this role, living members
serve as food for others, and their produce and residue serve as nutrients to soils and
gases to the atmosphere. This makes soil nutrient cycle, carbon and oxygen cycle and
water cycle possible and also for living things to continue procreation.
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B. Provision
Ecosystems are also the source of all foods, store of all energy, fibre, genetic resources,
medicines, fresh water and minerals. All natural resources that humans depend on, has its
source from ecosystems.
C. Regulation
The function of a healthy ecosystem ensures that there is balance and regulation in the
climate, regulation in fresh water, soils, rocks, and atmosphere. They function to regulate
animals and plant diseases and ensure that biodiversity is preserved.
D. Spiritual Value
Perhaps not exclusive to humans, ecosystems provide humans with deeper spiritual
enrichment and cognitive development. The wonder and breathtaking properties of
healthy ecosystems has recreational effects, as well as aesthetic value to us. From
land the remotest places on earth to the deepest places in the oceans, there are
millions of life forms that function in harmony, and provide humans with
meditative and healing benefits.
Threats to Ecosystems
Anything that attempts to alter the balance of the ecosystem potentially threatens
the health and existence of that ecosystem. Some of these threats are not overly
worrying as they may be naturally resolved provided the natural conditions are
restored. Other factors can destroy ecosystems and render all or some of its life
forms extinct. Here are a few:
Habitat Destruction
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for timber can expose the upper layers of the soil to the sun's heat, causing erosion
and drying. It can cause a lot of animals and insects that depended on the shade and
moisture from the tree to die or migrate to other places.
Pollution
Water, land and air pollution all together play a crucial role in the health of
ecosystems. Pollution may be natural or human caused, but regardless they
potentially release destructive agents or chemicals (pollutants) into the
environments of living things. ―In a lake, for example, it can create havoc on the
ecological balance by stimulating plant growth and causing the death of fish due to
suffocation resulting from lack of oxygen. The oxygen cycle will stop, and the
polluted water will also affect the animals dependent on the lake water‖ Source:
Study the effect of pollution on an ecosystem, WWF.
Eutrophication
This is the enrichment of water bodies with plant biomass as a result of continuous
inflow of nutrients particularly nitrogen and phosphorus. Eutrophication of water
fuels excessive plant and algae growth and also hurts water life, often resulting in
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the loss of flora and fauna diversity. ―The known consequences of cultural
eutrophication include blooms of blue-green algae (i.e., cyanobacteria, Figure 2),
tainted drinking water supplies, degradation of recreational opportunities, and
hypoxia. The estimated cost of damage mediated by eutrophication in the U.S.
alone is approximately $2.2 billion annually (Dodds et al. 2009) Source:
Eutrophication: Causes, Consequences, and Controls in Aquatic Ecosystems,
Michael F. Chislock
Invasive species
Any foreign specie (biological) that finds its way into an ecosystem, either by natural or
human introduction can have an effect on the ecosystem. If this alien has the ability to
prey on vulnerable and native members of that ecosystem, they will be wiped out, sooner
or later. One devastating impact of introducing alien Nile Perch and Nile Tilapia into
Lake Victoria in the 1970s was the extinction of almost half of the 350+ endemic species
of fish in the cichlid family.
Overharvesting
Fish species, game and special plants all do fall victim from time to time as a result of
over harvesting or humans over dependence on them. Overharvesting leads to reduction
in populations, community structures and distributions, with an overall reduction in
recruitment. Lots of fish species are know to have reached their maximum exploitation
level, and others will soon be. ―For example Oreochromis karongae is one of the most
valuable food fishes in Malawi, but populations collapsed in the 1990s due to
overfishing, and it is now assessed as Endangered.‖ Source: IUCN, Major Threats
UV Radiation
The sun‘s rays play an important role in living things. UV rays come in three main
wavelengths: UVA, UVB and UVC, and they have different properties. UVA has long
wavelengths and reaches the earth‘s surface all the time. It helps generate vitamin D for
living things. UVB and UVC are more destructive and can cause DNA and cell damage
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to plans and animals. Ozone depletion is one way that exposes living things to UVB and
UVC and the harm caused can wipe lots of species, and affect ecosystems members
including humans.
Preserving Ecosystems
Considering the threats facing ecosystems, we can begin to appreciate the importance of
policy, rules and regulation in human activity towards ecosystems. Here are a few ways
we can ensure the health and smooth functioning of ecosystems.
Habitat preservation
Economic activity should be managed and made sustainable. Tree cutting for example
must be regulated and best practices enforced.
Invasive Species
In many of the tragedies that ecosystems have faced with the introduction of alien species,
humans have caused that. It is crucial that proper inspection, regulation, research and monitoring
systems are in place to protect weaker native species in ecosystems, if new species are to be
introduced.
Eutrophication
One big cause of eutrophication is the runoff of surface chemicals and fertilizers applied
to plants during farming. Whiles we need food to survive, it is important that we
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encourage organic planting as against the heavy use of chemicals. Sewage also need to
be inspected and monitored such that when the waste water is deposited into water
bodies, it is properly filtered and treated to reduce the organic nutrient content.
Pollution
Air and land pollution together have effects on water bodies too. Acid rains and chemical
runoff all affect life forms in the water. Oils that are discharged into water bodies can
have a devastating effect. ―In aquatic ecosystems, air pollution acidifies surface waters,
reducing their ability to sustain native fish. In estuaries and coastal waters, it contributes
to nutrient over-enrichment, producing algal blooms, foul smells and low oxygen levels.
