Національна академія наук України

Державний природознавчий музей

Наукові колекції
Державного природознавчого музею
Випуск 2
Р.Й. Годунько, В.К. Войчишин, О.С. Климишин та ін.
НОМЕНКЛАТУРНІ ТИПИ І ТИПОВІ СЕРІЇ (1)

Scientific Collections
of the State Natural History Museum
Issue 2
R.J. Godunko, V.K. Voichyshyn, O.S. Klymyshyn et al.
NAME-BEARING TYPES AND TYPE SERIES (1)

Львів 2006
 26

NAME-BEARING TYPES AND TYPE SERIES OF FOSSIL AGNATHANS

(AGNATHA: HETEROSTRACI, OSTEOSTRACI)

The oldest documented collection of fossil agnathans in the State Museum of Natural
History (SMNH) consists of several specimens which were found in Podolia by
palaeontologist Dr. Мarian Łomnicki at the end of the 19th century. These findings were
included in the natural history assemblage of the Museum of Dzieduszycki (previous name
of SMNH). At least from 1924 to 1940 Professor Władysław Zych, who was working at the
Lviv University named in honor of Jan Kazimierz, collected a lot of fish fauna remains
from the Lower Devonian rocks in Podolia. Evidently his collection included much more
than one thousand specimens. A part of his findings W. Zych had donated to the British
Museum of Natural History in London and to the Naturhistoriska Riksmuseet in Stockholm
in gratitude for a help of English and Swedish sponsors (Janvier 1985). Some Podolian
specimens were sent by him to the Palaeontologisk Museum in Oslo (Zych 1931: 83,
footnote 1). After World War II a part of Zych’s collection was moved to Poland. The main
part of the rest collection, which had been left at the Department of Geology of the Lviv
University, was passed to the SMNH at the request of Dr. Pavlo Balabai (Balabai 1957:
305; Balabai 1962: 3). During 1948-1959 P. Balabai, who was studying Early Devonian
agnathans, supplemented the fossil fish collection by his own materials. At that time the
collection was also enriched by specimens which had been found by V. Shtym (in 1955),
S. Kotsubynskyi (in 1956), V. Gavrylyshyn (in 1958) and many others. Between 1986 and
2006 the SMNH collection of fossil fishes considerably increased in numbers owing to the
materials collected by the author of this paper, Dr. Daniel Drygant, S. Solodkyi and others.
The first (new to a science) agnathans species were established on Podolian materials
by A. Alth (1874), W. Zych (1927-1937) and F. Brotzen (1933-1936). However, the little is
known about the destiny of its type series. It is possible that some type materials on these
taxa have been lost. There is Alth’s collection of fossils from Podolia in Krakow, but at
present I have no information whether it consists also of type materials concerning
Podolian agnathans. A. Blieck (1984: 23 and footnote) has noted that Brotzen’s collection
is partly housed in Stockholm and partly in the Museum of Natural History of Humboldt
University in Berlin. L. Novitskaya (1986: 24, footnote 1) has stated that all attempts to
reveal the present location of Brotzen’s collection, which early was housed in Geological
and Palaeontological Institute in Berlin, did not give any results. Perhaps some type
specimens of Zych’s assemblage are kept in Poland. This year, owing to favour of Mr. Piotr
Szrek, I have seen at least plaster-cast copies of Pteraspis lerichei mut. latissima and
Pteraspis lerichei mut. major syntypes in the Collection of the Institute of Geology of the
Warsaw University. But the attempt to ascertain the whereabouts of the original specimens
at present has been failed.
The SMNH collection includes type series of the species which have been
designated by P. Balabai (1962) and V. Voichyshyn (1994-2006). It should be noted that
P. Balabai has cited some numbers of studied specimens only in captions to figures. In
general he has just indicated a total number of specimens he had studied. The authors of the
Reference Book dealt with palaeontological monographical collections of the Museum did
not possess full information on that question (Pasternak et al. 1963: 21). Thus, in order to
find out type series of the species established by P. Balabai, it was necessary to use the
Recommendation 72B of ICZN in some cases (see Voichyshyn, this volume).
 27

The collection of Fossil Fishes (ВР) belongs to the Section “Palaeozoology:

Vertebrates” (ПЗХ) within the Palaeontological SMNH Assemblage. Localities of type
specimens are marked on the schematic map of Podolia (Fig. 1). Since the old register
numbers on fossil fish specimens in the Museum collection beginning with figures “17***”
are related to Zych’s assemblage, it may be assumed that just he was a collector of those
specimens although the labels have been lost. The images of type specimens are given if in
previous publications they were of insufficient quality.
 28

Subphylum Agnatha Cope, 1889

Class Heterostraci Lankester, 1868
Order Pteraspidiformes Berg, 1940
Family Larnovaspididae Halstead in Novitskaya 1986

Genus Alaeckaspis Voichyshyn, 1999

Alaeckaspis ustetchkiensis Voichyshyn, 1999

The type series consists of single specimen (Voichyshyn 1999: 51).

