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Bread, beer, and Bordeaux: most of us love some or all of these!

But they would not exist if not for yeast, a eukaryotic microorganism that can metabolize sugars anaerobically
through a pathway called alcohol fermentation. Humans have been using yeasts to make these products for thousands of years, but only learned of their existence in the last
two hundred years. How exactly do these tiny creatures make these delicious food and drink items?
Alcohol fermentation, also known as ethanol fermentation, is the anaerobic pathway carried out by yeasts in which simple sugars are converted to ethanol and carbon dioxide.
Yeasts typically function under aerobic conditions, or in the presence of oxygen, but are also capable of functioning under anaerobic conditions, or in the absence of oxygen.
When no oxygen is readily available, alcohol fermentation occurs in the cytosol of yeast cells. Let's explore the process of alcohol fermentation then see what it means for
yeasts and for humans.

The Process of Alcohol Fermentation


The basic equation for alcohol fermentation shows that yeast starts with glucose, a type of sugar, and finishes with carbon dioxide and ethanol. However, to better understand
the process, we need to take a look at some of the steps that take us from glucose to the final products.
The process of alcohol fermentation can be divided into two parts. In the first part, the yeast breaks down glucose to form 2 pyruvate molecules. This part is known as glycolysis.
In the second part, the 2 pyruvate molecules are converted into 2 carbon dioxide molecules and 2 molecules of ethanol, otherwise known as alcohol. This second part is called
fermentation.
The main purpose of alcohol fermentation is to produce ATP, the energy currency for cells, under anaerobic conditions. So from the yeast's perspective, the carbon dioxide and
ethanol are waste products. That's the basic overview of alcohol fermentation. Now, let's examine each part of this process in greater detail.

In the first part of this process, each glucose molecule is broken down into 2 pyruvate molecules. Pyruvate, or pyruvic acid, is an amino acid and will help form ethanol. In the
process of breaking glucose down to form pyruvate, several molecules known as electron acceptors are involved.
Electron acceptors are molecules whose job is to give and take the electrons released when a chemical reaction takes place. During this first part, an electron acceptor
molecule called NAD+ is reduced to form NADH, gathering up the electrons released by breaking one glucose down to 2 pyruvate molecules. This exchange of electrons that
occurs while glucose is being broken down i

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