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Robert M.

Timm
Superintendent and Extension
Wildlife Specialist
HOUSE MICE
Hopland Research & Extension Center
University of California
Hopland, California 95449

Fig. 1. House mouse, Mus musculus

Damage Prevention and Repellents Fumigants


Control Methods Ro-pel® Practical use is limited to structures,
containers, and commodities; for
Moth flakes (naphthalene) not
Exclusion use only by trained personnel.
specifically registered, but may be
Seal all openings larger than 1/4 inch of some value. Trapping
(0.6 cm) wide. Toxicants Snap traps.
Habitat Modification
Anticoagulant rodenticides (slow- Live traps (Sherman-type, Ketch-All®,
Good sanitation practices reduce acting chronic-type toxicants). Tin Cat®, and others).
sources of food, water, and shelter. Brodifacoum (Talon®).
Glue boards.
Bromadiolone (Maki®, Contrac®).
Store foodstuffs in rodent-proof Other Methods
Chlorophacinone (RoZol®).
structures or containers.
Diphacinone (Ditrac®).
Predators: dogs and cats are of limited
Control weeds and remove debris Pindone (Pival®, Pivalyn®).
value in some situations.
from around structures. Warfarin (Final® and others).
Frightening Toxicants other than anticoagulants
(may be acute or chronic poisons).
Ultrasonic devices have not been
Bromethalin (Assault®, Vengeance®).
proven to control mice.
Cholecalciferol (Quintox®).
Zinc phosphide (Ridall Zinc®, ZP®).

PREVENTION AND CONTROL OF WILDLIFE DAMAGE — 1994


Cooperative Extension Division
Institute of Agriculture and Natural Resources
University of Nebraska - Lincoln
United States Department of Agriculture
Animal and Plant Health Inspection Service
Animal Damage Control
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Great Plains Agricultural Council
Wildlife Committee
Identification move into structures in search of loosely woven together. They are usu-
shelter and food. ally 4 to 6 inches (10.2 to 15.2 cm) in
The house mouse (Mus musculus, Fig. diameter.
1) is a small, slender rodent that has a Food Habits Litters of 5 or 6 young are born 19 to
slightly pointed nose; small, black, 21 days after mating, although females
somewhat protruding eyes; large, that conceive while still nursing may
House mice eat many types of food
sparsely haired ears; and a nearly have a slightly longer gestation period.
but prefer seeds and grain. They are
hairless tail with obvious scale rings. Mice are born hairless and with their
not hesitant to eat new foods and are
House mice are considered among the eyes closed. They grow rapidly, and
considered “nibblers,” sampling many
most troublesome and economically after 2 weeks they are covered with
kinds of items that may exist in their
important rodents in the United States. hair and their eyes and ears are open.
environment. Foods high in fat, pro-
Adult house mice weigh about 2/5 to tein, or sugar may be preferred even They begin to make short excursions
4/5 ounce (11 to 22 grams). They are when grain and seed are present. Such from the nest and eat solid food at 3
generally grayish brown with a gray or items include bacon, chocolate candies, weeks. Weaning soon follows, and
buff belly. Similar mice include the butter, and nutmeats. mice are sexually mature at 6 to 10
white-footed mice and jumping mice weeks of age.
Unlike Norway and roof rats, house
(which have a white belly), and Mice may breed year-round, but when
mice can survive with little or no free
harvest mice (which have grooved living outdoors, they breed mostly in
water, although they readily drink
upper incisor teeth). For more details spring and fall. A female may have 5
water when it is available. They obtain
on species identification, see a field to 10 litters per year. Mouse popula-
their water from the food they eat. An
guide such as that by Burt and tions can therefore grow rapidly under
absence of liquid water or food of
Grossenheider (1976). good conditions, although breeding
adequate moisture content in their
Native to central Asia, this species environment may reduce their breed- and survival of young decline mark-
arrived in North America with settlers ing potential. edly when population densities
from Europe and from other points of become high.
origin. A very adaptable species, the House mice have physical capabilities
house mouse often lives in close General Biology,
that enable them to gain entry to struc-
association with humans and therefore Reproduction, and tures by gnawing, climbing, jumping,
is termed one of the “commensal” Behavior and swimming. For more detailed
rodents along with Norway and roof information on their physical abilities
rats. House mice are much more House mice are mainly nocturnal, and the resulting need to design
common in residences and commercial although at some locations consider- rodent-proof structures, see the chap-
structures than are rats. Brooks (1973) able daytime activity may be seen. See- ter Rodent-Proof Construction and
regards them to be the most common ing mice during daylight hours does Exclusion Methods.
mammal in cities, next to humans. not necessarily mean that a high popu-
Studies indicate that during its daily
lation is present, although this is usu-
activities, a mouse normally travels an
Range ally true for rats.
area averaging 10 to 30 feet (3 m to
Mice have poor eyesight, relying on 9 m) in diameter. Mice seldom travel
Following their arrival on colonists’ their hearing and their excellent senses farther than this to obtain food or
ships, house mice spread across of smell, taste, and touch. They are water. Because of their limited move-
North America and are now found in considered color-blind; therefore, for ment and feeding behavior, both of
every state, including coastal areas of safety reasons, baits can be dyed dis- which differ from those of commensal
Alaska, and in the southern parts of tinctive colors without causing avoid- rats, they are much more difficult to
Canada. ance by mice, as long as the dye does control in some situations.
not have an objectionable taste or
Mice constantly explore and learn
odor.
Habitat about their environment, memorizing
House mice may burrow into the the locations of pathways, obstacles,
House mice live in and around homes, ground in fields or around structures food and water, shelter, and other ele-
farms, commercial establishments, and when other shelter is not readily avail- ments in their domain. They quickly
in open fields and agricultural lands. able. Nesting may occur in the ground detect new objects in their environ-
At times they may be found living far or in any sheltered location. Nests are ment but, unlike rats, do not fear them.
from human settlements, particularly constructed of shredded fibrous mate- Thus, they will almost immediately
where climates are moderate. The rials such as paper, burlap, or other enter bait stations and sample new
onset of cold weather each fall in similar items, and generally have the foods (baits). The degree to which
temperate regions may cause mice to appearance of a “ball” of material mice consume a particular food