Ho loty pe : BP.168 (old number 37682); incomplete mould and imprint of carapace
in dorsal view.
Image: Plate I: 1A-B.
Locality, age and collection data: left bank of the Dniester, the first ravine along
the country road from Ustechko village to the old broken bridge over the Dniester
(Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower Devonian, Old-Red
Formation (the Dniester Series, the lowest part of its Khmeleva Member); collected by
V. Voichyshyn on July 11, 1989.
Publication(s): Voichyshyn 1999: 52, fig. 3A-B.

Alaeckaspis verbivciensis Voichyshyn, 1999

The type series consists of twelve specimens (Voichyshyn 1999: 49). All of them
were found by unknown collector on September 9-10, 1960 in a quarry “on a bank over the
Zvyniach Stream” in the vicinity of Verbivtsi village (Terebovlia rayon, Ternopil’ oblast’
(Podolia), Ukraine). The horizon of exposure is unclear. Possibly the rocks may belong to
Ivanie horizon or (Old-Red Formation) Ustechko Member of the Dniester Series.

Ho loty pe : BP.167/1 (old number 31399/1); almost complete carapace.

Image: Plate I: 2.
Publication(s): Voichyshyn 1999: 50, fig. 1A.
Pa raty pe : BP.167/2 (old number 31399/2); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 51, fig. 2A.
Pa raty pe : BP.167/3 (old number 31399/3); incomplete carapace.
Publication(s): Voichyshyn 1999: 51, fig. 2B.
Pa raty pe : BP.180 (old number 37712); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 51, fig. 2C.
Pa raty pe : BP.187 (old number 31395/3); incomplete mould of carapace in dorso-
ventral view.
Publication(s): Voichyshyn 1999: 49.
Pa raty pe : BP.188 (old number 31412/1); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 49.
Pa raty pe : BP.218 (old number 37719); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 49.
Pa raty pe : BP.219 (old number 37718); a contour of carapace.
 29

Publication(s): Voichyshyn 1999: 49.

Pa raty pe : BP.220 (old number 31395/1); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 49.
Pa raty pe : BP.221 (old number 31395/2); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 49.
Pa raty pe : BP.222 (old number 31399/4); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 49.
Pa raty pe : BP.645 (old number 37971, erroneously has been designated in the
paper as 31412/2); incomplete mould of carapace.
Publication(s): Voichyshyn 1999: 49.

Pteraspidiformes indet.

Genus Pavloaspis Voichyshyn, 1999

Pavloaspis pasternaki Voichyshyn, 1999

The type series consists of single specimen (Voichyshyn 1999: 53).

Ho loty pe : BP.169 (old number 37585); incomplete internal mould of carapace

from the both dorsal and ventral side, and its imprint (lacking a posterior-lateral part) from
the dorsal side.
Image: Plate I: 3.
Locality, age and collection data: left bank of the Dniester, the outcrop in NW of
Ustechko village (Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower
Devonian, the Old-Red Formation (the Dniester Series, the boundary between its Ustechko
and Khmeleva Members); collected by V. Voichyshyn on July 13, 1989.
Publication(s): Voichyshyn 1999: 54, fig. 4A.

Class Osteostraci Lankester, 1868

Order Cephalaspidida Stensiö 1958
Family Mimetaspididae Stensiö 1958

Genus Mimetaspis Stensiö 1968

Mimetaspis concordis Voichyshyn, 1994

There are five specimens in the type series (Voichyshyn 1994: 20). All of them have
the same next data: left bank of the Dniester, the outcrop in NW of Ustechko village
(Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower Devonian, the Old-Red
Formation (the Dniester Series, the lowest part of its Khmeleva Member); collected by
V. Voichyshyn on July 17, 1990.

Ho loty pe : BP.982 (old number 35617); mould of the shield from dorsal view and
imprint of orbito-pineal + otico-occipital regions of the shield.
Image: Plate II: 3.
 30

Publication(s): Voichyshyn 1994: 20, table I, fig.2, table II, fig. 1.

Pa raty pe : BP.983 (old number 38616); imprint of the shield from dorsal view.
Image: Plate II: 4.
Publication(s): Voichyshyn 1994: 20, table I, fig. 1, table II, fig. 2.
Pa raty pe : BP.985 (old number 38615); fragmentary mould and imprint of the
shield, lacking the posterior part and cornual process of the right side.
Image: Plate II: 5.
Publication(s): Voichyshyn 1994: 20, table I, fig. 4.
Pa raty pe : BP.981 (old number 38618); mould and imprint of (mainly) contours
of the shield from ventral side.
Publication(s): Voichyshyn 1994: 20.
Pa raty pe : BP.984 (old number 38619); fragmentary mould and imprint of the
shield from dorsal side.
Publication(s): Voichyshyn 1994: 20.

Family Pattenaspididae Stensiö 1958

Genus Pattenaspis Stensiö (1958)

Pattenaspis rogalai (Balabai, 1962)

The type series was stated by P. Balabai to include twenty one specimens (Balabai
1962: 3). But one can see below that, at least, in two cases a mould and an imprint of the
same specimen were numbered as separate specimens. Although P. Balabai gave the picture
of one specimen collected in Horodnytsia (Balabai 1962: fig. 1a), it has not been found.