It also causes mercury to accumulate in aquatic food webs, threatening the health of both
people and wild animals2‖
Ozone
Ozone is a secondary pollutant. It is the result of the formation of precursors nitrogen oxides and
Volatile Organic Compounds (VOC). Biomass burning produces this. It is known that forest
cover act as a net sink of ozone. It is therefore important that we preserve natural vegetative
covers on earth and invest in energy forms that reduce the emissions of VOCs.
How are human activities affecting the amount of nitrogen in the environment?
How serious is the threat to biodiversity posed by invasive alien species?
FOCAL AREA | Addressing the major threats to biodiversity, including those arising
from invasive alien species, climate change, pollution, and habitat change
Five main threats to biodiversity are commonly recognized in the programmes of work of
the Convention: invasive alien species, climate change, nutrient loading and pollution,
habitat change, and overexploitation. Unless we successfully mitigate the impacts of
these direct drivers of change on biodiversity, they will contribute to the loss of
biodiversity components, negatively affect ecosystem integrity and hamper aspirations
towards sustainable use.
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Indicators for trends in nutrient loading and invasive alien species have been identified
under the focal area addressed here, and are described below. Information on habitat
change is provided by the indicator trends in extent of selected biomes, ecosystems and
habitats. Overexploitation is discussed under the focal area on sustainable use. While
there is no single indicator of the impacts of climate change on biodiversity, a number of
indicators, including those on trends in extent of selected biomes, ecosystems and
habitats (particularly applied to coral reefs, polar ice and glaciers, and certain types of
forests and drylands), abundance and distribution of selected species, and incidence of
human induced ecosystem failure, can serve to derive trends where specific data are
available. Because small, fragmented ecosystems are more affected by changes in
temperature and humidity than large contiguous ecosystems with a more balanced micro-
climate, trends in connectivity/fragmentation of ecosystems provide an indicator of the
vulnerability of ecosystems to climate change.
Case Study 1:
The ability of agriculture to produce far greater quantities of food and fibre than ever
before can be attributed to a number of factors, including the availability of fertilizers on
a commercial scale. However, excessive levels of the plant nutrients nitrogen and
phosphorus in natural ecosystems are now causing concern. While reactive nitrogen
occurs naturally in all ecosystems, the production of reactive nitrogen by humans, mostly
from manufacturing synthetic fertilizer to increase agricultural production, has changed
ecological balances, both locally and in far-distant ecosystems. Anthropogenic
production of reactive nitrogen leads to the release of nitrogen compounds into the
atmosphere, which are subsequently deposited onto the biosphere. Aerial deposition of
nitrogen increases levels in ecosystems such that those slow growing species that thrive
in nitrogen-poor environments cannot compete with faster-growing species that depend
on higher nutrient levels. Temperate grasslands are particularly vulnerable in this respect.
Moreover, soluble nitrogen leaches from soils into groundwater, resulting in increased
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Figure 2.15 Global trends in the creation of reactive nitrogen on Earth by human activity
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Figure 2.16 Estimated total reactive nitrogen deposition from the atmosphere (wet and dry)
(early 1990s)
To continue to meet global demand for food and fibre and minimize
environmental problems, significant improvements are required in the
efficiency with which nitrogen fertilizer is utilized within production
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Case Study 2:
Invasive alien species can have devastating impacts on native biota, causing extinctions
and affecting natural and cultivated ecosystems. Since the 17th century, invasive alien
species have contributed to nearly 40% of all animal extinctions for which the cause is
known. In the Fynbos biome of South Africa, 80% of the threatened species are
endangered because of invading alien species.
A proportion of invasive alien species are important pests or pathogens that can cause
enormous economic costs. The annual environmental losses caused by introduced pests in
the United States, United Kingdom, Australia, South Africa, India and Brazil have been
calculated at over US$ 100 billion. Invasive alien species can transform the structure and
species composition of ecosystems by repressing or excluding native species. Because
invasive species are often one of a whole suite of factors affecting particular sites or
ecosystems, it is not always easy to determine the proportion of the impact that can be
attributed to them. In the recent past, the rate and risk associated with alien species
introductions have increased significantly because human population growth and human
activities altering the environment have escalated rapidly, combined with the higher
likelihood of species being spread as a result of increased travel, trade and tourism.
A major source of marine introductions of alien species is hull fouling and the release of
ballast water from ships, although other vectors, such as aquaculture and aquarium
releases, are also important and less well regulated than ballast water. In the marine
ecosystem, the movement of non-native species has been well studied. Of the 150 species
that have recently arrived in the Great Lakes, 75% originated from the Baltic Sea.
Similarly, migration flow from the Red Sea to the Mediterranean through the Suez Canal
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continues unabated with nearly 300 species of these Lessepsian migrants, including
decapod crustaceans, molluscs and fishes, having entered the Mediterranean since 1891.
Equally long-term data available from five Nordic countries (Iceland, Denmark, Norway,
Sweden and Finland) that have recorded the cumulative number of alien species in
freshwater, marine and terrestrial environments since 1900 demonstrate the continuing
arrival of new immigrants of plants, vertebrates and invertebrates (Figure 2.17).
Invasive alien species are a global problem requiring responses at all levels. Many
countries have established systems to prevent and control invasive alien species and, as
part of risk assessments, to predict the likelihood of alien species becoming invasive and
the potential ecological and economic cost they may incur. To effectively communicate
the challenges posed by invasive alien species there is a need to develop a methodology
for integrating information quantifying the threat and its impacts on biodiversity into a
coherent indicator.