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depends on the flavor of the food in packaging materials in warehouses fungal skin disease) in humans. They
addition to its physiological effect. where food and feeds are stored. have also been found to act as reser-
Mice may reject baits simply because Much of this loss is due to contamina- voirs or transmitters of diseases of vet-
they do not taste as good as other tion with droppings and urine, making erinary importance, such as swine dys-
available foods. food unfit for human consumption. entery, a serious bacterial disease of
swine often called “bloody scours.”
If the bait contains poison or some House mice cause structural damage
other substance that produces an ill to buildings by their gnawing and Mouse Sign
effect (but not death) within a few nest-building activities. In livestock
hours, the bait will often become asso- confinement facilities and similar The presence of house mice can be
ciated with the illness. Bait shyness can structures, they may quickly cause determined by a number of signs
persist for weeks or months and may extensive damage to insulation inside described below:
be transferred to nontoxic foods of walls and attics. Such damage also Droppings may be found along run-
similar types. Prebaiting, that is, train- occurs in homes, apartments, offices, ways, in feeding areas, and near shel-
ing mice to feed repeatedly on non- and commercial buildings but usually ter. Differentiating between mouse
toxic bait for a period of days prior to at a slower rate because mouse popu- droppings and those of certain insects
applying the toxicant in the bait, will lations in such structures are smaller. may be difficult. Mouse droppings are
largely prevent sublethal doses and House mice often make homes in large about 1/4 inch (0.6 cm) long, whereas
thus bait shyness. It will also reduce electrical appliances, and here they those of cockroaches are usually 1/8 to
the number of mice left to be bait shy. may chew up wiring as well as insula- 1/4 inch (0.3 to 0.6 cm) long and under
Prebaiting is especially recommended tion, resulting in short circuits which a magnifying glass show distinct longi-
with zinc phosphide baits. All of the create fire hazards or other malfunc- tudinal ridges and squared-off ends. In
other toxic baits currently registered tions that are expensive to repair. Mice comparison, droppings of bats contain
for house mice are chronic or slow- may also damage stored items in insect fragments and are more easily
acting. Because of this slow action, the attics, basements, garages, or muse- crushed between the fingers.
mice’s subsequent illness is not associ- ums. Damaged family heirlooms,
ated with the bait even if a sublethal paintings, books, documents, and Tracks, including footprints or tail
dose is consumed; thus, bait shyness other such items may be impossible to marks, may be seen on dusty surfaces
does not usually occur. These baits, in replace. or in mud (Fig. 2). A tracking patch
effect, serve as their own prebait. made of flour, rolled smooth with a
Among the diseases mice or their para- cylindrical object, can be placed in
sites may transmit to humans are pathways overnight to determine if
Damage and Damage salmonellosis (food poisoning), rickett- rodents are present.
sialpox, and lymphocytic choriomen-
Identification Urine, both wet and dry, will fluo-
ingitis. Mice may also carry lepto-
spirosis, ratbite fever, tapeworms, and resce under ultraviolet light, although
When house mice live in or around so will some other materials. Urine
organisms that can cause ringworm (a
structures, they almost always cause
some degree of economic damage. In
homes and commercial buildings, they
may feed on various stored food items
or pet foods. In addition, they usually a
contaminate foodstuffs with their
urine, droppings, and hair. On farms,
they may cause damage to feed stor-
age structures and feed transporting
equipment. A single mouse eats only
about 3 grams of food per day (8
pounds [3.6 kg] per year) but destroys
considerably more food than it con-
sumes because of its habit of nibbling
on many foods and discarding par-
tially eaten items.
b
House mice living in fields may dig up
and feed on newly planted grain, or
may cause some damage to crops
before harvest. But losses in stored
foods are considerably greater. Mice
commonly damage containers and Fig. 2. Tracks left in dust by (a) Norway rat and (b) house mouse.

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Legal Status
House mice are not protected by law.
They may be controlled using any pes-
ticide registered by federal or state au-
thorities for this purpose, or they may
be controlled by use of mechanical
methods such as traps.

Damage Prevention and


Control Methods
Fig. 3. Rub marks along beams, rafters, or other travel routes give evidence of rodent activity. Mouse Effective prevention and control of
rub marks can be distinguished from those of rats by their smaller size. house mouse damage involves three
aspects: rodent-proof construction,
stains may occur along travelways or Odors may indicate the presence of sanitation, and population reduction
in feeding areas. house mice. A characteristic musky by means of traps, toxicants, or fumi-
odor is a positive indication that house gants. The first two are useful as pre-
Smudge marks (rub marks) may ventive measures, but when a house
mice are present, and this odor can be
occur on beams, rafters, pipes, walls, mouse infestation already exists, some
used to differentiate their presence
and other parts of structures. They are form of population reduction is almost
from that of rats.
the result of oil and dirt rubbing off always necessary. A flow chart outlin-
mice’s fur along frequently traveled Estimating Mouse Numbers ing steps in controlling house mice is
routes (Fig. 3). They may be less appar- found in figure 4.
ent than rub marks left by rats. Mouse sign and visual sightings are of
limited value in accurately estimating Control of house mice differs in impor-
Gnawing may be visible on doors, tant ways from the control of Norway
mouse numbers, but they are the sim-
ledges, in corners, in wall material, on plest and often the only practical or roof rats. Mice are smaller and
stored materials, or on other surfaces method available. Search premises therefore can enter narrower openings,
wherever mice are present. Fresh accu- thoroughly when looking for mice. In making rodent-proofing more difficult.
mulations of wood shavings, insula- structures, searches should include They have limited areas of movement
tion, and other gnawed material attics, basements, around foundations, (home range) and require little or no
indicate active infestations. Size of crawl spaces, and behind and under free water. While having a reproduc-
entry holes (often 1 1/2 inches [3.8 cm] stored materials. tive capability that is higher than that
in diameter or less for mice, 2 inches [5 of rats, house mice are usually less sen-
cm] or larger for rat) or tooth marks One method to detect the presence of sitive (often far less sensitive) to many
can be used to distinguish rat gnawing mice is to make nontoxic tracking-dust rodenticides. Persons who do not take
from mouse gnawing. Mice keep their patches of flour or talc at 20- to 30-foot these differences into account when
paired incisor teeth, which grow con- (6- to 9-m) intervals throughout a attempting house mouse control may
tinuously, worn down by gnawing on structure. The number of patches expect poor results.
hard surfaces and by working them showing tracks after 24 hours, and the
against each other. abundance of tracks in each patch, After rats are controlled at a given
indicate the size of the population. location, house mice may increase in
Sounds such as gnawing, climbing in numbers by moving in from elsewhere
Because house mice, unlike rats, do
walls, running across the upper sur- not travel far from their nests or shel- or by reproduction. This may be
face of ceilings, and squeaks are com- ter, the percentage of patches showing expected because habitats suitable for
mon where mice are present. tracks is a good indicator of the rela- rats are usually even more suitable for
Visual sightings of mice may be tive size and distribution of the mouse mice. One should anticipate that fol-
possible during daylight hours, and population. lowing rat control, the potential for
mice also can be seen after dark with house mouse problems may increase,
Snap trapping is also an excellent way and control measures should be taken
the aid of a flashlight or spotlight. to determine the presence of mice. A before mouse numbers reach high
Nests frequently are found when relative index of mouse abundance can levels.
cleaning garages, closets, attics, base- be calculated from the number of mice
ments, and outbuildings where mice trapped for a certain number of traps
are present. They consist of fine, shred- set during 1 or more nights (for ex-
ded fibrous materials. ample, 35 mice caught per 100 trap
nights).