Lectoty pe : BP.990 (old number 17311); fragmentary mould of the shield from
dorsal side, lacking cornual processes.
Locality, age and collection data: W vicinity of Horodnytsia village (Horodenka
rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower part of the
Old-Red Formation (the Dniester Series, most probably its Ustechko Member); collected by
W. Zych and E. Stensiö on September 6, 1935.
Publication(s): Afanassieva et al. 1991: 67, table IX, figs. 1-2.
Pa ra lectotype : BP.1359 (old numbers 17310 and 17312, mould and imprint of
the same individual’s shield); fragmentary mould and imprint of shield from dorsal side,
lacking cornual processes.
Locality, age and collection data: the vicinity of Horodnytsia village (Horodenka
rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower part of the
Old-Red Formation (the Dniester Series, most probably its Ustechko Member); collected by
W. Zych.
Publication(s): Afanassieva et al. 1991: 69, fig. 2.
Pa ra lectotype : BP.33 (old numbers 17316 and 17411, mould and imprint of the
same individual’s shield, correspondingly); fragmentary mould and imprint of shield,
lacking a central part and right cornual process.
Image: Plate II: 1A-B.
 31

Locality, age and collection data: W vicinity of Ostrivets’ village (Terebovlia

rayon, Ternopil’ oblast’, Ukraine); Lower Devonian, probably, uppermost part of the Ivanie
horizon or lowest part of the Old-Red Formation (Ustechko Member of the Dniester
Series); collected by W. Zych in 1936.
Publication(s): Afanassieva et al. 1991: 69.

Order Zenaspidida Stensiö 1958

Family Zenaspididae Stensiö, 1958

Genus Zenaspis Lankester, 1870

Zenaspis podolica (Balabai, 1962)

The type series was stated by P. Balabai to include ten specimens (Balabai 1962: 4).
It is possible to define only three specimens among them. The specimens BP.34 (17318)
and BP.35 (17323) originally labelled by the author of the species as “Cephalaspis podolica
sp. n.” then have been redefined by him as “C. microlepidota”. In that way, the author
caused some confusion, and it is unknown whether two latter specimens can be regarded
among initial ten ones.

Lectoty pe : BP.37 (old number 17538); it was single specimen of the species
figured in Balabai’s paper; mould of damaged shield, lacking a distal part of left cornual
process.
Locality, age and collection data: the vicinities of Horodnytsia village (Horodenka
rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower part of the
Old-Red Formation (the Dniester Series); probably collected by W. Zych.
Publication(s): Balabai 1962: 5, fig. 4; Voichyshyn, this volume, Plate I: 1.
Pa ra lectotype : BP.30 (old number 25561); a fragment of the shield mould and a
series of trunk segments (scales); left bank of the Dniester.
Locality, age, collection data and note: the vicinity of Ivane-Zolote village
(Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower Devonian, lowest part of
the Old-Red Formation (the Dniester Series, lowest part of its Ustechko Member, near the
boundary with the Ivanie horizon); collected by P. Balabai in 1956; the specimen was
labelled by P. Balabai as “C. podolica sp. n.”.
Pa ra lectotype :BP.36 (old number 25562); fragmentary mould of the shield.
Locality, age, collection data and note: left bank of the Dniester, the vicinity of
Ustechko village (Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower
Devonian, the Old-Red Formation (lower part of the Dniester Series); collected by
Tykhomyrov; old number of this specimen was wrongly used in caption of the lectotype
figure in the Balabai’s paper.
Publication(s): Voichyshyn, this volume, Plate I: 2.

Zenaspis major (Balabai, 1962)

The type series was stated by P. Balabai to include four specimens (Balabai 1962: 4).
At present two specimens are identified only.
 32

Lectoty pe : BP.995 (old number 25579); contours of the shield lacking an

abdominal part; right bank of the Dniester.
Locality, age and collection data: the vicinity of Horodnytsia village (Horodenka
rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower part of the
Old-Red Formation (lower part of the Dniester Series); collector and date of collecting are
unknown.
Publication(s): Balabai 1962: 4, fig. 3; Janvier 1985: 330, fig. 19C; Voichyshyn,
this volume, Plate II: 1.
Pa ra lectotype : BP.31/1 (old number 25575/1); imprints of cornual processes of
the shield; right bank of the Dniester.
Locality, age, collection data and note: the vicinity of Horodnytsia village
(Horodenka rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower
part of the Old-Red Formation (the Dniester Series); collector and date of collecting are
unknown; the specimen has been identified due to Balabai’s record on the label.
Publication(s): Voichyshyn, this volume, Plate II: 2.

Zenaspis dzieduszyckii Voichyshyn, 2006

The type series consists of three specimens (Voichyshyn 2006: 133).