Figure 2.17 Number of alien species recorded in the Nordic terrestrial, freshwater and marine
environment
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Case 3:
Ecosystems are dynamic and complex assemblages of organisms that interact with each
other and with the physical environment. Conversion, degradation, or the unsustainable
management of a natural ecosystem has far-reaching consequences: it results in a change
of the relative abundance of individual species, and frequently the loss of populations,
and also in the reduction or loss of ecosystem services. Over the last 50 years, humans
have changed ecosystems more rapidly and extensively than in any comparable period of
time in human history. Reducing the rate at which ecosystems are being degraded or lost
is therefore a key contribution towards the achievement of the 2010 Biodiversity Target.
For most of the world‘s main habitats and ecosystems, neither the current global extent
nor rates of change in that extent are known with high certainty. This is due in part to the
challenges of measuring global habitat extent, differences in definitions and classification
systems and the lack of historical data. The exception is forests, many of which have
direct commercial and/or scientific value, and are therefore regularly inventoried and
assessed in most countries. Even here, however, there are limitations in analyses to date
that make it difficult to assess, for example, changes in primary forests.
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In the absence of human influence, forests and woodlands covered approximately half of
the Earth‘s land surface. However, thousands of years of human activity have reduced
their extent to about 30% of total land area. Of this area only one-third is considered
primary forest—forest of native species where ecological processes are not significantly
disturbed by human activities. Deforestation, mainly conversion of forests to agricultural
land and pasture, continues at an alarmingly high rate: about 13 million hectares—
equivalent to the area of Greece or Nicaragua—are lost each year. At the same time, tree
planting, landscape restoration and natural expansion of forests have significantly offset
the loss of primary forest area. It should be borne in mind, however, that the biodiversity
value of forest plantations and secondary forests is generally much lower than that of
forests. Figure 2.1 presents the trends in net forest area by region. The net loss in forest
area in the period 2000–2005 is estimated at 7.3 million hectares per year, equivalent to
an annual loss of 0.18% of net forest area. This compares to 8.9 million hectares (0.22%)
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per year from the period 1990 to 2000. Over the last 15 years, primary forest has been
lost or modified at a rate of approximately six million hectares a year.
Africa and South America continue to have the largest net loss of forests. Oceania and
North and Central America also show a net loss of forests. The forest area in Europe
continued to expand, although at a slower rate. Asia, which had a net loss in the 1990s,
reported a net gain of forests in the period 2000–2005, primarily due to large-scale
afforestation reported by China. There is recent evidence, however, of increases in the
frequency and extent of natural disturbances (fire, insect outbreaks and disease) in boreal
forests, which negatively affect forest cover in those ecosystems.
Figure 2.2 Locations reported by various studies as undergoing high rates of change in forest
cover in the past few decades
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On the basis of various studies from the period of 1980 to 2000, the Millennium Ecosystem
Assessment prepared a map showing the areas undergoing high rates of change in forest cover
(Figure 2.2)
Figure 2.3 Change in live coral cover across the Caribbean basin (1977-2002)
General patterns of change in the extent of ecosystems across other biomes besides
forests show similar negative trends. The Millennium Ecosystem Assessment reported
that almost 70% of Mediterranean forests, woodlands and scrub, 50% of tropical and sub-
tropical grasslands, savannas and shrublands and 30% of desert ecosystems had been lost
by 1990. Coastal and marine ecosystems have been heavily impacted by human activities,
with degradation leading to a reduced coverage of kelp forests, seagrasses and corals. In
the Caribbean, average hard coral cover declined from about 50% to 10% in the last three
decades, equivalent to a loss of almost 7% of remaining area covered by live coral each
year since the 1970s (Figure 2.3). Some 35% of mangroves have been lost in the last two
decades in countries for which adequate data are available. This is equivalent to an
annual loss of 2% of the remaining area. There has been a widespread retreat of mountain
glaciers in non-polar regions during the 20thcentury, and decreases of about 10% in the
extent of snow cover since the late 1960s. In the Arctic the average annual sea ice extent
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has declined by about 8% in the past 30 years, with a loss of 15 to 20% in summer sea ice
extent over the same period.
Case. 4 :
Threatened species occur across all taxonomic groups and in all parts of the world. Over
the past few hundred years, it is estimated that humans have increased species extinction
rates by as much as 1,000 times the background rates typical over Earth‘s history.
Between 12% and 52% of species within well-studied higher taxa are threatened with
extinction, according to the IUCN Red List of Threatened Species.
Figure 2.6 Red List Index for birds in marine, freshwater and terrestrial ecosystems, and in
forest and shrubland/grassland habitats (1988-2004)
On the basis of Red List data, a Red List Index can be calculated for different taxonomic
groups or geographic regions to show trends in the proportion of species expected to
remain extant in the near future without additional conservation interventions. The index
is based on the number of species present in each Red List category, and on the number
that change categories over time (i.e., between assessments), as a result of genuine
improvement or deterioration in status. This index shows a continuing deterioration in the
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status of bird species, which have been completely assessed for the IUCN Red List four
times over the last two decades, across all biomes (Figure 2.6). Despite limitations in our
knowledge about the total number of species and their status, preliminary findings for
other major groups, such as amphibians and mammals, indicate that the situation is likely
worse than for birds.
The Red List Index is highly representative, being based on assessments of a high
proportion of species in a taxonomic group across the world, but it shows a coarse level
of resolution because of the width of the Red List categories. Some of the Red List
criteria are based on absolute population size or range size, while others are based on
rates of decline in these values or combinations of absolute size and rates of decline.