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Can mice’s food source
Exclusion
be removed or limited?
Physical barriers can prevent mice
YES NO from gaining entry to structures where
Remove or
food and shelter are available. Rodent-
limit food proofing is an important and often
source neglected aspect of rodent control. It is
Can mice’s shelter be
removed or limited? a relatively permanent form of control
YES NO that can prevent damage from occur-
ring.
Reduce or
limit To exclude mice, seal all holes and
shelter.
openings larger than 1/4 inch (0.6 cm)
NO across. Rodent-proofing should be
Are mice numerous?
done with heavy materials that will
YES resist rodent gnawing. These include
concrete mortar, galvanized sheet
Is a quick reduction in metal, and heavy-gauge hardware
mouse numbers
needed? cloth. For more detailed information
YES NO on techniques of mouse-proof con-
struction, see the chapter Rodent-
Proof Construction and Exclusion
Will the presence of dead Methods.
mice cause an odor or
sanitation problem?
NO YES

Are the mice within a


building or structure that
can safely be fumigated?

NO YES

Contact a
pest control Use
operator for traps
fumigation.
Can a single-dose
toxicant be used Glue boards
safely? Use single- (if not apt to
dose toxicant entrap
NO YES (assistance of
Snap children, pets,
a pest control or wildlife)
operator may traps
be required).

After using a single-


dose toxicant, do some
mice remain? Multiple-
Can an anticoagulant NO capture
YES
rodenticide be used traps
safely? Use
NO YES anticoagulant After using an anti-
rodenticide coagulant, do some
mice remain?

NO YES

Fig. 4. A flow chart of steps in controlling house mouse populations. Additional factors, such as the cost of particular
control methods, must be taken into account when planning a control program (see text).

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12" white band

Fig. 5. A 12-inch (30.5-cm) white painted band makes inspection for rodent sign easier and reminds personnel not to store commodities too close to walls.

Habitat Modification consumed will support at least a few heavy gravel placed adjacent to build-
mice. For this reason, a constant watch ing foundations or other structures
Sanitation, which includes good
must be kept for mice that may invade will reduce rodent burrowing at these
housekeeping practices and proper
the premises. locations. In any event, keep the
storage and handling of food materi-
perimeter of buildings and other struc-
als, feed, and garbage, is often stressed Where possible, store bulk foods in
tures clean of weeds and debris
as a method of rodent control. Unfor- rodent-proof containers or rooms.
(including stacked lumber, firewood,
tunately, even the best sanitation will Stack sacked or boxed foods in orderly
and other stored materials) to discour-
not eliminate house mice. It will, how- rows on pallets in a way that allows
age rodent activity and to allow easier
ever, aid in control by permitting for thorough inspection for evidence of
detection of rodent sign.
easier detection of mouse sign, increas- mice. In such storage areas, keep
ing effectiveness of traps and baits by stored materials away from walls. A Frightening
reducing competing food items, and 12-inch (30.5-cm) white band painted
by preventing mice from flourishing on the floor next to the wall serves as a Mice are somewhat wary animals and
and reaching high populations. reminder to keep items away from can be frightened by unfamiliar
walls. It also will allow you to detect sounds or sounds coming from new
Although house mice are less depen-
rodent droppings or other sign more locations. Most rodents, however, can
dent upon humans for their existence
easily (Fig. 5). Sweep floors frequently quickly become accustomed to new
than are Norway rats, they are much
to permit ready detection of fresh sounds heard repeatedly.
more adaptable to living with people.
droppings. For years, devices that produce ultra-
They require very little space and only
small amounts of food. Mice have been When storing foods or feed on pallets, sonic sound that is claimed to control
known to inhabit buildings even keep in mind that mice can jump up rodents have come and gone on the
before construction has been complete, more than 12 inches (30.5 cm) from a market. There is little evidence to sug-
living off the crumbs and scraps of flat surface. They are also good climb- gest that rodents’ responses to nonspe-
worker’s lunches. In offices, mice may ers and can walk up surfaces such as cific, high-frequency sound is any
live behind cabinets or furniture and wood or concrete (unless the surfaces different from their response to sound
feed on scraps or crumbs from lunches have a slick finish). Mice can live for within the range of human hearing.
and snacks and on cookies or candy considerable periods of time within a What is known about rodents and
bars kept in desks. In homes, they may pallet of feed without coming down to sound?
find ample food in kitchens, and in the the floor.
garage they will eat sacked or spilled —Unusually loud, novel, or ultrasonic
Regular removal of debris and control sounds, which rodents can hear, will
pet food, grass seed, or insects such as
of weeds from around structures will frighten them and may cause tempo-
cockroaches. Thus, no matter how
reduce the amount of shelter available rary avoidance lasting from a few min-
good the sanitation, most buildings in
to rodents. In some instances, a strip of utes to a few weeks.
which food is stored, prepared, or