Ho loty pe : ВР.103 (old number 35631); mould and impress of the shield, lacking
a posterior part and cornual process of the right side.
Locality, age and collection data: right bank of the Dniester, the old quarry in SW
of Horodnytsia village (Horodenka rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine);
Lower Devonian, lower part of the Old-Red Formation (the Dniester Series, most probably
its Ustechko Member); collected by V. Voichyshyn on June 18, 1992.
Publication(s): Voichyshyn 2006: 134, fig. 3.
Pa raty pe :ВР.1215 (old number 37796); contour of the shield in ventral view,
lacking the cornual process of the right side.
Locality, age and collection data: right bank of the Dniester, the vicinity of
Horodnytsia village (Horodenka rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine);
Lower Devonian, lower part of the Old-Red Formation (the Dniester Series, most probably
its Ustechko Member); collected by W. Zych.
Publication(s): Voichyshyn 2006: 133.
Pa raty pe : ВР.1193; partial mould of the shield, lacking its contours and the right
side; right bank of the Dniester.
Locality, age and collection data: the vicinity of Horodnytsia village (Horodenka
rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower part of the
Old-Red Formation (the Dniester Series, most probably its Ustechko Member); collected by
M. Solodkyi in July 2004.
Publication(s): Voichyshyn 2006: 133.

Genus Zychaspis Janvier, 1985

Zychaspis elegans (Balabai, 1962)

The type series was stated by P. Balabai to include one specimen (Balabai 1962: 8).
 33

Ho loty pe :BP.41 (old number 17329); partial contours of the shield in ventral
view.
Locality, age and collection data: left bank of the Dniester, the vicinity of
Khmeleva village (Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); lower part of
the Old-Red Formation (the Dniester Series, its Khmeleva Member); very probably
collected by W. Zych in 1935.
Publication(s): Balabai 1962: 8, fig. 10; Janvier 1985: 318, fig. 7A; Voichyshyn,
this volume, Plate III: 2.

Zychaspis granulata Voichyshyn, 1998

The type series consists of one specimen (Voichyshyn 1998: 27).

Ho loty pe :BP.49 (old number 35899); imprint of dorsal side of the shield, lacking
distal part of cornual processes, imprint of fragmentary contours of abdominal part of the
shield in ventral view, and fragmentary mould of abdominal part of the shield in dorsal
view.
Image: Plate III: 1A-B.
Locality, age and collection data: left bank of the Dniester, the outcrop in NW of
Ustechko village (Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower
Devonian, lower part of the Old-Red Formation (the Dniester Series, lowest part of its
Khmeleva Member); collected by V. Voichyshyn in July 1990.
Publication(s): Voichyshyn 1998: 27, table, figs. 1-2.

Zychaspis concinna Voichyshyn et Solodkyi, 2004

The type series consists of three specimens those are remains of three individuals
preserved on one rock fragment (Voichyshyn et al. 2004: 173). Thus, all of them share the
next data: right bank of the Dniester, a quarry in the vicinity of Horodnytsia village
(Horodenka rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine); Lower Devonian, lower
part of the Old-Red Formation (the Dniester Series); collected by S. Solodkyi in 2003.

Ho loty pe : BP.966/1; almost complete imprint of the shield, lacking a distal part
of left cornual process.
Image: Plate II: 2.
Publication(s): Voichyshyn et Solodkyi 2004: 174, fig. 1.
Pa raty pe : BP.966/2; an imprint of the cornual process and of a fragment of the
shield abdominal part.
Publication(s): Voichyshyn et Solodkyi 2004: 173.
Pa ra lty pe : BP.966/3; imprint of a shield fragment in orbit region.
Publication(s): Voichyshyn et Solodkyi 2004: 173.

Zenaspidida indet.

“Cephalaspis” microlepidota Balabai, 1962

 34

The type series was stated by P. Balabai to include four specimens (Balabai 1962: 6).
As it has been noted above the specimens BP.34 (17318) and BP.35 (17323) originally
labelled by the author of the species as “Cephalaspis podolica sp. n.” then have been
redefined by him as “C. microlepidota”. In such circumstances it is not sure what type
series should accommodate these specimens in Balabai’s meaning.

Lectoty pe : BP.989 (old number 25564; in the Balabai’s paper its old register
number was wrongly marked as 17323 and locality as Ustechko); mould of shield from
dorsal side (lacking the right cornual process and dorsomedial crest) with remains of trunk
segments.
Locality, age and collection data: right bank of the Dniester, the vicinity of
Horodnytsia village (Horodenka rayon, Ivano-Frankivs’k oblast’ (Podolia), Ukraine);
Lower Devonian, lower part of the Old-Red Formation (the Dniester Series); collector and
date of collecting are unknown.
Publication(s): Balabai 1962: 6, fig. 8 ; Afanassieva et al. 1991: 69, table IX, figs. 3-4.

Osteostraci indet.

Genus Wladysagitta Voichyshyn, 2006

Wladysagitta janvieri Voichyshyn, 2006

The type series consists of three specimens, including British Museum (Natural
History) P.20508 (Voichyshyn 2006: 138).