Because the Red List Index is based on a proportional change in a measure and its values
relate to the rate at which species are slipping towards extinction at particular points in
time, a downward trend, even if becoming less steep, shows that the slide of the species
towards extinction is accelerating, rather than slowing down. The 2010 Biodiversity
Target would therefore only be met when a positive trend is achieved.
Land use pattern in India relates to the physical characteristics of land, the institutional
and other resources framework like labour, capital available. All these aspects are
associated with the economic development. India has a total land area of approximately
328 million hectares. Mostly, land utilisation statistics are obtainable for almost 93 % of
the entire area that is around 306 million hectares. It is considerable to note that every
forefather over the past 8,000 years or so have been successful in harbouring nearly 140
million hectares of land from the natural ecosystem to agriculture. From the time of
independence, people have been successful to add another 22 million hectares. As a
result, 162 million hectares of land excels as the net sown area at present. It forms a
stupendous percentage of as high as 51%. No other large country is as fortunate as India
in this regard.
The land for which the data on classification of land-use is available is known as
Reporting Land. In some cases the reporting land is that land, where the land use
pattern figures are supported on land records and are based on village records or
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papers. These records are preserved by village revenue agency and here the data is
completely based on details of entire areas. In cases, where the records are not
preserved, the estimates are mostly based on sample survey. Thus, the statistics of land
use pattern are based on these two methods. On the other hand, the lands where no
data is available are known as Non-Reporting Lands.
Uncultivated Land
According to the available land use statistics, there has been a slight increase in the net
sown area. Almost 28 million hectares have been added over the passing few decades.
Around 1.3 % of the land is under fruit trees. Nearly 5 % of the land falls in the category
of uncultivated land which is cultivated once every 2 to 3 years. Thus, near about 51% of
the whole area, on an average, is cultivated once a year. The uncultivated lands are
subsidiary lands and are kept so to re-establish their richness. Its use depends upon
high-quality and timely rains also.
The area separated as cultivable waste, has remained stationary at around 6.4 % for
several decades. The land under permanent pastures is despondently low and suggests
a remarkable population pressure on the land. Also, credit must be confirmed on the
farmers that with so modest land under pastures, they have the biggest number of
cattle. They are nurtured mainly on husk, grain chaff, farm waste and few fodder crops.
This is definitely the most economical way to have a larger number of drought animals
and bovine cattle. Areas that have been classed under forests are also used for cattle
grazing.
Forested land in India is far less in scientific norm. For a self-sufficient economy and
accurate ecological steadiness, at least one third of the total land area must be kept
under forests and natural vegetation. In India, it is as low as 19.27%. Photographic
proof, gained from satellites has confirmed that only about 46 million hectares come
under real forests, as opposed to the estimated 63 million hectares, according to the
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figures of land use pattern. However, this outline establishes a tiny rise from 40 million
hectares.
Forest Lands
It is important for the populace to ensure that they increase the area under forests for
reasons more than one. A bigger area under forests is an obligation, to maintain the
ecological balance and for absorption of carbon dioxide, the assemblage of which is
likely to heighten the green house effect. This in turn would raise atmospheric
temperature at the global stage. It may lead to thawing of ice caps and equivalent rise in
sea level, jeopardising low-lying densely populated parts of the world. Forests supply
home to wildlife and help their continuation. They help in enhancing the level of rainfall,
minimising cases of famine. Forested lands also help in permeation of rainwater in the
subsoil and modulating the flow of river waters in both rainy and dry seasons. Forests
safeguard not only water but soil as well. They, thus, help in plunging the volume of
floodwaters and their ferocity.
Wasteland
A part of the land that is not utilised for the moment is classified as wasteland. This
embraces the baked and rocky deserts. High mountainous and uneven lands also fall
into this category. At times humankind has also been responsible to add to such areas
by deforestation and overgrazing.
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The mounting population and advanced standards of living have resulted in an ever
increasing demand for residential land, both in villages and towns. Cities and towns are
obligated to grow vertically rather than horizontally. Land is needed to develop industry,
commerce, transport and recreational facilities. In view of mounting pressure on land
for numerous purposes, it is customary to plan appropriate use of all the obtainable
land. This may be done by following fitting measures to control soil erosion,
desertification etc. which turns cultivatable land into wildernesses. In addition, some of
the barrens may be brought around for different uses. Likewise, with the help of up-to-
date and scientific methods of farming, productivity of land can also be amplified. All
endeavours should be made to strike a balance amongst diverse use of land.
In India the capacity for expansion of cultivation to further new areas is very restricted.
As of now, 49% of the entire reporting land is cultured. Fallow and other waste lands,
including grazing pastures, which are not currently cultivated, is presumed around 42
million hectares, and further expansion of cultivation to such lands would be expensive
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In the dynamic context, keeping in view the natural endowments and the recent
advances in technology, the overall interests of a country may dictate a certain
modification of or a change in the existing land-use pattern of a region. A proper study
of the present land-use patterns and the developing trends will help to suggest the
scope for planned shifts in the patterns in India.
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As you develop and grow there are certain predictable changes that will occur, and they usually
happen in a specific order. As a baby, you first learn to roll over, then crawl, then walk. You also
learn new skills such as how to feed yourself and how to talk, but each skill you learn builds on
those learned before it. Your body also changes as you grow taller, your muscles and bones
develop, your hair grows longer, etc.
