Professional Documents
Culture Documents
INTRODUCTION
biological variation. It is the variety of life and it plays an important role as collectively to
variations at all levels of biological organizations (Gaston and Spicer, 1998). All species are
interconnected and are seemingly dependent on one another. According to ClientEarth (2021),
biodiversity is the natural world around us, and the variety of all of the different kinds of organisms
– the plants, animals, insects and micro-organisms that live on our planet it is one of the most
precious and important things we have. The word biodiversity originates from the Greek word
Bios means life and the Latin word Diversitas means variety or difference; therefore, the word
biodiversity means the variety of life. Definitely, a lot of aspects we can deal with biodiversity and
its ecosystem. Without biodiversity, our entire support system for human, as well as animal life,
would collapse. We rely on nature to provide us with food and clean water, for a lot of medicines,
and to prevent flooding and other extreme weather effects. Our biodiversity is being dominated by
plants and herbivores that consumes them (Futuyma et al., 2009). Ecological niche breadth
(specialization) and niche differentiation (complementarily) play a crucial role for species
coexistence and hence biodiversity (Bluthgen et al., 2011). Despite the fact that species may seem
redundant when considering one function under a particular set of environmental variables, many
species are required to maintain several roles at various times and locations in a changing world
earth’s terrestrial surface (Suttie et al., 2005). Grasses are a plant with narrow leaves growing from
the base and some common grasses are used to cover the ground. These grasses are the most
valuable horizontal green plant with about 11,000 species, and more than 650 types. It is one of
the largest families of the flower plants (Almusaed, 2017). Grasses are the low, green, nonwoody
plants belonging to the grass family Poaceae, the sedge family Cyperaceae, and the rush family
Juncaceae (Lotha, 2023). They are usually seen in steppes or prairies, tundra, coastal marshes,
pioneer or early successional communities, disturbed sites and in certain aquatic communities (Lu
& Liu, 2002). Grasses that comes from the family of Graminea, has an important group of plants
that are deemed important in a variety of environment (Gould & Shaw, 1983).
Grasses are agriculture’s most important plat family (Keller and Feuillet, 2000). A well-
managed grass is globally recognized for their high biodiversity and for their social and cultural
values. These grasses could supply additional non-agricultural services, such as the water supply
and for flow regulation, carbon storage, erosion control, climate mitigation, pollination, and
cultural ecosystem services (Bengtsson et al., 2019). They also support an array of native
herbivores; they are usually affected for the energy consumption of this organisms. Different
species are usually capable of withstanding moderate grazing, but the frequent overgrazing causes
Grass is first and foremost a sustainable plant because it can store so much carbon in the
soil, thus, it can help reduce the CO2 levels in the atmosphere. The blending of different types and
varieties has led to a varied plant population that can stand up to all kinds of extremes (Barenburg,
2023). A plant species’ contaminant tolerance and growth potential are directly related to its
biochemistry and physiology (Huang et al., 2004). Thus, grasses are not only constituting staple
food crops but are grown as fodder for livestock, grown in ornamental landscape, soil conservation
This study which focuses on the inventory of grass species that could be a help in the future
research in effective environmental management. Although grasses are not deemed of great
importance as they may seem too common for people, but they are naturally a part of an ecosystem.
This inventory assessment on grasses can provide information that may be used to convince people
This study aims to conduct and document different grass species in the Visayas State
University – Alangalang Campus, Alangalang, Leyte. Specifically, this study ought to;
2. Determine the species diversity of grasses in terms of richness, percent cover, percent
frequency and coefficient similarities indices (Sorenson’s coefficient and Jaccard’s Coefficient
similarity Index).
This study is limited only to the inventory of grass species on the campus of Visayas State
University, Alangalang, Leyte. This inventory is sought to determine the diversity, evenness, and
the similarities of species among sites – woodland, agricultural land, and grassland. Furthermore,
grasses were the only species considered in recording the data, and the study delimits on the
inventory of other plants species like herbs, shrubs, and ferns are not considered.