Ho loty pe : BP.102 (old number 37523); fragmentary natural mould of a shield

from the dorsal side and its imprint.
Locality, age and collection data: left bank of the Dniester, the outcrop in NW of
Ustechko village (Zalishchyky rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower
Devonian, the Old-Red Formation (the Dniester Series, lowest part of its Khmeleva
Member); collected by V. Voichyshyn on July 26, 1994.
Publication(s): Voichyshyn 2006: 137, fig. 7A.
Pa raty pe : BP.100 (old number 37792); a fragment of the shield, exposed in
ventral view.
Locality, age and collection data: the same locality and horizon as of the holotype;
collected by V. Voichyshyn in July 1990.
Publication(s): Voichyshyn 2006: 137, fig. 7B.

"Cephalaspis" djurinensis Balabai, 1962

The type series was stated by P. Balabai to include two specimens, but in fact they
are presented by the mould and the imprint of the shield of a single individual (Voichyshyn
this volume). Therefore, the question is about one specimen representing the holotype.

Ho loty pe : BP.42 (old numbers 25563 and 25565); an incomplete mould and
imprint of the shield.
 35

Locality, age and collection data: left bank of the Dzhuryn River, the vicinity
(possibly, an old quarry) of Buriakivka village (Zalishchyky rayon, Ternopil’ oblast’
(Podolia), Ukraine); Lower Devonian, the Old-Red Formation (the Dniester Series);
collected by P. Balabai on September 5, 1955.
Publication(s): Balabai 1962: 6, fig. 7; Janvier 1985: 318, fig. 7B; Voichyshyn, this
volume, Plate III: 1A-B.

"Cephalaspis" feliciae Balabai, 1962

The type series consisted of one specimen (Balabai 1962: 4).

Ho loty pe :BP.991 (old number 17331); contours of the shield lacking the pectoral
sinuses and an abdominal part.
Locality, age and collection data: the vicinities of Bily Potik village (Chortkiv
rayon, Ternopil’ oblast’ (Podolia), Ukraine); Lower Devonian, probably, the lowest part of
the Old-Red Formation (the Dniester Series), or the uppermost part of Ivanie horizon;
collected by W. Zych.
Publication(s): Balabai 1962: 4, fig. 2; Janvier 1985: 330, fig. 19B; Voichyshyn,
this volume, Plate III: 3.

"Cephalaspis" sjostremi Balabai, 1962

The type series was stated by P. Balabai to include five specimens (Balabai 1962: 5).
At present only two of them are believed to be identified. The specimen BP.200 (old
number 17333; mould of the shield poorly preserved) from Ustechko was labelled by
W. Zych as “C. sjöstremi n. sp.”, later P. Balabai labelled it as “C. brevicornis sp.n.”.
Although P. Balabai has mentioned the last species name in his paper on pteraspid fauna
(Balabai 1959: 89, footnote), this species has never been designated by him, possibly
because the species with the same name had been established earlier (Cephalaspis
brevicornis Stensiö, 1927).

Sy ntyp e : BP.992 (old number 17325); incomplete contour of the shield;

Locality, age and collection data: the vicinities (on Zych’s label it had been marked
“E II”) of Rizdviany village (Terebovlia rayon, Ternopil’ oblast’ (Podolia), Ukraine);
Lower Devonian, probably, uppermost part of the Ivanie horizon or lowest part of the Old-
Red Formation (Ustechko Member of the Dniester Series); collected by W. Zych in 1936.
Publication(s): Balabai 1962: 5, fig. 5; Janvier 1985: 331, fig. 19D; Voichyshyn, this
volume, Plate III: 4.
Sy ntyp e : BP.994 (old number 17327); incomplete contour of the shield.
Locality, age and collection data: the same as of previous (BP.992) specimen.
Publication(s): Balabai 1962: 5, fig. 5; Janvier 1985: 331, fig. 19D; Voichyshyn, this
volume, Plate III: 5.
 36

Acknowledgments
I wish to express my gratitude to Professor Hubert Szaniawski (Institute of
Palaeobiology, Polish Academy of Science, Warsaw) for the all-round aid in searching
traces of Zych’s collection in Poland. I am very obliged to Mr. Piotr Szrek for placing to
my disposal the collections of Podolian agnathans of the Museum of the Earth (Warsaw)
and of the Institute of Geology, Warsaw University, to my colleague Iren Konovalova
(SMNH, Lviv) for help in English version of the paper.

Plate I

Fig. 1. Alaeckaspis ustetchkiensis Voichyshyn, 1999, holotype, SMNH ПЗХ-ВР.168,

incomplete mould (A) and imprint (B) of carapace.
Fig. 2. Alaeckaspis verbivciensis Voichyshyn, 1999, holotype, SMNH ПЗХ-ВР.167/1,
almost complete carapace.
Fig. 3. Pavloaspis pasternaki Voichyshyn, 1999, holotype, SMNH ПЗХ-ВР.169,
incomplete imprint of carapace.
Scale bars in all plates equal 10 mm.

Plate II

Fig. 1. Pattenaspis rogalai (Balabai, 1962), paralectotype, SMNH ПЗХ-ВР.33,

fragmentary mould (A) and imprint (B) of shield.
Fig. 2. Zychaspis concinna Voichyshyn et Solodkyi, 2004, holotype, SMNH ПЗХ-
ВР.966/1, almost complete imprint of the shield.
Fig. 3. Mimetaspis concordis Voichyshyn, 1994, holotype, SMNH ПЗХ-ВР.982,
mould of the shield.
Fig. 4. Mimetaspis concordis Voichyshyn, 1994, paratype, SMNH ПЗХ-ВР.983,
imprint of the shield.
Fig. 5. Mimetaspis concordis Voichyshyn, 1994, paratype, SMNH ПЗХ-ВР.985,
fragmentary mould of the shield.

