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Contents-
Introduction
Origin of life
Evolution of life forms- A theory
What are the evidences for evolution
What is adaptive Radiation?
Biological evolution
Mechanism of evolution
Hardy –Weinberg Principle
Origin and Evolution of Man
Introduction:
- Evolutionary biology is the study of history of life
forms on earth.
To understand the changes in the flora and fauna that
have occurred over million of years on earth, we must
have an understanding of context of origin of life i.e.
evolution of earth, of stars and indeed the universe
itself.
Evolution means act of unfolding and unrolling or
evolution is orderly ‘change’ from one form to another.
 Figure of milky way
Figure of early earth
 Origin of life- The
Universe:
Life is an inherent capacity that an organism possesses to maintain
and reproduce itself.
Stellar distances are measured in light years.
The universe is very old – almost 20 billion years old.
The Big Bang theory attempts to explain to us the origin of
universe.
The Big Bang theory:
A singular huge explosion unimaginable in physical term.
The universe expanded hence the temperature came down and
Hydrogen and Helium were formed later.
The gases condensed under gravitation and formed the galaxies
of the present day universe.
In the solar system of the Milky Way galaxy, earth was
supposed to have been formed about 4.5billion years back.
 Condition of early earth:
Earth formed 4.5 billion years back.
There was no atmosphere on early earth.
Water vapour, methane, carbon dioxide and ammonia released
from molten mass covered the surface.
The UV rays from the sun broke up water into Hydrogen and
oxygen and lighter H2 escaped.
Oxygen combined with ammonia and methane to form water,
CO2 and others.
The ozone layer was formed.
As it cooled, the water vapour fell as rain, to fill all the
depressions and formoceans.
Life appeared 500 Million years after the formation of
earth.
Theories of Origin of Life-
Early Greek thinkers thought units of life called spores were
transferred to different planets including earth.

Various theories were proposed to explain the origin of life:

1.Theory of Special Creation
2. Theory of Panspermia/ Cosmozoic theory
3. Theory of Spontaneous Generation
4. Theory of Chemical Evolution
1. Theory of Special Creation:
•Conventional religious literature talks about the theory of special
creation, according to which God, the supernatural being created the
earth, light, plants and animals
• This theory has three suggested views:
I All living organisms (different species or types of plants & animals)
that we see today were created as such.
ii. The diversity of life forms was always the same since the time of
creation & will remain the same in future also.
iii. Earth is about 4000 years old
• All these ideas underwent drastic changes during 19th century when
Charles Darwin showed that the existing life forms share similarities
to varying degree among themselves and also with those life forms
that existed millions of years ago.
•Extinctions also took place during at different phase of geological
History.
2. Theory of Panspermia/ Cosmozoic theory:
• Some scientist believe that life has come on earth from
other planets or outer space.
• Early Greek philosophers thought units called ‘spores’ or
‘pan sperms’ came on earth along with meteorites & they
might have evolved into present day forms.
3. Theory of Spontaneous Generation:
• It was also believed that living organisms (life) arose from
decaying matter like straw.
• But Louis Pasteur demonstrated that life can arise only from
pre-existing life.
• He showed that no life arose from the heat- killed yeast broth
that was kept in pre- sterilized closed flask, while new
organisms arose from the heat- killed yeast that was kept in
the flask left open in the air.
4. Theory of Chemical Evolution:
• This theory was proposed by Oparin
& Haldane
• They proposed that first life form
could have come from pre- existing
non- living organic molecules (like
RNA, proteins, etc.) and that
formation of life was preceded by
chemical evolution, i.e., formation
of diverse organic molecules from
inorganic constituents.
• The conditions on earth that
favoured chemical evolution were
very high temperature, volcanic
storms & reducing atmosphere that
contained methane, ammonia,
water vapour etc.
Experimental Proof for Chemical Evolution of
Life..
• Stanley Miller & Urey created conditions similar to the
primitive
atmosphere in the laboratory using glass apparatus & tubes.
• They created electric discharge using electrodes in a conical
flask containing methane, ammonia, hydrogen & water vapour
at 800ºC.
• The water containing chamber was heated to provide water
vapour.
• After a week, they observed the formation of amino acids.