CHAPTER 2
methodologies and others. Those that were included in this chapter helps to familiarizing
Grassland’s Biodiversity
The most unique feature of Earth is the existence of life, and the most extraordinary feature
of life is its diversity (Cardinale et al., 2012). Biodiversity refers to all the different kinds of living
organisms within a given area. It considers all types and kinds of organisms existing on the planet
including plants, animals, and other microorganisms (Cadre, 2013). All species are interconnected
and are seemingly dependent on one another. Biodiversity is more than something beautiful to
look at, it is also what provides us with so many of our basic needs (Turnhout et al., 2020). Despite
the fact that species may seem redundant when considering one function under a particular set of
environmental variables, many species are required to maintain several roles at various times and
locations in a changing world (Cardinale et al., 2011). Diversity matters because it has a big effect
on the productivity and stability of natural ecosystems, and on the services, they provide (Roe,
2019). It has been observed that the planet’s biodiversity is steadily decreasing.
Grasses are rarely vining or aquatics; they are perennial or annual, terrestrial, and typically
free-standing. Grasses include all the cereal crops and 75% of cultivated forage crops (Barnes et
al., 1995). Instead of a taproot, lie in many dicotyledons, the root system comprises of fine, fibrous
roots. There are numerous explanations for why there are currently so many genera and species of
grasses, due to adaptations that make grasses ecologically successful, the grass spikelet has
apparently been a competent means of protecting the flower, developing the fruit, and dispersing
the seed (Watson, 1992). It has evolved into a myriad of forms y addition, loss, and modification
of parts. The seven families that make up the grass group within Poales are Poaceae, Restionaceae,
2008). Grasses occur in almost every habitat around the world from the equator to the polar
regions, sea level to mountain heights and aquatic to desert environments (Wheeler et al., 1982).
Grassland make up one of the planet’s predominant biomes and can cover up to 40% of its
land surface (Peterman and Buzhdygan, 2021). In terms of species variety, it may even dwarf
tropical rainforests as one of the planet’s most species-rich environments. Grasslands may support
Importance of Grasses
Ecological Importance
Grass is first and foremost a sustainable plant. Because it can store so much carbon in the
soil, it helps reduce the CO2 levels in the atmosphere. Grasses have acquired characteristics that
enable them to survive harsh climates, particular soil types, fires, and herbivory. Grasses faces
tougher challenges than in past especially due to climate change and increasing water shortages.
Grass with long roots, however, can easily extract more water from the soil, thus reducing the need
for irrigation (Almussaed et al., 2015). These factors all help to maintain grasslands by preventing
the establishment, survival, expansion, and dominance of woody plants. Grassland ecosystem
A study by Tomaskin and Tomaskinova (2012), have the results that confirms the
environmental functions, the grass swards root system constitutes some biological barrier fro crop
contaminations by heavy metals concentrated in soil. There has been the highest concentration in
Agricultural Importance
Grass species are by far the most significant among plant crops. The first domesticated
plants were wheats (Triticum sp.) and barley (Hordeum vulagare) (Zohary and Hopf, 2000).
Today, the main crops for humans are bread wheat (Triticum aestivum), rice (Oryza sativa), maize
(Zeamays), sorghum (Sorgum bicolor), and pearl millet (Pennisetum glaucum). Although most of
these are currently considered minor crops; many additional grasses were domesticated as cereals.
(Simmonds, 1976). From an agricultural, economic, and ecological standpoint, the grass family is
without a doubt the most significant plant family to humanity (Wheeler et al., 1982).
Economic Importance
Grasses are of many importance economically, for instance, with food, fodder, beverages,
and even with industrial use. Several grass species are grown for ornamental purposes, extract and
process to be used in making various perfumes, medicines, soaps, and many others (Koirala, 2018).
Selection of particular grasses for landscape areas plays a major role in the activation of recreation
function throughout the walking and visiting od some picturesque regions, historic gardens.