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What is known about ultrasonic commercial devices are unsubstan- toxicants and multiple-dose (chronic)
sound? tiated by scientific research. Since com- toxicants. However, the complexity in
mercial ultrasonic devices are often mode of action of newer rodenticides
—It is very directional and does not
expensive and of questionable effec- makes these classifications outdated. A
travel around corners well; thus,
tiveness, they cannot be recommended classification into two groups, the first
sound shadows or voids are created.
as a solution to rodent problems. group including all anticoagulants and
—Ultrasound does not travel very far. the second group all other compounds
It loses its intensity rapidly as it leaves Repellents (“non-anticoagulants”), is currently
the source. Rodents find some types of tastes and more useful.
—Ultrasound has not been shown to odors objectionable, but chemical Anticoagulants (slow-acting,
drive established rodents out of build- repellents are seldom a practical solu- chronic toxicants). House mice are
ings or areas, nor has it been proven to tion to mouse infestations. Substances susceptible to all of the various anti-
cause above-normal mortality in their such as moth balls (naphthalene) or coagulant rodenticides (Table 1), but
populations. While it is possible to household ammonia, in sufficient con- they are generally less sensitive (often
cause convulsions or permanent physi- centration, may have at least tempo- far less sensitive) to the active ingre-
ological damage to rodents with ultra- rary effects in keeping mice out of dients than are Norway or roof rats. It
sound, the intensity of such sounds certain enclosed areas. These are not usually requires a few more feedings
must be so great that damage to specifically registered by the EPA as to produce death with the first-genera-
humans or domestic animals would mouse repellents, however. tion anticoagulants (such as warfarin,
also be likely. Commercial ultrasonic diphacinone, and chlorophacinone)
Ro-pel® is registered for use in repel-
pest control devices do not produce than with the second-generation anti-
ling house mice and other rodents
sound of such intensity. coagulants (such as brodifacoum and
from gnawing on trees, poles, fences,
bromadiolone). All anticoagulants pro-
Recent tests of commercial ultrasonic shrubs, garbage, and other objects.
vide good to excellent house mouse
devices have indicated that rodents Little information is currently available
control when prepared in acceptable
may be repelled from the immediate on its effectiveness against house mice.
baits. A new second-generation anti-
area of the ultrasound for a few days, Other solutions to rodent problems, coagulant, difethialone, is presently
but then will return and resume nor- including rodent-proof construction being developed and EPA registration
mal activities. Other tests have shown and methods of population reduction, is anticipated in the near future. The
the degree of repellency to depend are usually more permanent and cost- characteristics of the various antico-
upon the particular ultrasonic frequen- effective than the use of repellents. agulant rodenticides are described
cies used, their intensity, and the pre-
further under Anticoagulants in the
existing condition of the rodent Toxicants Pesticides section, and in the chapter
infestation. Ultrasonic sound has very
Rodenticides were formerly classified Norway Rats.
limited usefulness in rodent control.
The advertising claims for many into two groups, single-dose (acute)

Table 1. Anticoagulants used for house mouse control in the United States.
Usual types of formulations Percent
Common name and Food Tracking active ingredient
typical trade names Chemical name bait Liquid powder used in food bait
Hydroxycoumarins
Warfarin (Final®
and others) 3-(α-acetonylbenzyl)-4-hydroxycoumarin X X 0.025
Brodifacoum 3-[3(4'-bromo[1,1’biphenyl]-4-yl)-1,2,3,4-tetrahydro-
(Talon®)* 1-naphthalenyl]-4-hydroxy-2H-1-benzopyran-2-one X 0.005
Bromadiolone (Maki®, 3-[3-(4'-bromo[1,1’biphenyl]-4-yl)-3-hydroxy-1-
Contrac®)* phenylpropyl]-4-hydroxy-2H-1-benzopyran-2-one X 0.005
Difethialone* [(bromo-4'-[biphenyl-1-1']-yl-4) 3-tetrahydro-1,2,3,4-napthyl-1] X 0.0025
3-hydroxy-4, 2H-1-benzo-thiopyran-2-one
Indandiones
Chlorophacinone
(RoZol®) 2-[(p-chlorophenyl)phenylacetyl]-1,3-indandione X X 0.005
Diphacinone (Ditrac®) 2-diphenylacetyl-1,3-indandione X X 0.005
Pindone (Pival®, Pivalyn®) 2-pivalyl-1,3-indandione X X 0.025
* Second-generation anticoagulants especially useful for the control of warfarin-resistant rats and mice.

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Because of their similarity in mode of individuals are less sensitive to — Improperly placed bait stations.
action, all anticoagulant baits are used anticoagulants than others. Where Other foods are more convenient to
in a similar fashion. Label directions anticoagulants have been used over the mice.
commonly instruct the user to “main- long periods of time at a particular
— Abundance of other food choices.
tain a continuous supply of bait for 15 location, there is an increased potential
days or until feeding ceases,” thus for the existence of a population that is — Tainted bait: the bait has become
ensuring that the entire mouse popula- somewhat resistant to the lethal effects moldy, rancid, insect-infested, or
tion has ample opportunity to ingest a of the baits. Such resistant populations contaminated with other material
lethal dose of the bait. Anticoagulants of house mice have been identified at a that reduces acceptance. Discard
have the same effect on nearly all number of locations throughout the old bait periodically, and replace it
warm-blooded animals, but the sensi- United States. Although not common, with fresh.
tivity to these toxicants varies among resistance may be underestimated
Occasionally, mice accept bait well and
species. If misused, anticoagulant because relatively few resistance
an initial population reduction is suc-
rodenticides can be lethal to nontarget studies have been conducted on house
cessful. Then bait acceptance appears
animals such as dogs, pigs, and cats. mice. Nevertheless, resistance is of lit-
to stop although some mice remain. In
Additionally, residues of anticoagu- tle consequence in the control of house
such instances, it is likely that the re-
lants which are present in the bodies of mice with the newer rodenticides
maining mice never accepted the bait,
dead or dying rodents can cause toxic available. When anticoagulant resis-
either because of its formulation or
effects to scavengers and predators. In tance to the first-generation anticoagu-
placement. The best strategy is to
general, however, the secondary poi- lants is known or suspected, use of
switch to a different bait formulation,
soning hazard from anticoagulants is these compounds should be avoided
place baits at different locations, and/
relatively low. in favor of the second-generation
or use other control methods such as
anticoagulants or one of the non-
Brodifacoum and bromadiolone baits, traps.
anticoagulant products.
because of their potential to be lethal in
Other Rodenticides. The older
a single feeding, can be more effective Anticoagulant Bait Failure.
rodenticides, formerly referred to as
than the other anticoagulants in certain Resistance is only one (and perhaps
acute toxicants, such as arsenic triox-
situations. Chlorophacinone (RoZol®) the least likely) reason for failure in the
ide, phosphorus, strychnine, and Com-
and diphacinone (Ditrac®) are similar control of mice with anticoagulant
pound 1080, are no longer registered
to each other in potency and are more baits. Control with baits that are highly
for house mice. Newer rodenticides
toxic than the anticoagulant com- accepted may fail for one or more of
are much more effective and have
pounds developed earlier. Thus, they the following reasons:
resulted in the phasing out of these
are formulated at lower concentra- — Too short a period of bait exposure. older materials over the last 20 years.
tions. Chlorophacinone and diphaci-
none may kill some mice in a single — Insufficient bait and insufficient At present, three non-anticoagulant
feeding, but multiple feedings are replenishment of bait (none remains rodenticides (Table 2) are registered by
needed to give adequate control of a from one baiting to the next). EPA against house mice: bromethalin,
mouse population. — Too few bait stations and/or too far cholecalciferol (vitamin D3), and zinc
apart. For mice, stations should be phosphide. All are potentially useful
Pindone (Pival®, Pivalyn®) is also less
within 6 feet (2 m) of one another in for controlling anticoagulant-resistant
potent than chlorophacinone or
areas where mice are active. populations of house mice.
diphacinone, and is similar to warfarin
in effectiveness against house mice. It — Too small a control area, permitting Of these active ingredients, brometha-
has some properties that resist insects mice to move in from untreated lin and cholecalciferol are formulated
and growth of mold in prepared baits. adjacent areas. to serve as chronic rodenticides,
applied so that house mice will have
Warfarin (Final® and other trade — Genetic resistance to the anticoagu-
the opportunity to feed on the baits
names) was the first marketed anti- lant. Although this is unlikely, it
one or more times over the period of
coagulant and is, therefore, the best should be suspected if about the
one to several days. Bait acceptance is
known and most widely used. It is same amount of bait is taken daily
generally good when formulations
effective against house mice, although for several weeks.
appropriate for house mice are select-
some warfarin contains small quanti-
Reasons for failure to achieve control ed. Zinc phosphide differs from the
ties of contaminants that apparently
with anticoagulant baits that are other two compounds in that prebait-
can reduce bait acceptance. This has
poorly accepted: ing (offering mice similar but nontoxic
been resolved with the development of
— Poor bait choice, or bait formulated bait prior to applying the zinc phos-
encapsulated warfarin.
improperly. Other foods are more phide-treated bait) is recommended to
Anticoagulant Resistance. Within attractive to the mice. increase bait acceptance. Zinc phos-
any population of house mice, some phide baits are not designed to be left