Ecological succession is the same idea. It is the observed changes in an ecological community
over time. These changes are fairly predictable and orderly. Within an ecological community, the
species composition will change over time as some species become more prominent while others
may fade out of existence. As the community develops over time, vegetation grows taller, and
the community becomes more established. (Or)Ecological succession is the observed process of
change in the species structure of an ecological community over time. The time scale can be
decades (for example, after a wildfire), or even millions of years after a mass extinction.
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For those unfamilire with the subject matter, there are two types of Ecological
Succession. First there is Primary Succession and then there is Secondary
Succession. The primary difference between the two types of succession lies in the
oragins of the ground upon which the plants take root.
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enough conditions. Included in the following are the general processes of both
primary and seccondary succession.
Primary Succession:
These lichens and algea secrete digestive acids which help break down the
rock's surface material for better nutrient absorption. These digestive acids help
form tiny cracks in the rocks surface which are then widened over time as the
mositure undergoes natural freezing and thawing processes. These cracks
eventually grow large enough to trap enough organic materal and moisture for
various moss species to take hold. These mosses also secrete digestive acids which
assist in the breaking down of the rock's surface helping to form even larger cracks
as they grow.
These cracks eventually collect enough soil to support grasses, flowers, and
even small shrubs. Over time the roots of these plants burrow even further into the
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cracks in the rock's surface. This allows the water to burrow further into the rock as
well where it also freezes and thaws further widening the cracks in the rock. With
the rooting of each new plant. the size of both the crack in the rock's surface and
the size of the next plant able to take root there increases. They have a direct
corrolation to each other. Eventually, several of the larger cracks form together
creating a basin where trees can then take root. The eventual end result of primary
succession is a climax community. However, it can often take hundreds, or in
worse cases, even thousands of years for a climax community to become a reality.
Secondary Succession:
Secondary succession is the redevelopment of an ecological community where
an ecological community has already previously existed but had since been wiped
out for some reason or another. There are several primary causal sources of
seccondary succession. These primary causal sources include things such as Lake
Succession, Dune Succession, Fire Succession, Clear Cutting Succession, and Old
Field Succession. For the general purpose of my assignment I am only going to go
into detail about the proccess of Old Field Succession since this is the only one of
the five which falls under the scope of my project.
While the strip of land in my back yard is not an old field, its environment has
resulted from a churning of earth much like that which happens in a farmers field.
Like the old field abandoned by the farmer, the strip of earth in my back yard has
also been left to seed. In old fields, grasses are usually the first plants to take root.
These plants are usually followed by low lying plants such as dandylions and other
short flowering plants. As these cycle through taller plants such as milkweeds,
goldenrods, and asters take root. These are then followed up by taller and taller
plants.
The reason for this is due to the competition for light sources. Any new plants
that want to take root can not be content to grow in the shadow of the plants that
have come before it as it will not be able to undergo photosynthesis. Inorder to
compeat for the light source it must be able to rise above and out distance the
canopy of the shorter plants which would otherwise crowd it out. As the plants get
taller they often become more woody inorder to combat gravity. This process
eventually leads to the planting of shrubs and other small trees such as Sassafrases
and Hawthornes.
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As the plants grow taller they have a tendency to shade out the shorter plants.
These shorter plants will eventually die off leaving more space for taller plants
which must breach the canopy of the medium sized plants that came before it. This
escalating cycle eventually leads to the growth of larger trees such as Oaks,
Maples, Hickories,and Beeches. As with primary succession, this proccess will
eventually result in a mature forest, better known as a climax community. Also
known as old growth forests, such communities have a layering of trees and shrubs
in varying hights and sizes. Unlike the way it is with primary succession, if left to
itself the secondary succession proccess does not usually take more than 75-150
years to become a climax community.
Succession may be initiated either by formation of new habitat (landslide or lava flow) or
disturbance of already existing habitat (fires, land clearance). There are three recognized
stages to ecological succession. Each covers a gradual process of change and
development. They do not have hard and defined boundaries, and it is possibly for an
ecological system to be in both stages at once during the transition period from one to
another. The 3 stages of ecological succession are:
1. Primary – This is when an ecological community first enters into a new form of
habitat that it has not been present in before. A good example of this would be the habitat
created when granite is removed in a quarry. The rock face that is left behind is altered
and becomes a new habitat. The environment that then grows within that habitat is
considered to be in its primary stage.
2. Secondary – The secondary succession stage occurs after a habitat has been
established, but it is then disturbed or changed in some fashion and a new community
moves in. To use the example from before – let us say that a primary stage develops on
the face of a newly quarried granite cliff. That habitat grows undisturbed, until there is a
forest fire that then burns and changes a portion of the habitat that has been growing on
the rock face. That ecological habitat has now entered its secondary stage.
3. Climax – the climax stage is the last stage of an ecosystem. It is when the ecosystem
has become balanced and there is little risk of an interfering event or change to mutate the
environment. Several rainforests and deserts qualify as being in the climax stage. What is
tricky about a climax stage is that given human development, any ecosystem that is in the
climax stage now holds the risk of being destroyed and going backward in the stages.
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When talking about the types of ecological succession it is important to remember that
the ―types‖ occur within the stages, but they may not necessarily be unique to that stage.
What determines the stage that an ecosystem is in is dependent on its energy balance –
which is discussed in the next section. There are four main types of ecological
succession:
Pioneer – pioneer types are the new lifeforms that enter into a primary
succession and begin to take hold. This can be anything from a seed to a bacteria
to an insect or to an animal wandering into a new area and bedding down to
make it their home. The pioneer has no connection to the environment, but it
does find enough present in the new ecosystem to begin to establish its life.