Recreation function is an organic and spiritual necessity for human beings to escape from the
artificial environment of the cities (Al-samaraee, 2012). Bermuda grass for example, is a highly
variable, hardy, long-lived perennial grass, and is one of the most used warm-season forages in the
Many studies suggest that species diversity and abiotic factors promote ecosystem
diversity remains controversial (Shu et al., 2022). Climate, topography, and human activities are
known to influence plant diversity. Human civilization’s expansion has a specific impact on
grassland biomes, which are defined by large areas of land where grasses are the primary form of
plant life. Human expansion into these areas are frequently threatens the grazing land for many
species of animals, which in turn provide a food source for larger predators (Carr, 2018). Due to
human population growth e.g. human activity-induced environmental changes and natural
environmental changes causing the agricultural and industrial production to extensively increasing
(Moloi et al., 2021). Almost half of all in temperate grassland and for about 16% of tropical
grassland have been converted to agricultural or industrial uses (Walters, 2019). The uniqueness
of grassland from most other biomes wherein grasslands are relatively simple in structure but rich
in number of species. However, most areas of the prairie have experienced serious declines in
biodiversity. Grasses have been especially impacted by decades of human induced alterations to
the prairies that have allowed for the invasion of non-native species. Today, non-native grasses
account for 13 - 30 percent of prairie grass species. Loss of diversity continues as forested patches,
tilled soil, and woody corridors increase on the prairie landscape. Grassland ecosystems worldwide
have been threatened by woody encroachment, which is a result of overgrazing, fire suppression,
grasslands has reached ∼37% of grassland area globally, causing gradual losses of grassland
structure and functions. With this, the long-term intensive human activities caused vegetation
degradation, species loss and a decline in plant richness (Hussein et al., 2021).
Conservation of Grasses
Much of Earth’s grassland has been lost to agricultural development, threatening wildlife
but there are many emerging solutions in conserving grasses (Wolters, 2019). These plantlets can
be used in several ways to fulfil conservation goals including the repopulation of regions with
declining densities of threatened species that hold cultural meaning. However, the potential of in
vitro technologies in the conservation of plants that hold cultural meaning is understudied (Kulak,
et al., 2022). Due to excess degradation in the environment, grasses play a vital role in conservation
sequestration, resistance of abiotic stress, soil conservation, nutritional security for livestock with
higher fodder value, and medicinal and aromatic property (Singh et al., 2022). The current levels
of grassland protection are not sufficient to maintain biodiversity and functioning of the grassland
biome. Temperate grasslands are, in fact, the least protected habitat in the world, with only ∼4%
of the biome under protected status (Petermann & Buzhdygan, 2021). Fodders and grasses can be
preserved either as hay (dried fodder) or as silage (wet fodder), depending on the weather
conditions and the available resources (Singh et al., 1995). Most especially, these grasses have
been directly impacted by decades of human induced alterations to the prairies that have allowed
for the invasion of non-native species. Certainly, the grassland biome is much different today than
before extensive alteration. But with continued persistence and action by dedicated conservation
METHODOLOGY
Study Site
The municipality of Alangalang, Leyte is landlocked town with an area of 151 km2. It is
bounded on the North Barugo and San Miguel, on the South by Pastrana, on the Northeast by
Tacloban City, on the East by Santa Fe, and on the west by Jaro. The study will be conducted on
Visayas State University – Alangalang Campus that located on Barangay Binongtoan, Alangalang,
Leyte. It is approximately at north 110134.51” and east of 124049’43” with an elevation of 43.21
meter (141.78 ft) above sea level. Consequently, three (3) sampling sites will be selected through
a topography and existing vegetation in the area and labeled as sampling location 1, 2, and 3.
Sampling location 1 is randomly selected, then followed by subsequent sampling locations 2 and
3 guided by 5-meter interval. Each sampling locations display different types of soil
(Sanglahi,2022).
For establishment of transect lines, two (2) bamboo stands and a 23-meter transect line was
measured is utilizing by a tape measure. The bamboo stands serve as the indicators of the start and
end points of the transect lines, respectively. On the other hand, 0.5-meter × 0.5-meter quadrat grid
was used for data sampling. To measure the distance or interval between each quadrat along the
transect lines there is a use of meter tape to measure. Thus, the same equipment is used to measure
the interval between the sampling locations. (The Quadrant Cut Method to Measure Grass, 2023).