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Table 2. Other (non-anticoagulant) rodenticides used to control house mice in the United States.
Acute oral Percent
LD50 for Time active
mice, to ingredient Relative
Common Name Chemical Name mg/kg death Odor Taste in food bait hazard Mode of Action
Bromethalin N-methyl-2,4-dinitro-N- 5.25-8.13 2-4 days None Slight 0.01 Moderate Central nervous
(Assault®, (2,4,6-tribromophenyl)- system depression
Vengeance®) 6-(trifluoromethyl) and paralysis
benzenamine
Cholecalciferol 9,10-Seocholesta-5,7,10 42.5 3-4 days Slight None 0.075 Low to Mobilizes calcium
(vitamin D3, (19)-trein-3 betaol moderate resulting in death
Quintox®) from hypercalcemia
Zinc phosphide Zinc phosphide 40 1/2-20 hours Strong Strong 1.0-2.0 Moderate Phosphine gas enters
circulatory system;
heart paralysis,
gastrointestinal and
liver damage

available to mice for more than a few accustomed to eating are often a good available in bulk or packaged in small
days, as continued exposure is likely to choice, particularly if their normal plastic, cellophane, or paper “place
result in bait shyness within the foods are limited or can be made less packs” (Fig. 6). These packets keep bait
population. Be sure to follow label rec- available to them. In past years, many fresh and make it easy to place baits
ommendations on any specific product people involved in house mouse con- into burrows, walls, or other locations.
to achieve best success. trol preferred to mix their own baits so Mice will gnaw into these bags to feed
as to tailor them to the food preference on an acceptable bait. Pelleted baits
Bait Selection and Formulation of a specific mouse population. Today, can more easily be carried by mice to
Oatmeal, ground or rolled wheat, there is a wide selection of ready-to- other locations. Such hoarding of food
rolled barley, ground or rolled milo, use baits which are commercially by mice is not uncommon. It may
and corn have been successfully used available. It is still important, particu- result in amounts of bait being moved
as chief ingredients of toxic baits for larly in moderate- to large-scale mouse to places where it is undetected or dif-
house mice. Grass seed, such as whole control programs, to check for differ- ficult to recover and may, if accessible,
canary grass seed (Phalaris canarienses), ences in bait acceptance among candi- be hazardous to nontarget species. On
is often highly accepted by house mice date baits prior to investing time and the other hand, pelleted bait avoids
and can be very effective as a principal money in a specific bait product. Place some problems common to loose baits
bait ingredient. In general, the fresher about 1/2 ounce (14 g) of each of sev- — settling out of different-sized par-
the bait, the better it will be accepted eral ready-to-use baits about 4 inches ticles during shipment and uneven
by mice. Rodent baits should always (10 cm) apart in several locations mixing of the toxicant. Pellets are eas-
be made from high-quality food mate- where mice are present. Check baits ily manipulated by mice, increasing
rials, and baits should be replaced or the next day to see which ones are the attractiveness of this form of bait.
replenished regularly. preferred.
Anticoagulant baits have also been for-
Food preferences may vary among Ready-to-use baits come in a variety of mulated into wax and extruded blocks
mouse populations and individuals. formulations. Grain-based baits in a (Fig. 7). These are particularly useful
Bait materials similar to foods mice are loose meal or pelleted form are where moisture may cause loose grain

Fig. 6. Various types of place packs containing Fig. 7. Wax and extruded bait blocks are useful in damp locations where loose baits become spoiled
rodenticides are commercially available. quickly.

B-39
baits to spoil. Mice accept paraffin — provide a protected place for Kinds of Bait Stations. Bait sta-
block baits less readily than loose or rodents to feed, allowing them to tions can contain solid baits (food
pelleted grain baits, but acceptance of feel more secure. This is an impor- baits), liquid baits, or both. Bait boxes
extruded bait blocks is high. tant advantage when baiting mice, can be purchased from commercial
which apparently like to spend time suppliers or made at home. Manufac-
Where no water is available, water or
feeding inside such bait boxes; tured bait boxes made of plastic, card-
food items of high moisture content
board, or metal are sold to pest control
are often more readily accepted than — keep other animals (pets, livestock,
companies and to the public (Fig. 9) in
dry baits. Sodium salts of anticoagu- desirable wildlife) and children
sizes for rats or mice. Some farm sup-
lants are available as concentrates to be away from hazardous bait;
ply and agricultural chemical supply
mixed with water, making a liquid bait
— allow placement of bait in locations stores have them in stock or can order
(Fig. 8). Although mice require little or
where it would otherwise be diffi- them. Recent research suggests mice
no water to survive, they will readily
cult because of weather or potential may prefer to feed in cardboard bait
drink it when available. Water baits
hazards to nontarget animals; stations rather than plastic ones.
can be an effective supplement to other
control measures where water is — help prevent the accidental spilling Bait boxes can be built from scrap ma-
scarce. They are particularly useful in of bait; terials, and homemade stations can be
grain storage structures, warehouses, designed to fit individual needs. Make
— allow easy inspection of bait to see
and other such locations. Rodents are them out of sturdy materials so they
if rodents are feeding on it.
more easily able to detect anticoagu- cannot be easily knocked out of place
lants in water baits than in food baits;
therefore, up to 5% sugar is sometimes
added to liquid baits to increase
rodents’ acceptance of the bait solu-
tion. Since water is attractive to most
animals, use water baits in ways that
prevent nontarget animals from drink-
ing them.