Sustaining – Sustaining type means that life in the ecosystem has begun to enter
into a pattern that allows for a cycle of life to continue. This means that birth and
death are occurring, and there is little migration outside of the ecosystem – this is
most common in the climax succession.
Producing – the producing type occurs during the secondary succession. This is
when lifeforms are breeding and growing, but there is migration because what is
produced is also not capable of being supported within the ecosystem. There are
also more areas of overgrowth or overpopulation due to seed levels.
Pioneer species are the ones that thrive the new habitat at the beginning of ecological
succession. Pioneer species are ‗r-selected‘ species that are fast growing and well-
dispersed. Early succession is therefore dominated by so called ‗r-selected‘ species. As
succession continues, more species enter the community and begin to alter the
environment. These are called ‗k-selected‘ species. They are more competitive and fight
for resource and space. The species that are better suited for the modified habitat then
begin to succeed the other species. These are superseded by newer set of species. This
goes on till the stage of climax or equilibrium is achieved.
When succession reaches a climax, where community is dominated by stable and small
number of prominent species and no other species can be admitted, that is called the state
of equilibrium or the climax community.
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The climax stage of ecological succession is defined by the energy balance that is
achieved. This means that within this very stable ecological system, there is a balance
between the life that is produced, and the life that is consumed. For example, there are
enough animals to eat the extra seeds to prevent overgrowth that could choke out plants,
but not enough to prevent some of the seeds from growing and continuing their cycle of
life. The climax stage is stable, but not static. During the other stages, the balance of
energy is not in place and there may be crises that develop as a result which will prolong
the secondary stage.
Each stage of ecological succession can take 100s to 1,000s of years – if not more. That
is true, but only in a forensic sense. The assumption of ecological succession is that it is a
forward moving, and linear path. As more of humankind encroaches on the natural world,
the linear progression of this methodology is changing itself. That someone seems fitting
for a theory that talks about the inevitability of change.
To best illustrate this, let us return to our first example – the rock face. Let us suppose
that the granite wall was quarried by man, and then abandoned once they had what
they needed. This allows for a primary stage to begin. Left alone by man, it could quickly
pass into a secondary stage within a hundred years or so. Another few centuries after
that, the old quarry is slowly entering its stable climax stage – except – now man has
returned to build a road. One thing that ecological succession recognizes is the death of
an ecosystem. That is what occurs when a climax stage ecosystem like the rain forest is
destroyed by logging. Yet when a climax stage ecosystem is only interrupted, it is not yet
understood whether it returns to the secondary stage, or would still be considered at its
climax of ecological succession.
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Bioindicators
Introduction
Bioindicators are organisms, such as lichens,birds and bacteria, that are used to monitor
the health of the environment. The organisms and organism associations are monitored
for changes that may indicate a problem within their ecosystem. The changes can be
chemical, physiological or behavioural.
What can the canary in the coal mine tell us? Historically, canaries accompanied coal
miners deep underground. Their small lung capacity and unidirectional lung ventilation
system made them more vulnerable to small concentrations of carbon monoxide and
methane gas than their human companions. As late as 1986, the acute sensitivity of these
birds served as a biological indicator of unsafe conditions in underground coal mines in
the United Kingdom. Since human health concerns continue to drive the development
and application of bioindicators, the loss of ecosystem services (e.g., clean air, drinking
water, plant pollinators) has increasingly focused our attention on the health of natural
ecosystems. All species (or species assemblages) tolerate a limited range of chemical,
physical, and biological conditions, which we can use to evaluate environmental quality.
Despite many technological advances, we find ourselves turning to the biota of natural
ecosystems to tell us the story of our world.
Figure 1: Comparison of environmental tolerances of (a) bioindicators, (b) rare species, and (c)
ubiquitous species
However, not all biological processes, species, or communities can serve as successful
bioindicators. Physical, chemical, and biological factors (e.g., substrate, light,
temperature, competition) vary among environments. Over time, populations evolve
strategies to maximize growth and reproduction (i.e., fitness) within a specific range of
environmental factors. Outside an individual's environmental optima, or tolerance range,
its physiology and/or behavior may be negatively affected, reducing its overall fitness
(Figure 1). Reduced fitness can subsequently disrupt population dynamics and alter the
community as a whole (Figure 2). Bioindicator species effectively indicate the condition
of the environment because of their moderate tolerance to environmental variability
(Figure 1). In contrast, rare species (or species assemblages) with narrow tolerances are
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In common usage, the terms "biomonitoring" and "bioindication" are interchangeable, but
in the scientific community these terms have more specific meanings. Bioindicators
qualitatively assesses biotic responses to environmental stress (e.g., presence of the
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Each element is represented by a different set of colored dots (red, Aluminum; yellow,
Zinc; green, Lead; blue, Cadmium). The greatest concentration of each element occurred
close to the road and declined with distance from the road, demonstrating a marked
impact of overland transport of mined ore on the biota.
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Considering the 1.7 million species that currently documented on Earth, how do
we chose just one as a bioindicator? The answer is simple: No single species can
adequately indicate every type of disturbance or stress in all environments.
Depending upon the specific environment, the species present, and local
disturbances, appropriate bioindicator species or groups of species need to be
selected. Ecologists have established a broad set of criteria that species must
exhibit to be considered good bioindicators (see Table 1).