Plate III

Fig. 1. Zychaspis granulata Voichyshyn, 1998, holotype, SMNH ПЗХ-ВР.49, imprint of

dorsal side of the shield (A) and the same with fragmentary mould of abdominal part (B).
 37

Plate I

2 3
 38

Plate II

1B
1A

5
 39

Plate III

1A

1B
 40

References

Afanassieva O.B. and Voichyshyn V.K. On the revision of the Podolian osteostracans (Agnatha) //
Palaeontologičeskij žurnal, 1991, 2: 65-72. [in Russian]
Balabai P.P. On the question of vertical distribution of pteraspids of Podolian Old-Red //
Geologičeskij sbornik Lvovskogo Geologičeskogo Obŝestva, 1957, 4: 305-309. [in Russian]
Balabai P.P. On the fauna of the cephalaspids of Podolian plate // Naukovi zapysky Naukovo-
pryrodoznavčogo muzeû AN URSR, 1962 10: 3-8. [in Ukrainian]
Blieck A. Les Hétérostracés Pteraspidiformes. Systématique, phylogénie, biostratigraphie,
biogéographie. Paris: CNRS, 1984. 199 p.
International Code of Zoological Nomenclature. Fourth edition. Sankt-Petersburg, 2000,
222 p. [in Russian]
Janvier P. Preliminary description of Lower Devonian Osteostraci from Podolia (Ukrainian S.S.R.) //
Bulletin British Museum (Natural History) Geol., 1985, 38(5): 309-334.
Novitskaya L.I. The oldest agnathans of the USSR. Heterostracans: cyathaspids, amphiaspids,
pteraspids. Moscow, Nauka, 1986. 160 p. [in Russian]
Pasternak S.I., Levytskyi V.T. The monographical collections of palaeontological assemblage of
Scientific Natural History Museum of AN USSR. Kyiv, ed. AN USSR, 1963, 36 p. [in Ukrainian]
Voichyshyn V.K. New species of the genus Mimetaspis (Agnatha) from the Lower Devonian of
Podolia // Palaeontologičnyj zbirnyk, 1994, 30: 19-24. [in Ukrainian]
Voichyshyn V.K. New species of the genus Zychaspis (Agnatha) from the Dniester Series (Early
Devonian) of Podolia // Palaeontologičnyj zbirnyk, 1998, 32: 25-29. [in Ukrainian]
Voichyshyn V.K. The new forms of pteraspids (Agnatha, Heterostraci) from Podolian Early
Devonian // Vestnik zoologii, 1999, 33 (3): 47-56.
Voichyshyn V. and Solodkyi S. New materials concerning the genus Zychaspis (Agnatha,
Osteostraci) from Horodnytsia (Podolia, Ukraine) // Naukovi zapysky Deržavnogo pryrodoznavčogo
muzeû, 2004, 19: 171-182. [in Ukrainian]
Voichyshyn V. New osteostracans from the Lower Devonian terrigenous deposits of Podolia,
Ukraine // Acta Palaeontologica Polonica, 2006, 51(1): 131-142.
Voichyshyn V. On the revision of Balabai’s collection of Early Devonian Osteostraci (Agnatha)
from Podolia // Scientific collections of the State Natural History Museum. Name-bearing types and
type series (I), 2006, issue 2: 9-25.
Zych W. Fauna ryb dewonu i downtonu Podola. Pteraspidomorphi: Heterostraci. Lwów, 1931. Cz. I.
A. 92 s.

Department of Taxonomy of Modern and Fossil Biota, State Museum of Natural History
NASU, Teatralna Str. 18, 79008, Lviv, Ukraine, e-mail: victor@museum.lviv.net
 107

Анотації

Палеозоологічний фонд
Диденко Ю.В. Новые виды остракод (Arthropoda: Ostracoda) из верхнемеловых
отложений Волыно-Подолья // Научные коллекции Государственного
природоведческого музея / Годунько Р.Й., Войчишин В.К., Климишин А.С. и др.
Номенклатурные типы и типовые серии (1). Вып. 2. – Львов, 2006. – С. 4-8.
Приводится монографическое описание трех новых видов остракод,
относящихся к семействам Veeniidae и Progonocytheridae из отложений верхнего мела
Волыно-Подолья. Для описанных видов характерно узкое стратиграфическое
распространение.
Didenko, J. New species of Ostracods (Arthropoda: Ostracoda) from the Upper
Cretaceous deposits of Volyno-Podolia // Scientific collections of the State Natural
History Museum / Godunko R.J., Voichyshyn V.K., Klymyshyn O.S. et al. Name-bearing
types and type series (1). Issue 2. – Lviv, 2006. – P. 4-8.
Monographic description of three new Ostracoda species belonging to families
Veeniidae and Progonocytheridae from the Upper Cretaceous deposits of Volyno-Podolia is
given. New species are characterized by a limited stratigraphical distribution.