• In similar experiments, many other scientists observed the
formation of sugar, nitrogen bases, lipids, amino acids & even
pigments.
• Analysis of the meteorites also revealed the presence of similar
compounds, indicating the occurrence of similar processes
elsewhere in the space.
• The chemical evolution of life was more or less accepted.
ORIGIN OF FIRST
• The first non- cellular form of life could have originated three
LIFE
billion years ago.
•They would have been giant molecule like RNA, proteins or
polysaccharides
• These capsules were able to replicate/ reproduce
• The first cellular forms of life must have appeared about less than 2
billion years ago, they must have been single cells
• All life forms have appeared in water bodies.
•This is theory of Abiogenesis i.e., the first form of life arose slowly
through evolutionary forces from non- living molecules; it is
accepted by the scientists
• Once formed, these single cells must have evolved into the diverse
complex organisms oftoday
 Organism has undergone gradual evolution due to variation of
characteristics which enable some to survive better in natural
conditions (climate, food, physical factors, etc.) would
outbreedothers that are less-endowed to survive under such
natural conditions.
•It was referred as fitness of individual/ population which means
reproductive fitness.
• Those who are better fit in an environment, leave more progeny
than others which will survive more and hence are selected by
nature. He called it natural selection and implied it as a
mechanism of evolution.
• Alfred Wallace, a naturalist who worked in Malay Archipelago
had also come to similar conclusions that Darwin had.
•All the existing life forms share similarities and share common
ancestors.
• They also concluded is that earth is very old, not thousand of years as was
thought earlier but billions of years old.
 EVIDENCES FOR
EVOLUTION
Evidences for evolution come from, Palaeontology,
Embryology, Comparative anatomy and morphology,
Molecular homology & Biogeography
1. Palaeontology:
• Is a study of fossils found in the rocks to support organic
evolution.
• Study of fossils from different sedimentary layers indicates:
I . The geological time period in which the organisms
existed.
ii.The life forms varied over time & certain life forms are
restricted to certain geological life- span.
iii.The new forms of life that have appeared at different
times in the history of earth.
Contd.
.•2.ItEmbryology:
is a study based upon the observation of certain common
features during embryonic stages of all vertebrates.
• Ernst Haeckel proposed biogenetic law which states that
‘ontogeny recapitulates phylogeny’, i.e., developmental/
embryological stages of organisms (ontogeny) recapitulate
theirevolutionary history (phylogeny).
• Vertebrate embryos shows similarities which are:
1. All vertebrate embryos develop a row of gill slits, but they are
functional only in fish & not found in any other vertebrates.
2. Notochord is present in all vertebrate embryos.
• But this proposal was disapproved by Ernst Von Baer; he noted
that embryos never pass through the adult stage of other
animals.
Comparative Anatomy &
Morphology:
Comparative anatomy and morphology shows both similarities &
differences among the present day organisms & those existed long
before.
i. Homology
•Homology is the relationship among organs of different groups of
organism, that show similarity in the basic structure and embryonic
development, but perform different functions.
•Homology of organs of different organisms indicates their common
Ancestry.
•Homology is found in the bones of forelimbs of whales, cheetah,
birds, amphibians and human; they have similar basic anatomical
structure with the bones humerus, radius, ulna, carpals, matacarpals
& phalanges.
• The thorns of Bougainvillea & tendrils of Cucurbitarepresents
homology
Homology/ homologous organs are the result of divergent evolution,
i.e., the evolutionary process where the same structure develops
along different directions due to adaptations for different needs.
Analog
•Analogy is the relationship among organs of different groups of
yorganisms performing the same function, irrespective of structural or
anatomical differences.
• E g.- Eyes of octopus & those of mammals; wings of butterfly & those of
birds; flippers of whales or dolphins & those of penguins; tubers of
sweet potato (root modified) & those of potato (stem modified)
•Analogy is the result of convergent evolution, i.e., the evolutionary
process, where anatomically different structures in different groups of
organisms evolve towards the same function.
• It is the similar habitat conditions that have selected similar adaptive
features in different groups of organisms towards the same function.
Molecular Homology:
•Molecular homology refers to the similarities in the
bio molecules of different groups of organisms.