The numerous benefits of bioindicators have spurred legislative mandates for their use in
countries around the world and their inclusion in several international accords. Yet
bioindicators are not without their problems. Like the canaries in the coal mine, we rely
upon the sensitivity of some bioindicators to function as early-warning signals. In some
instances, we cannot discriminate natural variability from changes due to human impacts,
thus limiting the applicability of bioindicators in heterogeneous environments.
Accordingly, populations of indicator species may be influenced by factors other than the
disturbance or stress (e.g., disease, parasitism, competition, predation), complicating our
picture of the causal mechanisms of change. A second criticism of the use of
bioindicators is that their indicator ability is scale-dependent. For example, a large
vertebrate indicator (e.g., a fish) may fail to indicate the biodiversity of the local insect
community. Third, bioindicator species invariably have differing habitat requirements
than other species in their ecosystem. Managing an ecosystem according to the habitat
requirements of a particular bioindicator may fail to protect rare species with different
requirements. Finally, the overall objective of bioindicators is to use a single species, or a
small group of species, to assess the quality of an environment and how it changes over
time, but this can represent a gross oversimplification of a complex system.
Like all management tools, we must be conscious of its flaws. However, the limitations
of bioindicators are clearly overshadowed by their benefits. Bioindicators can be
employed at a range of scales, from the cellular to the ecosystem level, to evaluate the
health of a particular ecosystem. They bring together information from the biological,
physical, and chemical components of our world that manifest themselves as changes in
individual fitness, population density, community composition, and ecosystem processes.
From a management perspective, bioindicators inform our actions as to what is and is not
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biologically sustainable. Without the moss in the tundra, the cutthroat in the mountain
stream, and the canary in the coal mine, we may not recognize the impact of our
disturbances before it is too late to do anything to prevent them.
Conservation status
The conservation status of a species indicates the likelihood that it will become extinct.
Many factors are considered when assessing the conservation status of a species; e.g.,
such statistics as the number remaining, the overall increase or decrease in the population
over time, breeding success rates, or known threats.[2] The IUCN Red List of Threatened
Species is the best-known worldwide conservation status listing and ranking system.[3]
Extinct (EX)
Examples: aurochs
Bali tiger
blackfin cisco
Caribbean monk seal
Carolina parakeet
Caspian tiger
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dodo
dusky seaside sparrow
eastern cougar
golden toad
great auk
Japanese sea lion
Javan tiger
Labrador duck
passenger pigeon
Schomburgk's deer
Steller's sea cow
thylacine
toolache wallaby
western black rhinoceros
Examples: addax
African wild ass
Alabama cavefish
Amur leopard
Arakan forest turtle
Asiatic cheetah
axolotl
Bactrian camel
black rhino
blue-throated macaw
Brazilian merganser
brown spider monkey
California condor
Chinese alligator
Chinese giant salamander
gharial
Hawaiian monk seal
Javan rhino
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kakapo
Leadbeater's possum
Mediterranean monk seal
mountain gorilla
northern hairy-nosed wombat
Philippine eagle
red wolf
saiga
Siamese crocodile
Malayan tiger
Spix's macaw
southern bluefin tuna
South China tiger
Sumatran orangutan
Sumatran rhinoceros
Sumatran tiger
vaquita
Yangtze river dolphin
northern white rhinoceros
hawksbill sea turtle
Kemp's ridley sea turtle
Endangered (EN)
Faces a high risk of extinction in the near future.
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Vulnerable (VU)
Faces a high risk of endangerment in the medium term.
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mountain zebra
polar bear
red panda
sloth bear
takin
yak
great white shark
American crocodile
dingo
king cobra
Near-threatened (NT)
May be considered threatened in the near future.
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brown-throated sloth
Canada goose
cane toad
common wood pigeon
cougar
common frog
giraffe
grey wolf
house mouse
wolverine[6]
human
palm cockatoo
mallard
meerkat
mute swan
platypus
red-billed quelea
red-tailed hawk
rock pigeon
scarlet macaw
southern elephant seal
milk shark
red howler monkey
1. An observed, estimated, inferred or suspected population size reduction of ≥ 70% over the last
10 years or three generations, whichever is the longer, where the causes of the reduction are
clearly reversible AND understood AND ceased, based on (and specifying) any of the following:
1. direct observation
2. an index of abundance appropriate for the taxon
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1. Extent of occurrence estimated to be less than 5,000 km², and estimates indicating at least two
of a-c:
1. Severely fragmented or known to exist at no more than five locations.
2. Continuing decline, inferred, observed or projected, in any of the following:
1. extent of occurrence
2. area of occupancy
3. area, extent and/or quality of habitat
4. number of locations or subpopulations
5. number of mature individuals
3. Extreme fluctuations in any of the following:
1. extent of occurrence
2. area of occupancy
3. number of locations or subpopulations
4. number of mature individuals
2. Area of occupancy estimated to be less than 500 km², and estimates indicating at least two of a-
c:
1. Severely fragmented or known to exist at no more than five locations.
2. Continuing decline, inferred, observed or projected, in any of the following:
1. extent of occurrence
2. area of occupancy
3. area, extent and/or quality of habitat
4. number of locations or subpopulations
5. number of mature individuals
3. Extreme fluctuations in any of the following:
1. extent of occurrence
2. area of occupancy
3. number of locations or subpopulations
4. number of mature individuals
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C) Population estimated to number fewer than 2,500 mature individuals and either:
1. An estimated continuing decline of at least 20% within five years or two generations,
whichever is longer, (up to a maximum of 100 years in the future) OR
2. A continuing decline, observed, projected, or inferred, in numbers of mature individuals AND at
least one of the follow (a-b):
1. Population structure in the form of one of the following:
1. no subpopulation estimated to contain more than 250 mature individuals, OR
2. at least 95% of mature individuals in one subpopulation
2. Extreme fluctuations in number of mature individuals
E) Quantitative analysis showing the probability of extinction in the wild is at least 20%
within 20 years or five generations, whichever is the longer (up to a maximum of 100 years).