Войчишин В.К. До ревізії колекції ранньодевонських остеостраків (Agnatha)

П. Балабая з Поділля (Україна) // Наукові колекції Державного природознавчого
музею / Годунько Р.Й., Войчишин В.К., Климишин О.С. та ін. Номенклатурні типи і
типові серії (1). Вип. 2. – Львів, 2006. – С. 9-25.
Дана робота завершує ревізію монографічної колекції ранньодевонських
остеостраків П. Балабая з Поділля, започатковану у статті О.Б. Афанасьєвої і
В.К. Войчишина (1991). Крім раніше переописаних видів Pattenaspis rogalai (Balabai,
1962) та “Cephalaspis” microlepidota Balabai, 1962, інші два види Zenaspis podolica
(Balabai, 1962) і Zychaspis elegans (Balabai, 1962) можуть вважатися такими, що
дійсно існували. З огляду на те, чим представлений типовий матеріал Zenaspis major
(Balabai, 1962), ця видова назва може виявитись синонімом Zenaspis podolica.
“Cephalaspis” djurinensis Balabai, 1962 теж розглядається як окремий вид, що,
ймовірно, належить до невідомого роду. Обидві видові назви “Cephalaspis” feliciae
Balabai, 1962 і "C." sjostremi Balabai, 1962 слід вважати nomina dubia.
Войчишин В.К. К ревизии коллекции раннедевонских остеостраков (Agnatha)
П. Балабая из Подолья (Украина) // Научные коллекции Государственного
природоведческого музея / Годунько Р.Й., Войчишин В.К., Климишин А.С. и др.
Номенклатурные типы и типовые серии (1). Вып. 2. – Львов, 2006. – С. 9-25.
Настоящая работа завершает ревизию монографической коллекции
раннедевонских остеостраков П. Балабая из Подолья, начатую в статье
О.Б Афанасьєвой и В.К. Войчишина (1991). Кроме ранее переописанных видов
Pattenaspis rogalai (Balabai, 1962) и “Cephalaspis” microlepidota Balabai, 1962, два
других вида Zenaspis podolica (Balabai, 1962) и Zychaspis elegans (Balabai, 1962) могут
считаться реально существовавшими. Исходя из того, чем представлен типовой
материал Zenaspis major (Balabai, 1962), это видовое название может оказаться
 108

синонимом Zenaspis podolica. “Cephalaspis” djurinensis Balabai, 1962 также

рассматривается как отдельный вид, который вероятно входит в состав неизвестного
рода. Оба видовых названия “Cephalaspis” feliciae Balabai, 1962 и "C." sjostremi
Balabai, 1962 следует считать nomina dubia.
Voichyshyn, V. On the revision of P. Balabai’s collection of Early Devonian Osteostraci
(Agnatha) from Podolia (Ukraine) // Scientific collections of the State Natural History
Museum / Godunko R.J., Voichyshyn V.K., Klymyshyn O.S. et al. Name-bearing types and
type series (1). Issue 2. – Lviv, 2006. – P. 9-25.
Present work is a result of the revision of the P. Balabai’s monographic collection of
Early Devonian osteostracans from Podolia which begin by Afanassieva and Voichyshyn in
1991. Besides the earlier redescribed species Pattenaspis rogalai (Balabai, 1962) and
“Cephalaspis” microlepidota Balabai, 1962, another two species Zenaspis podolica
(Balabai, 1962) and Zychaspis elegans (Balabai, 1962) should be considered as those
existed in fact. Proceeding from the type material on Zenaspis major (Balabai, 1962) it is
likely that the species name is synonymous to Zenaspis podolica. “Cephalaspis”
djurinensis Balabai, 1962 also is regarded as separate species, probably belonging to an
unknown genus. Both species names “Cephalaspis” feliciae Balabai, 1962 and
"C." sjostremi Balabai, 1962 should be considered as nomina dubia.

Войчишин В. Номенклатурні типи і типові серії викопних безщелепних (Agnatha:

Heterostraci, Osteostraci) // Наукові колекції Державного природознавчого музею /
Годунько Р.Й., Войчишин В.К., Климишин О.С. та ін. Номенклатурні типи і типові
серії (1). Вип. 2. – Львів, 2006. – С. 26-40.
Колекція “Викопні риби” Державного природознавчого музею НАН України
(Львів) містить номенклатурні типи наступних ранньодевонських видів: Alaeckaspis
ustetchkiensis Voichyshyn, 1999, A. verbivciensis Voichyshyn, 1999, Pavloaspis
pasternaki Voichyshyn, 1999, Mimetaspis concordis Voichyshyn, 1994, Pattenaspis
rogalai (Balabai, 1962), Zenaspis podolica (Balabai, 1962), Z. major (Balabai, 1962),
Z. dzieduszyckii Voichyshyn, 2006, Zychaspis elegans (Balabai, 1962), Z. granulata
Voichyshyn, 1998, Z. concinna Voichyshyn et Solodkyi, 2004, Wladysagitta janvieri
Voichyshyn, 2006, “Cephalaspis” microlepidota Balabai, 1962, "C." djurinensis Balabai,
1962, "C." feliciae Balabai, 1962 та "C." sjostremi Balabai, 1962. Колекція нараховує 42
типові зразки, а саме 11 голотипів, 4 лектотипи, 20 паратипів, 5 паралектотипів і 2
синтипи. Наводиться характеристика кожного зразка, включаючи інформацію про
місце збору, стратиграфічний горизонт, автора та дату знахідки.
Войчишин В.К. Номенклатурные типы и типовые серии ископаемых
бесчелюстных (Agnatha: Heterostraci, Osteostraci) // Научные коллекции
Государственного природоведческого музея / Годунько Р.Й., Войчишин В.К.,
Климишин А.С. и др. Номенклатурные типы и типовые серии (1). Вып. 2. – Львов,
2006. – С. 26-40.
Коллекция “Ископаемые рыбы” Государственного природоведческого музея
НАН Украины (Львов) содержит номенклатурные типы следующих раннедевонских
видов: Alaeckaspis ustetchkiensis Voichyshyn, 1999, A. verbivciensis Voichyshyn, 1999,
Pavloaspis pasternaki Voichyshyn, 1999, Mimetaspis concordis Voichyshyn, 1994,
Pattenaspis rogalai (Balabai, 1962), Zenaspis podolica (Balabai, 1962), Z. major (Balabai,
1962), Z. dzieduszyckii Voichyshyn, 2006, Zychaspis elegans (Balabai, 1962), Z. granulata
 109

Voichyshyn, 1998, Z. concinna Voichyshyn et Solodkyi, 2004, Wladysagitta janvieri

Voichyshyn, 2006, “Cephalaspis” microlepidota Balabai, 1962, "C." djurinensis Balabai,
1962, "C." feliciae Balabai, 1962 и "C." sjostremi Balabai, 1962. Коллекция насчитывает
42 типовых образца, а именно 11 голотипов, 4 лектотипа, 20 паратипов, 5
паралектотипов и 2 синтипа. Приводится характеристика каждого образца, включая
информацию о месте сбора, стратиграфическом горизонте, авторе и дате сбора.
Voichyshyn, V. Name-bearing types and type series of Fossil agnathans (Agnatha:
Heterostraci, Osteostraci) // Scientific collections of the State Natural History Museum /
Godunko R.J., Voichyshyn V.K., Klymyshyn O.S. et al. Name-bearing types and type series
(1). Issue 2. – Lviv, 2006. – P. 26-40.
The collection of Fossil Fishes of the State Museum of Natural History NASU (Lviv)
contains name-bearing types of following Early Devonian species: Alaeckaspis
ustetchkiensis Voichyshyn, 1999, A. verbivciensis Voichyshyn, 1999, Pavloaspis
pasternaki Voichyshyn, 1999, Mimetaspis concordis Voichyshyn, 1994, Pattenaspis
rogalai (Balabai, 1962), Zenaspis podolica (Balabai, 1962), Z. major (Balabai, 1962),
Z. dzieduszyckii Voichyshyn, 2006, Zychaspis elegans (Balabai, 1962), Z. granulata
Voichyshyn, 1998, Z. concinna Voichyshyn et Solodkyi, 2004, Wladysagitta janvieri
Voichyshyn, 2006, “Cephalaspis” microlepidota Balabai, 1962, "C." djurinensis Balabai,
1962, "C." feliciae Balabai, 1962 and "C." sjostremi Balabai, 1962. There are 42 type
specimens in the collection, namely 11 holotypes, 4 lectotypes, 20 paratypes, 5
paralectotypes and 2 syntypes. Description of an every specimen, including information on
locality, horizon, author and date of its collecting is given.

Палеоботанічний фонд
Мамчур А.П. Типы видов ископаемых растений // Научные коллекции
Государственного природоведческого музея / Годунько Р.Й., Войчишин В.К.,
Климишин А.С. и др. Номенклатурные типы и типовые серии (1). Вып. 2. – Львов,
2006. – С. 41-49.
В музее хранятся образцы с типами 12 видов и 1 вариетета ископаемых растений
из мелового и неогенового периодов. Каталог иллюстрирован фотоизображениями.
Mamtchur, A. The types of fossil plant species // Scientific collections of the State Natural
History Museum / Godunko R.J., Voichyshyn V.K., Klymyshyn O.S. et al. Name-bearing
types and type series (1). Issue 2. – Lviv, 2006. – P. 41-49.
At present the type specimens of 12 species and 1 variety of fossil plants from
Cretaceous and Neogene are kept in the museum collection. The images of type specimens
are given.

Ботанічний фонд
Климишин А.С., Кузярин А.Т. Типовой материал в Гербарии сосудистых растений
(LWS) // Научные коллекции Государственного природоведческого музея / Годунько
Р.Й., Войчишин В.К., Климишин А.С. и др. Номенклатурные типы и типовые серии
(1). Вып. 2. – Львов, 2006. – С. 50-61.