Bait Stations
Bait stations (bait boxes) may increase
both the effectiveness and safety of
rodenticides. They came into general
use after the development of the first-
generation anticoagulants, which
require that a continuous supply of
bait be made available to rodents. Bait
stations are useful because they:
— protect bait from moisture and
dust;

Fig. 8. Liquid baits can be placed in fonts or


other similar containers. Fig. 9. Examples of commercially manufactured rodent bait stations.

B-40
or damaged. Where children, pets, or
livestock are present, be careful to
construct the stations so that the bait is
18"
accessible only to rodents. Locks, seals,
or concealed latches are often used to
make bait boxes more tamperproof.
Clearly label all bait boxes or stations
with “Poison” or “Rodent Bait — Do
Not Touch,” or with a similar warning.
Some rodenticides or situations may
require use of tamper-resistance bait
stations. If so, use only bait boxes or
stations which are so designated, and
also be sure to secure them to build-
ings by nailing or gluing them to walls
or floors in a way that will not permit
a person or animal to knock them over
or shake the bait out.
Bait Station Design. Bait stations
should be large enough to allow sev-
eral rodents to feed at once. They can
Fig. 10. A flat board nailed to a wall protects rodent bait from nontarget animals and allows rodents be as simple as a flat board nailed at an
to feed in a sheltered location. The board should be at least 18 inches (45.7 cm) long to keep pets and angle to the bottom of a wall (Fig. 10),
children from reaching the bait. or a length of pipe into which bait can
be placed (Fig. 11). More elaborate sta-
tions are completely enclosed and can
contain liquid as well as solid rodent
baits (Fig. 12). A hinged lid with a
child-proof latch can be used for con-
venience in inspecting permanent
stations.

18"

Fig. 11. Rodent bait station made from a length of pipe. Pipe
diameter can be 2 to 3 inches (5.1 to 7.6 cm) for mice; 3 1/2 to
6 inches (8.9 to 15.2 cm) for rats.

2 1/2"

Fig. 12. A homemade rodent bait station can contain liquid as


well as solid baits.

B-41
Bait stations for mice should have at
least two openings approximately 1
inch (2.5 cm) in diameter. Locate the
two holes on opposite sides of the sta-
tion so that mice can see an alternate
escape route as they enter the station.
Bait Station Maintenance. Baits
must be fresh and of high quality.
Mice may reject spoiled or stale foods.
Provide enough fresh bait to allow
rodents to eat all they want. When
using rodenticides designed for con-
tinuous bait application (such as anti-
coagulants), bait station maintenance is
essential to a successful baiting effort.
When bait boxes are first put out,
check them daily and add fresh bait as
needed. After a short time, as rodent
numbers and feeding decline, check
the boxes once every 2 to 4 weeks. If
the bait becomes moldy, musty, soiled,
or insect-infested, empty the box and
clean it, and then refill it with fresh
bait. Dispose of spoiled or uneaten bait
Fig. 13. Rodent bait box attached to the top of open dividing wall in a swine confinement facility.
in accordance with the label. Follow all When used in such locations, bait boxes must be securely fastened and out of pigs’ reach.
label directions for the product you are
using.
Placement of Bait Stations. House For example, in swine confinement toxic powder, they pick some of it up
mice are active in a small area and lack buildings it may be possible to attach on their feet and fur and later ingest it
notable food preferences. Therefore, bait boxes to wall ledges or the top of while grooming. Tracking powders are
proper placement of baits or bait sta- pen dividing walls. Bait boxes may be useful in controlling mice where food
tions is often more important than the placed in attics or along floors or alleys is plentiful and good bait acceptance is
type of bait used. Mice will not visit where rodents are active (Fig. 13). difficult to achieve. Mice are more
bait stations, regardless of their con- Rodent tracks visible on dusty surfaces likely to ingest a lethal amount of a
tents, if not conveniently located in and their droppings often give clues to poorly accepted toxicant applied by
areas where they are active. where they are active. this method than if it is mixed into a
bait material. There is little likelihood
Where possible, place bait between the Never place bait stations where live-
of toxicant shyness developing when
rodents’ source of shelter and their stock, pets, or other animals can knock
using tracking powders.
food supply. Put bait boxes near them over. Spilled bait may be a
rodent burrows, against walls or along potential hazard, particularly to Because the amount of material a
travel routes. Where mice are living in smaller animals. mouse may ingest while grooming is
sacked or boxed feed on pallets, baits small, the concentration of active
Where buildings are not rodent-proof,
or traps may have to be placed on top ingredient in tracking powders is con-
permanent bait stations can be placed
of stacks or wedged in gaps within the siderably higher than in food baits that
inside buildings, along the outside of
stacks. In such situations, this “three utilize the same toxicant. Therefore,
building foundations, or around the
dimensional” bait placement is impor- these materials can be more hazardous
perimeter. Bait stations will help keep
tant to obtain good control. Caution than food baits. For the most part,
rodent numbers at a low level when
should be used in selecting control tracking powders are used by profes-
maintained regularly with fresh anti-
methods in such situations. Do not use sional pest control operators and
coagulant bait. Rodents moving in
baits that will contaminate foodstuffs. others trained in rodent control. Track-
from nearby areas will be controlled
For safety, it may not be wise to use ing powders containing either zinc
before they can reproduce and cause
toxic baits in the vicinity of certain phosphide or anticoagulants are com-
serious damage.
foodstuffs. Traps or glue boards may mercially available, although some are
be used instead. Tracking Powders. Toxic dusts or Restricted Use Pesticides.
powders have been successfully used
On farmsteads, bait station placement Place tracking powders along run-
for many years to control mice and
depends on building design and use. ways, in walls, behind boards along
rats. When mice walk over a patch of

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Single trap set with trigger next to wall. The double set increases your success. Double set placed parallel to the wall with
triggers to the outside.

Wrong—trigger not next to wall. Wrong—parallel set with triggers on the inside. Wrong—trap too far from wall.