sample of grass species and will be placed in plastic containers, labeled, and recorded. Local names
were gathered through interview with residents in the locality. Identification of the grass species
using taxonomic references like “Guide to Grassland Plants” by Santos (1983), and “Guide to
Philippine Flora and Fauna” (Vol. IV). Photographs of the grasses are taken to facilitate
Data Analysis
The richness, percent cover and the similarity index of the grass species was derived.
ecological community.
presence in both compared paired groups rather than their mismatches of species occurrences. The
index is also termed as the Dice Similarity Index. SCSI is use for the pairwise comparison of
2Sc
𝑆𝐶𝑆𝐼 = 𝑥100
Sa + Sb + 2Sc
Where:
Jaccard’s coefficient similarity index (JCSI), is another form of similarity index for binary
data which has an equal relevance to SCSI, however, it is use to have a clear state of the presence
and absence of species between different paired forest areas. It is easily computed as
Sc
𝐽𝐶𝑆𝐼 = 𝑥 = 𝑥100
Sa + Sb + Sc
Where:
This chapter presents the results and discussion of the data gathered on the assessment of
grass species composition, richness, percent cover (%), and the similarity index from three areas
such as Site 1 (woodland), site 2 (cropland), and site 3 (grassland) in Visayas State University
Alangalang Campus.
A total of eight (8) species belonging to three (3) families were identified and counted in
the different land-use types in Visayas State University, Alangalang Campus. As shown in table
1, the grass species recorded and identified have been dominated by the Poaceae family having six
(6) species all over which are composed of Chrysopogon aciculate, Cynodon nlemfuensis,
scrobiculatom. On the other hand, only one species each has been recorded for both Cyperaceae
and Fabaceae family which are Cyperus rotundus and Minosa pudica, respectively.
Table 1. List of Grass Species identified in Visayas State University – Alangalang Campus
There were four (3) grass species found in woodland inside the campus of Visayas State
Paspalum scrobiculatum, belonging to family Poaceae. While there are five (4) grass species found
in cropland namely; Cyperus rotundus, Digitaria sanguinalis, Mimosa pudica, and Paspalum
conjugatom, belonging to family Cyperaceae, Fabaceae, and Poaceae respectively. In the grassland
site, a total of four (4) grass species were found and recorded, namely; Cyperus rotundus,
Cyperaceae and Poaceae respectively. Among the identified grass species, only Paspalum
conjugatom appeared on all thrre (3) sites. The species found in all sites probably are adaptive in
different land area they grow and are able to tolerate certain conditions (delos Angeles, 2020).
According to a study by Cayssials and Rodriguez (2013) grass species are more favor in
growing in open habitats, as light is a basic requirement for grass growth. Warm-season grasses
are are generally more heat and drought tolerant than cool-season grasses, but they are less tolerant
on shade. Although shade can increase herbaceous biomass productivity under individual tree
crowns (Belsky, 1994) it can also lead to the exclusion of warm-season grass species (Parr et al.,
2014). Roots on shaded grass grow shallow, and the growth slows. Paspalum conjugatom is one
example of warm season grasses, to which it is less tolerant to shade (Martin et al., 2017).
Table 2. Distribution of Grass Species across the sampling site in different land-use types of
Visayas State University, Alangalang Campus (2023)
Table 3 indicates that in site 1, which is the woodland, most plots were covered by
Paspalum conjugatum, with total average of 25.2% from all the plots established. It is then
scrobiculatum only covered about 0.4% from all the plots in site 1. In site 2, which is the cropland,
the grass species Digitaria sanguinalis, covered 45% of all plots established, followed by Mimosa
pudica with 29%, Paspalum conjugatum with 23.4%, and Cyperus rotundus with 2.6% cover. In
site 3, which is the grassland, most plots were covered by Paspalum conjugatom, with a total
average of 83.4% from all the plots established. It is then followed by Chrysopogon aciculatas
with 9.4%, then by Cynodon nlemfuensis with 4.2% and Cyperus rotundus with a percent cover of
3%. The visual estimates were made with the aid of 0.5m x 0.5m quadrat that were subdivided into
25 small squares marked off within the quadrat frame. Each small square ‘filled’ by a species was
counted as 4% cover.
