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Diptera
General identification characters:
Flies range in size from midges of little more than 1 mm to robber flies more than
7 cm long with antennae generally shorter than their head.
Body is often covered with short bristles
A large, moveable head with compound eyes that are often very large
Haustellate mouthparts
Front wings membranous; hind wings modified into knob like structure called
halteres
Thorax enlarged in the middle to give it a rather hunch-backed look
Immatures:
1. Culiciform
2. Head capsule present with chewing mouthparts
3. Legs absent
4. Vermiform (maggots)
5. Without legs or a distinct head capsule
6. Mouthparts reduced; only present as mouth hook
Adults:
1. Antennae filiform, stylate, or aristate
2. Mouthparts suctorial (haustellate)
3. Mesothorax larger than pro- or metathorax
4. One pair of wings (front); hind wings reduced (halteres)
5. Tarsi 5-segmented
Economic importance:
The economic importance of the group is immense. Members of this order are the most
ecologically diverse and probably have a greater economic impact on humans than any
other group of insects. Many species are particularly important as vectors of disease in
man, other animals, and plants. The abundance, worldwide distribution, and habits of
flies combine to make them a nuisance to humans. They are also responsible for
carrying different disease causing agents. Many members in the order cause damage to
plants and field crops resulting in economic losses.
On the other hand, many dipterans serve roles that are useful to humans. Houseflies,
blowflies and fungus gnats are scavengers and aid in decomposition. Robber flies,
dagger flies and balloon flies are predators and parasitoids of other insects, helping to
control a variety of pests. Many dipterans such as bee flies (Bombyliidae) and hoverflies
(Syrphidae) are pollinators of crops and have essential roles in maintaining our
ecosystem. In addition, much of our knowledge of animal genetics and development
has been acquired using the vinegar fly Drosophila melanogaster (family Drosophilidae)
as an experimental subject.