Fig. 14. Placement of snap traps.

walls, or on the floor of bait stations. efficiently or economically because nutmeat, chocolate candy, dried fruit,
Placement can be aided by using vari- they are small and often difficult to or bacon tied securely to the trigger.
ous types of sifters, shakers, or blow- find. Generally, control of house mice Peanut butter or marshmallows also
ers. Dampness may cause the powder by fumigation is only practical and may be used as bait. Because mice are
to cake and lessen its effectiveness. cost-effective in a very limited number always in search of nesting materials, a
Care must be taken to place tracking of situations. Fumigants are hazardous small cotton ball will also work as a
powders only where they cannot con- materials and should be applied only bait when attached securely to the trig-
taminate food or animal feed, or where by persons well trained in their use ger. Food baits that become stale lose
nontarget animals cannot come into and who possess the necessary safety their effectiveness.
contact with them. Do not place track- equipment.
Set traps close to walls, behind objects,
ing powders where mice can track the
Trapping
in dark places, and in locations where
material onto food intended for use by
mouse activity is seen. Place the traps
humans or domestic animals. Tracking Trapping can be an effective method of so that when mice follow their natural
powders are not generally recom- controlling mice, but it requires more course of travel (usually close to a
mended for use in and around homes labor than most other methods. Trap- wall) they will pass directly over the
because of the potential hazards to ping is recommended where poisons trigger (Fig. 14). Set traps so that the
children and pets. Where possible, seem inadvisable. It is the preferred trigger is sensitive and will spring eas-
remove tracking powder after the method to try first in homes, garages, ily. Effectiveness can be increased by
rodent control program is completed. and other small structures where there enlarging the trigger. Attach a square
Tracking powders used in conjunction may be only a few mice present. of cardboard, metal, or screen wire
with baiting can provide very effective
Trapping has several advantages: (1) it that fits just inside the wire deadfall
mouse control.
does not rely on inherently hazardous (Fig. 15).
Fumigants rodenticides; (2) it permits the user to Use enough traps to make the cam-
view his or her success; and (3) it paign short and decisive. Mice seldom
Fumigants (toxic gases) are most com-
allows for disposal of the mice, thereby venture far from their shelter and food
monly used to control mice in struc-
eliminating odor problems from supply, so traps should be spaced no
tures or containers such as feed bins,
decomposing carcasses that may more than about 6 feet (1.8 m) apart in
railway cars, or other enclosed areas.
remain when poisoning is done within areas where mice are active. Although
Aluminum phosphide, chloropicrin,
buildings. mice are not nearly as afraid of new
and methyl bromide are currently reg-
istered for this purpose. Some fumi- The simple, inexpensive, wood-based objects as rats are, leaving the traps
gant materials are registered for use in snap trap is available in most hard- baited but unset until the bait is taken
rodent burrows; however, house ware and farm supply stores. Traps at least once will reduce the chance of
mouse burrows cannot be fumigated should be baited with a small piece of mice escaping the trap and becoming
trap-shy.

B-43
Multiple-capture (automatic) mouse
traps such as the Ketch-All® and
Victor Tin Cat® (Fig. 16) are available
from some hardware and farm supply
stores as well as from pest control
equipment distributors. These traps
work on the principle that mice enter
Expanded trigger small holes without hesitation. The
Ketch-All® has a wind-up spring that
powers a rotating mechanism. When
triggered, the mechanism entraps mice
in a holding compartment. The Tin
Cat® has one-way doors that mice can-
not exit. Such traps may catch many
mice in a single setting, but should be
checked and emptied periodically so
A box or board placed to that mice do not die of starvation or
advantage may guide mouse exposure in the traps.
into trap.
Various types of box-type traps
(Sherman-type and others) that cap-
ture one mouse at a setting are used
primarily for research purposes. The
desire to “build a better mousetrap”
keeps a variety of traps of variable
effectiveness coming and going on the
retail market.
Place traps across Keep traps reasonably clean and in
obvious runways, good working condition. They can be
or where runs are
cleaned with a hot detergent solution
confined.
and a stiff brush. Human and dead-
mouse odors on traps are not known
Fig. 15. Expanded-trigger traps, when properly placed, can be very effective. to reduce trapping success.
An alternative to traps are glue boards,
which catch and hold mice attempting
to cross them, much the way flypaper
catches flies. Place glue boards wher-
ever mice travel—along walls or in
established runways. Do not use glue
boards where children, pets, or desir-
able wildlife can contact them. Glue
boards lose their effectiveness in dusty
areas unless covered, and temperature
extremes may affect the tackiness of
some glues. They are considered less
effective for capturing rats than for
mice. Glue boards can be purchased
ready-to-use, or they can be made.
Euthanize live, trapped rodents by car-
bon dioxide asphyxiation or use a stick
to kill them with sharp blows to the
base of the skull. For further informa-
tion on glue boards, see the section
Supplies and Materials.
Fig. 16. Automatic multiple-capture mouse traps are commercially available (for example: left, Tin
Cat®; right, Ketch-All®).