•The sequences of nucleotides in nucleic acid & many
proteins are similar in apes & humans
•The biochemical similarities point to the same/
common ancestry of diverse organisms
Biogeography:
•The differential geographical distribution of different
organisms also indicates common/ shared ancestry in
that restricted region.
•Habitat isolation has probably restricted these
organisms to particular geographical regions on the
earth.
Convergent
evolution:
Wings of butterfly and of birds look alike.
They are anatomically similar structure though they
perform similar function.
Hence analogous structures are a result of
convergent evolution.
Eye of octopus and eye of mammals.
Flippers of Penguins and Dolphins.
Sweat potato (root modification) and potato
(stem modification).
 Theory of Natural
• Darwin
Selection
made a sea voyage round the world in a sail ship, named H.
M. S Beagle
•Based on the observations he made during this voyage, he
concluded the following:
i. The existing living organisms share similarities to varying degrees
not only among themselves but also with the life forms that
existed millions of years ago.
ii.There has been gradual evolution of life forms during different
periods of geological history and there has been extinctions also.
iii. Any population has built- in variations in characteristics.
iv.Individuals with those characteristics which enable them to
survive better (fitness of the individual) in the natural conditions,
would overcome the others, who are less adapted under the
same natural conditions
Evolution by anthropogenic
action:
Excess use of herbicides, pesticides etc., has only resulted
in selection of resistant varieties in a much lesser time
scale.
This is also true for microbes against which we employ
antibiotics or drugs against eukaryotic organisms/cell.
Hence resistance organisms/cells are appearing in a time
scale of months or years and not in centuries.
These are the examples of evolution by anthropogenic
action.
Evolution is a stochastic process based on chance
events in nature and chance mutation in the
organisms.
WHAT IS ADAPTIVE
Darwin’s Finches:
RADIATION?
In Galapagos Islands Darwin observed small black birds later
called Darwin’s Finches.
He realized that there were many varieties of finches in the
same island.
All the varieties, he came across, evolved on the island itself.
Form the original seed-eating features, many other forms with
altered beaks arose, enabling them to become insectivorous
and vegetarian finches
This process of evolution of different species in a given
geographical area starting from a point and literally radiating
to other areas of geography (habitats) is called adaptive
radiation.
DARWIN’s
FINCHES
Australian
marsupial:
A number of marsupials each different from the other
evolved from an ancestral stock. But all within the
Australian island continent.
When more than one adaptive radiation appeared to have
occurred in an isolated geographical area (representing
different habitats), one can call this convergent
evolution.
Placental mammals in Australia also exhibit adaptive
radiation in evolving into varieties of such placental
mammals each of which appears to be ‘similar’ to a
corresponding marsupial(e.g. placental wolf and
Tasmanian wolf-marsupial).
 BIOLOGICAL EVOLUTION:
•The essence of Darwinian Theory about evolution is natural
selection.
•The rate of appearance of new forms is linked to the life cycle or
the life span.
•There must be a genetic basis for getting selected and to evolve.
•Some organisms are better adapted to survive in an otherwise
hostile environment.
•Adaptive ability is inherited.
•It has genetic basis.
•Fitness is the end result of the ability to adapt and get selected by
nature.
•Branching descent and natural selection are the two key
concepts of Darwinian Theory of Evolution.
MECHANISM OF
In the first decade of twentieth century, Hugo de Vries
EVOLUTION:
based on his work on evening primrose brought fourth
the idea of mutations.
Mutation is the large difference arising suddenly in a
population.
How de Vries theory of mutation differs from Darwin’s
theory of natural selection?
It is the mutation which causes evolution and not the minor
variations that Darwin talked about.
Mutations are random and directionless while Darwinian
variations are small and directional.
Evolution for Darwin was gradual while de Vries believed
mutation caused speciation and hence called it
saltation(single step large mutation).
HARDY –WEINBERG
In a given population one can find out the frequency of occurrence
PRINCIPLE:
of alleles of a gene on a locus.
This frequency is supposed to remain fixed and even remain the
same through generations.
Hardy-Weinberg principle stated it using algebraic equations.
The principle states that allele frequencies in a population are stable
and is constant from generation to generation.