1.
Near-critically endangered.
Particularly sensitive to poaching levels.
1. Habitat destruction
2. Degradation and
3. Over exploitation of resources
The RED-data book contains a list of endangered species of plants and animals. It contains a list
of species of that are endangered but might become extinct in the near future if not protected.
Some of the rarest animals found in India are:
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1. Asiatic cheetah
2. Asiatic Lion
3. Asiatic Wild Ass
4. Bengal Fox
5. Gaur
6. Indian Elephant
7. Indian Rhinocerous
8. Marbled Cat
9. Markhor
Endangered or threatened species is one whose number has been reduced to a critical number.
Unless it is protected and conserved, it is in immediate danger of extinction.
Vulnerable species is one whose population is facing continuous decline due to habitat
destruction or over exploitation. However, it is still abundant.
Rare species is localized within a restricted area or is thinly scattered over an extensive area.
Such species are not endangered or vulnerable.
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An international treaty to help protect endangered wildlife is, "Convention on International Trade
in Endangered Species 1975" (CITES). This treaty is now signed by 160 countries.
1. CITES lists 900 species that cannot be commercially traded as live specimens or wildlife
products as they are in danger of extinction.
2. CITES restricts trade of 2900 other species as they are endangered.
DRAWBACKS OF CITES
1. This treaty is limited as enforcement is difficult and convicted violators get away by
paying only a small fine.
2. Member countries can exempt themselves from protecting any listed species.
Species that are found only in a particular region are known as endemic species. Almost 60% the
endemic species in India are found in Himalayas and the Western Ghats. Endemic species are
mainly concentrated in:
1. North-East India
2. North-West Himalayas
3. Western Ghats and
4. Andaman & Nicobar Islands.
1. Sapria Himalayana
2. Ovaria Lurida
3. Nepenthis khasiana etc
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1. Habitat loss and fragmentation due to draining and filling of inland wetlands.
2. Pollution also plays an important role.
Ex:
1. Frog eggs, tadpoles and adults are extremely sensitive to pollutants especially pesticides.
2. Over-hunting and
3. Populations can be adversely affected by introduction of non active predators and
competitors. Disease producing organisms also play an important adversary in reducing
populations of endemic species.
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Suggested Readings:
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Borman, F.H. and G.E. Likens. 1970. "The nutrient cycles of an ecosystem." Scientific
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Wessells, N.K. and J.L. Hopson. 1988. Biology. New York: Random House. Ch. 44.
Burgess, Ernest W. (1925) 1961 The Growth of the City: An Introduction to a Research
Project. Pages 37–44 in George A. Theodorson (editor), Studies in Human Ecology.
Evanston, Ill.: Row, Peterson.
Christaller, Walter 1933 Die zentralen Orte in Siiddeutschland: Eine okonomisch-
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Duncan, Otis Dudley 1959 Human Ecology and Population Studies. Pages 678–716 in
Philip M. Hauser and Otis Dudley Duncan (editors), The Study of Population: An
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Hawley, Amos H. 1950 Human Ecology: A Theory of Community Structure. New York:
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McKenzie, Roderick D. (1924) 1925 The Ecological Approach to the Study of the
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American Journal of Sociology.
Park, Robert E. (1936) 1952 Human Ecology. Pages 145–158 in Robert E. Park, Human
Communities: The City and Human Ecology. Collected Papers, Vol. 2. Glencoe, Ill.: Free
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Quinn, James A. 1950 Human Ecology. Englewood Cliffs, N.J.: Prentice-Hall.
Schnore, Leo F. 1958 Social Morphology and Human Ecology. American Journal of
Sociology 63:620–634.
Steward, Julian H. 1955 Theory of Culture Change: The Methodology of Multilinear
Evolution. Urbana: Univ. of Illinois Press.
golley, frank. a primer for ecological literacy. new haven, conn.: yale university press,
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leopold, aldo. "the land ethic." in a sand county almanac. new york: oxford university
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northcott, michael s. the environment and christian ethics. cambridge, uk: cambridge
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pimentel, david; westra, laura; and noss, reed f., eds. ecological integrity: integrating
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rolston, holmes, iii. "the bible and ecology." interpretation: journal of bible and theology
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soulé, michael e., and lease, gary, eds. reinventing nature? responses to postmodern
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Billington, Elizabeth T. Understanding Ecology. New York: F. Warne, 1971.
Curtis, Helena, and N. Sue Barnes. Biology, 5th ed. New York: Worth Publishing, 1989.
Darwin, Charles. On the Origin of Species. Danbury, CT: Grolier Enterprises, 1981.
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Haeckel, Ernst. The Riddle of the Universe, trans. Joseph McCabe. New York: Buffalo
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Miller, G. Tyler, Jr. Living in the Environment, 6th ed. Belmont, CA: Wadsworth, 1990.
Pringle, Laurence. Lives at Stake: The Science and Politics of Environmental Health.
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Purves, William K., and Gordon H. Orians. Life: The Science of Biology. Sunderland,
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Rosenberg, D. M. & Resh, V. H. Freshwater Biomonitoring and Benthic
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