B-44
Other Methods increased amount of rodent damage. Burt, W. H., and R. P. Grossenheider. 1976. A
field guide to the mammals, 3rd ed.
Mice, in particular, are very destruc-
Some dogs and cats will catch and kill Houghton Mifflin Co., Boston. 289 pp.
tive to rigid foam, fiberglass batt, and
mice and rats. There are few situa- Chitty, D., and H. N. Southern. 1954. Control of
other types of insulation in walls and
tions, however, in which they will do rats and mice, vol. 1-3. Clarendon Press,
attics of such facilities. In one small
so sufficiently to control rodent popu- Oxford.
swine finishing building near Lincoln,
lations. Around most structures, mice Corrigan, R. M., C. A. Towell, and R. E.
Nebraska, rodent damage required the
can find many places to hide and rear Williams. 1992. Development of rodent
producer to spend $5,000 in repairs to control technology for confined swine
their young out of the reach of such
the facility only 3 years after initial facilities. Proc. Vertebr. Pest Conf. 15:280-
predators. Cats probably cannot elimi- 285.
construction.
nate existing mouse populations, but
in some situations they may be able to Crowcroft, P., and J. N. R. Jeffers. 1961.
prevent reinfestations once mice have Acknowledgments Variability in the behavior of wild house
mice (Mus musculus L.) towards live traps.
been controlled. Farm cats, if sufficient Proc. Zool. Soc. London 137:573-582.
I thank Rex E. Marsh for reviewing a previous
in number and supplementally fed, version of this chapter and providing many Davis, D. E. 1981. Environmental control of
may serve this function. helpful comments. Portions of the rodents. Pages 493-498 in D. Pimentel, ed.
recommendations on toxicant use are taken CRC handbook of pest management in
In urban and suburban areas, it is not directly from his chapter Roof Rats in this agriculture, vol. 1. CRC Press, Inc., Boca
uncommon to find rodents living in manual. Other material contained in this Raton, Florida.
close association with cats and dogs, chapter is derived from Brooks (1973), Marsh
and Howard (1981), and Pratt et al. (1977), Fitzwater, W. D. 1982. Bird limes and rat glues
relying on cat and dog food for nour- among other sources. — sticky situations. Proc. Vertebr. Pest Conf.
ishment. Mice frequently live beneath 10:17-20.
Figure 1 is from Schwartz and Schwartz (1981).
dog houses and soon learn they can Frantz, S. C., and D. E. Davis. 1991. Bionomics
feed on their food when they are Figures 2, 5, and 15 were adapted from Pratt et and integrated pest management of
absent or asleep. al. (1977) by Jill Sack Johnson. commensal rodents. Pages 243-313 in J. R.
Gorham, ed. Ecology and management of
Figures 3 and 14 were adapted from Howard
food-industry pests. Food Drug Admin.
and Marsh (1981) by Jill Sack Johnson.
Tech. Bull. 4, Assoc. Official Analytical
Economics of Damage Figure 4 from Hygnstrom and Virchow (1992). Chem. Arlington, VA.
and Control Figures 6, 7, and 8 were developed by Jill Sack Haines, H., and K. Schmidt-Neilsen. 1967. Water
Johnson. deprivation in wild house mice. Physiol.
Accurate data on mouse damage, Zool. 40:424-431.
Figures 9, 12, and 13 are by Frances I. Gould,
control, and their cost are difficult to University of Nebraska-Lincoln, Cooperative Howard, W. E., and R. E. Marsh. 1981. The rat:
obtain. Estimates of losses of food- Extension. its biology and control. Div. Agric. Sci. Univ.
stuffs, structural damage, and the California, Leaflet 2896 (revised). 30 pp.
Figures 10 and 11 were adapted from Pratt et al.
amount of labor and materials expend- (1977) by Frances I. Gould. Humphries, R. E., A. P. Meehan, and R. M. Sibly.
ed to control mice are usually only 1992. The characteristics and history of
educated guesses. behavioural resistance in inner-city house
For Additional mice (Mus domesticus) in the U.K. Proc.
In one survey of corn in a midwestern Vertebr. Pest Conf. 15:161-164.
state, 76% of about 1,000 grain samples Information
Hygnstrom, S. E., and D. R. Virchow. 1992.
were contaminated with rodent Controlling house mice. Univ. Nebraska
Berry, R. J. 1981. Town mouse, country mouse: Coop. Ext. NebGuide G92-1105-A. 4 pp.
droppings. Mouse droppings outnum- adaptation and adaptability in Mus
bered rat droppings twelve to one. A domesticus (M. musculus d). Mammal Rev. Jackson, W. B. 1990. Rats and mice. Pages 9-85 in
house mouse produces about 36,000 11:91-136. A. Mallis, ed. Handbook of pest control.
droppings in a year’s time. Mouse in- Franzak and Foster Co., Cleveland, OH.
Berry, R. J., ed. 1981. Biology of the house
festations are so widespread that drop- mouse. Symp. Zool. Soc. London, No. 47. Kaukeinen, D. E. 1982. A review of the
pings and hairs often end up in many 715 pp. secondary poisoning hazard potential to
wildlife from the use of anticoagulant
types of food commodities intended Bohills, S. T., A. P. Meehan, and S. P. Leonard. rodenticides. Proc. Vertebr. Pest Conf.
for human use. Certain levels of rodent 1982. Advantages of bait boxes in house 10:151-158.
contamination are grounds for con- mouse control. International Pest Control
24(2):34, 35, 37. Kaukeinen, D. E. 1984. Resistance: what we
demning food commodities. need to know. Pest Manage. 3(3):26-30.
Bronson, F. H. 1979. The reproductive ecology of
Structural damage caused by rodents the house mouse. Quarterly Rev. Biol. Knote, C. E. 1988. Stopping house mice building
can be expensive. In recent years, the 54(3):265-299. infestations through exterior control. Proc.
trend toward use of insulated confine- Vertebr. Pest Conf. 13:107-111.
Brooks, J. E. 1973. A review of commensal
ment facilities to raise swine in the rodents and their control. CRC Critical Rev.
northern Great Plains has led to an Environ. Control 3:405-453.

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Labov, J. B. 1981. Male social status, physiology, Robbins, R. J. 1980. Taste-aversion learning and
and ability to block pregnancies in female its implications for rodent control. Proc.
house mice (Mus musculus). Behav. Ecol. Vertebr. Pest Conf. 9:114-121.
Sociobiol. 8:287-291.
Rowe, F. P. 1966. Economic importance of the
Marsh, R. E., and W. E. Howard. 1981. The house mouse (Mus musculus L.). World
house mouse: its biology and control. Div. Health Organiz., Vector Control, Pub. 66.217,
Agric. Sci. Univ. California, Leaflet 2945 paper 1.4:21-26.
(revised). 30 pp.
Schwartz, C. W., and E. R. Schwartz. 1981.
Meehan, A. P. 1984. Rats and mice: their biology House mouse. Pages 248-252 in The wild
and control. Rentokil Ltd., E. Grinstead, U.K. mammals of Missouri, rev. ed. Univ.
383 pp. Missouri Press, Columbia. 356 pp.
Morris, K. D., and D. E. Kaukeinen. 1988. Shenker, A. M. 1973. The house mouse in
Comparative evaluation of tamper-proof London. Mammal Rev. 3:64-69.
mouse bait stations. Proc. Vertebr. Pest Conf.
13:101-106. Weber, W. J. 1982. Diseases transmitted by rats
and mice. Thomson Pub. Fresno, California.
Petras, M. L., and J. C. Topping. 1981. Studies of 182 pp.
natural populations of Mus. VI. Sizes of
populations inhabiting corn cribs in
southwestern Ontario. J. Mammal. 62:146-
153.
Editors
Pratt, H. D., B. F. Bjornson, and K. S. Littig. 1977.
Scott E. Hygnstrom
Control of domestic rats and mice. Public
Robert M. Timm
Health Serv., US Dep. Health, Educ. Welfare,
Gary E. Larson
Pub. No. (CDC) 77-841. 47 pp.

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