The gene pool (total genes and their alleles in a population) remains
a constant. This is called genetic equilibrium:
Sum total of all the allelic frequencies is 1.
(p + q)2= p2+ 2pq + q2= 1.
When frequency measured, differs from expected values, the
difference (direction) indicates the extent of evolutionary change.
Disturbance in genetic equilibrium, or i.e. change of frequency of
alleles in a population would then be interpreted as resulting in
evolution.
Five factors are known to affect Hardy-Weinberg equilibrium:

Gene migration or gene flow.

Genetic drift.
Mutation.
Genetic recombination.
Natural selection.

Gene migration : When migrations of a section of population to

another place occur, gene frequencies change in the original as
well as in the new population. New genes /alleles are added to
the new population and these are lost from the old population.
Gene flow: Gene migration occurs many time is termed as gene
flow.
Genetic drift: change in gene frequency takes place by chance.
Founder effect : sometimes the change in allelic frequency is so
different in the new sample of population that they became a
different species. The original drifted population becomes
founder species and the effect is called founder effect.
Operation of natural selection on different trait:
Natural selection can lead to :
Stabilization: in which more individuals acquire
mean charactervalue.
Directional changes i.e. more individuals acquire
value other than the mean character value.
Disruption : more individuals acquire peripheral
character value at both ends of the distribution
curve.
A BRIEF ACCOUNT OF
About 2000 million years ago (mya) the first cellular forms of life
EVOLUTION:
appeared on earth.
Some cellular form had the ability to release oxygen.
Slowly single cell organisms became multi-cellular life forms.
By the time of 500 mya invertebrates were formed and active.
Jawless fish probably evolved around350 mya.
Sea weeds and few plants existed probably around 320 mya.
Coelacantha lobe finned fish discovered in South Africa in
1938 evolved into first amphibians that lived on both land
and water. These were ancestors of modern day frogs and
salamanders.
The amphibian evolved into reptiles.
Reptiles’ lays eggs which don not dry up in sun unlike those of
amphibians.
Giant ferns(pteridophytes) were present but they fell to
form coal deposits slowly.
Some of the reptiles went back into water to evolve into fish
like reptiles probably 200 mya(Ichthyosaurs)
The land reptiles were the dinosaurs.
The biggest dinosaurs are Tyrannosaurus rexwas about 20
feet in height and had huge fearsome dagger like teeth.
About 65 mya the dinosaurs suddenly disappeared from the
earth.
Some of them evolved into birds.
The first mammals were like shrews. Their fossils were small
sized.
Mammals were viviparous and protected their unborn young
inside the mother’sbody.
Due to continental drift pouched mammals of Australia
survived because of lack of competition from any other
mammals.
Evolution of Man
•The common ancestor of apes and man is a primate Dryopithecus, that lived 15 million
years ago
• At the same time, another genus Ramapithecus also existed
•Both Dryopithecus & Ramapithecus were hairy and walked like gorillas & chimpanzees;
Dryopithecus was more ape like, but Ramapithecus was more man-like and is the
forerunner of hominid evolution
The human evolution is as follows:
1. Australopithecines
•They probably lived 2 million years ago, in the east Africa grass
lands
• They has brain capacity of 450- 600 cc
• They hunted with stone weapons but essentially ate fruits.
2. Homo habilis
• This is called first human like being, the hominid
• They had brain capacity of 650- 800 cc.
• They probably did not eat meat
3. Homo erectus
• Their fossils were found in java (Java man) in 1981
• They probably lived about 1.5 mya
• They had a brain capacity of about 900cc
• They probably ate meat.
4. Homo sapiens (Primitive man)
• Their fossils were found in near East & Central Asia
• He must have lived between 1,00,000- 40,000 years before
• Neanderthal man (Homo sapiens neanderthalensis) had a brain
capacity of about 1400 cc.
• They used hides to protect the body & buried the dead
• They moved across continents & developed into distinct races
5. Homo sapiens sapiens (Modern Man)
• Homo sapiens sapiens arose during ice age between 75000- 10000
years ago
•He spread all over the globe & learned to cultivate plants &
domesticate animals
• Pre- historic cave art developed about 18,000 years before
• Agriculture started around 10,000 years back
• Human settlements and cultivations started
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