Table 3. Percent cover by Grass Species in all Land-use Types of Visayas State University,
Alangalang, Campus (2023).
Plot (% Cover)
Site Species Average (% Cover)
1 2 3 4 5
1 (WL) Microstegium viminuem 3 2 0 0 0 1
Paspalum conjugatum 17 22 28 24 35 25.2
Paspalum scrobiculatom 0 0 2 0 0 0.4
2 (CL) Cyperus rotundus 0 0 5 0 8 2.6
Digitaria sanguinalis 0 75 55 80 15 45
Mimosa pudica 90 0 31 15 9 29
Paspalum conjugatum 10 25 9 5 68 23.4
3 (GL) Chrysopogon aciculatas 18 5 14 0 10 9.4
Cynodon nlemfuensis 0 6 0 0 15 4.2
Cyperus rotundus 0 0 5 10 0 3
Paspalum conjugatom 82 89 81 90 75 83.4
As shown in table 4, the Paspalum conjugatum is the most frequent grass species to appear
on all plots, having a percent frequency of 100% from all the sites. In site 1, Microstegium
Paspalum scrobiculatum appearing on 1 out of 5 plots in site 1, resulting on having only 20%
frequency percentage. In site 2, Mimosa pudica and Digitaria sanguinalis appeared on 4 out of 5
plots, resulting on having 80% frequency percentage, followed by Cyperus rotundus appearing on
2 out of 5 plots resulting on having 40% frequency percentage. In site 3, Chrysopogon aciculatas
nlemfuensis and Cyperus rotundus appearing on both 2 out of 5 plots, resulting on 40% frequency
percentage.
In the Philippines, Paspalum conjugatum is widely known and is usually consumed for
livestock as well as landscaping (Garduque et al., 2019) Since Paspalum conjugatum has a 100%
percent frequency based on the study of Rojas-Sandoval (2018) P. conjugatum tolerates high levels
of disturbances and pollution, grows vigoriously under partial shade, spreads easily by its seeds
and stolons, and can withstand acidic and low-nutrient soils. Around Pacific region, it has been
stated that some native forests have become extinct due to this grass pest.
Table 4. Frequency percentage of grass species in different land-use types of Visayas State
University, Alangalang Campus (2023).
Plot (Distribution)
Site Species Average (% Frequency)
1 2 3 4 5
Microstegium viminuem + + - - - 40
1 (WL) Paspalum conjugatum + + + + + 100
Paspalum scrobiculatom - - + - -- 20
2 (CL) Cyperus rotundus - - + - + 40
Digitaria sanguinalis - + + + + 80
Mimosa pudica + - + + + 80
Paspalum conjugatum + + + + + 100
3 (GL) Chrysopogon aciculatas + + + - + 80
Cynodon nlemfuensis - + - - + 40
Cyperus rotundus - - + + - 40
Paspalum conjugatom - - - - - 100
Legend: (+) = Present (-) = Absent
Index (JCSI)
The table below summarizes that paired Location 1 (A) X Location 2 (B) as well as the
paired Location 1 (A) and Location 3 (B) showed lower SCSI and JSCI percentages of less than
50% of having shared species. The former accounts SCSI and JCSI percentages for 28.57% and
16.67%, respectively. On the other hand, the latter accounts SCSI and JSCI percentages for
28.57% and 16.67%, respectively. Nonetheless, paired Location 2 (A) X Location 3 (B) showed
greater SCSI percentages of more than 50%, and a relatively high JSCI percentage. To be exact,
this pair accounts for 50% and 33.33% which represent higher shared similar species.
Table 5. Comparison of Species Composition of the Three Sampling Locations using Sorensen’s
Coefficient
Similarity
Presence of Species
Indices
Paired No. of No. of No. of Shared Total No. of
areas Unique Unique Species of Unique and SCSI JCSI
Species to Species to Location A & Shared (%) (%)
Location A Location B B Species
Location 1
(A) X
2 3 1 6 28.57 16.67
Location 2
(B)
Location 1
(A) X
2 3 1 6 28.57 16.67
Location 3
(B)
Location 2
(A) X
2 2 2 6 50 33.33
Location 3
(B)
CHAPTER V
Summary
Grass, the world's most important crop and it is first and foremost a sustainable plant.
Accordance to its importance grasses also has important ecological functions; it protects against
soil erosion, it absorbs water, it purifies the air we breathe and so much more, however, this
study is intended at assessing the condition of grass species in the advent of anthropogenic
activities. Hence, this is also conducted to assess the presence of grass species composition
significant information and assists in developing innovative and effective strategies on how to
conserve and manage the university’s grass community. This study particularly sought to
determine the species compositions of grass species and to determine the species diversity of
grasses in terms of richness, percent cover, percent frequency and coefficient similarities indices
(Sorenson’s coefficient and Jaccard’s Coefficient similarity Index) of the three sampling locations
The results showed that a total of eight (8) species belonging to three (3) families were
identified and counted in the three sampling locations – Poaceae, Fabaceae, and Cyperaceae.
Herewith, of all the grass species encountered, only Paspalum conjugatom appeared on all
sampling locations while the least species can be seen among all sampling locations are Cyperus
The grass distribution among all sampling locations found out that there were four (3) grass
in cropland namely; Cyperus rotundus, Digitaria sanguinalis, Mimosa pudica, and Paspalum
conjugatom, belonging to family Cyperaceae, Fabaceae, and Poaceae respectively. In the grassland
site, a total of four (4) grass species were found and recorded, namely; Cyperus rotundus,
The percent cover in each sampling loactions indicates that in site 1 (woodland), from the
plots that being established most of it were covered by Paspalum conjugatum with total average
of 25.2%, followed by Microstegium vimineum, with a percent cover of 1% while the Paspalum
scrobiculatum only covered about 0.4%. In site 2 (cropland), the grass species Digitaria
sanguinalis, covered 45% of all plots established, followed by Mimosa pudica with 29%,
Paspalum conjugatum with 23.4%, and Cyperus rotundus with 2.6% cover. In site 3 (grassland),
most plots were covered by Paspalum conjugatom, with a total average of 83.4% then followed
by Chrysopogon aciculatas with 9.4%, Cynodon nlemfuensis with 4.2% and Cyperus rotundus
The results in the percentage frequency shows that from all the sites the grass species,
Paspalum conjugatum is the most frequent grass species to appear on all plots, having a percent
of 5 plots having only 20% frequency percentage. In site 2 (cropland), Mimosa pudica and
Digitaria sanguinalis appeared on 4 out of 5 plots, resulting on having 80% frequency percentage,
followed by Cyperus rotundus appearing on 2 out of 5 plots resulting on having 40% frequency
Lastly, the result showed that paired Location 1 (A) X Location 2 (B) as well as the paired
Location 1 (A) and Location 3 (B) showed lower SCSI and JSCI percentages of less than 50% of
having shared species. On the other hand, paired Location 2 (A) X Location 3 (B) showed greater
SCSI percentages of more than 50%, and a relatively high JSCI percentage.
Conclusions
Based on the outcome of the study, the researchers found that there are eight (8) grass
species belonging to three (3) families in the different land-use types in Visayas State University,
Alangalang Campus. Among the grass family, the Poaceae have the higher count of grass species
while the family that have a least grass species would be Cyperaceae and Fabaceae family.
Meanwhile, on the grass species distribution there were a presence of Paspalum conjugatom on
all sites.
Furthermore, the researchers concluded that Visayas State University, Alangalang Campus
has a high species richness as revealed by the values of Percent Cover and Percentage Frequency.
Additionally, the similarity indices revealed that there two out of three paired locations that have
a lower SCSI and JCSI percentages of less than 50% of having shared species. The researchers
conclude that having a less than 50% similarities among all sites indicate that there has a high
Based on the findings and the results that were obtained, the future researchers are suggests
to study other parameters that might be considered for the improvement of the study:
b.) Other types of plant species should also be inventoried aside from grasses only;
c.) Aside from a simple inventoried, examine or assess also the environmental factors that
d.) Include also the abundance and other diversity indices; and
DOCUMENTATION
DATA GATHERED
Researched by:
Apostol, Kylamarie T.
Cadorna, Lailane B.
Caones, Abadiano A.
June 2023