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a
Institute of Food Science and Technology, Chinese Academy of Agricultural Sciences (CAAS), Beijing 100193, China
b
Department of Food Science and Engineering, Jilin Agricultural University, Changchun 130118, China
c
Division of Soybean Processing, Soybean Research & Development Center, Chinese Agricultural Research System, Changchun 130118, China
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Fig. 1 /GVCDQNKUOQH&(KPJWOCPDQF[CPFFKUGCUGRTGXGPVKQP
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10 3
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Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10
Table 2
'HHGEVQH&(KPUQOGV[RGUQHECPEGT
6[RGUQHECPEGT #PCN[VKECNOGVJQFU 'ZRGTKOGPVCNOQFGN /GEJCPKUO 4GHGTGPEGU
1XCTKCPECPEGT /GVCCPCN[UKU %NKPKECNTGUGCTEJGUQXCTKCPECPEGT 4GFWEGEQPEGPVTCVKQPUQHEKTEWNCVKPIJQTOQPCN =?
2CPETGCVKEECPEGT /GVCCPCN[UKU %NKPKECNTGUGCTEJGURCPETGCVKEECPEGT GUVTQIGPJQTOQPCNRCPETGCVKEECTEKPQIGPGUKU =?
2TQUVCVGECPEGT 2TQURGEVKXGCPCN[UKU %NKPKECNTGUGCTEJGURTQUVCVGECPEGT GUVTQIGPUQTCPFTQIGPU =?
$TGCUVECPEGT %CUGEQPVTQNUVWF[ %NKPKECNTGUGCTEJGUYQOGPDTGCUVECPEGT =?
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%QNQTGEVCNECPEGT 1XGTXKGYQHGZRGTKOGPVCNFCVC %NKPKECNTGUGCTEJGUCPFCPKOCNGZRGTKOGPV =?
acids
*GCFCPFPGEMECPEGT %CUGEQPVTQNUVWFKGU %NKPKECNTGUGCTEJGUJGCFCPFPGEMECPEGT =?
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10
Gut microbiota
DF Fermentation Feaces
Satiety
SCFAs L cell
PYY
GLP -1
Intestinal disease
Fig. 2 6JGGHHGEVQH&(QPRTGXGPVKQPCPFVTGCVOGPVFKUGCUGUKUTGNCVGFYKVJVJG5%(#U
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10
Table 3
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%NCUUKſECVKQP 2TQFWEVU (KDGTV[RG &(5QWTEGU 2J[UKEQEJGOKECNRTQRGTVKGU (WPEVKQPCN 4GHGTGPEGU
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VQFGVGTOKPGVJGQRVKOCNKPVCMGQHFKHHGTGPVV[RGU#HVGTVJGCDQXG JVVRU//FQKQTI//LENPW
RTQDNGOUCTGUQNXGF&(YKNNDGOQTGEQPFWEKXGVQVJGRTGXGPVKQP =? ,#4QDGTVUQP(&&/QPTGFQP2&[UUGNGTGVCN*[FTCVKQPRTQRGTVKGUQH
CPFCNNGXKCVKQPQHJWOCPFKUGCUGUYJKNGRTQOQVKPIVJGFGXGNQROGPV FKGVCT[ſDTGCPFTGUKUVCPVUVCTEJC'WTQRGCPEQNNCDQTCVKXGUVWF[.96(QQF
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Acknowledgments =? *% %JCQ /9 .CK /5 -QPI GV CN %WVQHH XQNWOG QH FKGVCT[ HKDGT
VQ COGNKQTCVG EQPUVKRCVKQP KP EJKNFTGP , 2GFKCVT
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QH /1( CPF /#4# %#45 #PF YG CNUQ VJCPM VJG 0CVKQPCN =? ,'.CODGTV,#2CTPGNN,/6WPPKENKHHGGVCN%QPUWOKPI[GNNQYRGC
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QH %JKPC CPF VJG -G[ .CDQTCVQT[ HQT #ITKEWNVWTCN 2TQFWEV &GGR JVVRU//FQKQTI//LENPW
2TQEGUUKPIHQTUWRRQTV =? 9,&CJN/.5VGYCTV2QUKVKQPQHVJGCECFGO[QHPWVTKVKQPCPFFKGVGVKEU
JGCNVJ KORNKECVKQPU QH FKGVCT[ HKDGT , #ECF 0WVT &KGV
References JVVRU//FQKQTI//LLCFC
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=? /,)KFNG[ )';CMWDQX(WPEVKQPCN ECVGIQTKUCVKQPQHFKGVCT[HKDTG KP HNGUJGF RQVCVQGU Heimeiren YKVJ JKIJ J[FTQUVCVKE RTGUUWTG CPF JKIJ
HQQFUDG[QPFµVROXEOH¶XU‘KPUQNWDNG¶6TGPFU(QQF5EK6GEJ RTGUUWTGJQOQIGPK\CVKQPRTQEGUUKPICEQORCTCVKXGUVWF[+PPQX(QQF5EK
JVVRU//FQKQTI//LVKHU 'OGTI JVVRU//FQKQTI//LKHUGV
=? #1NUQP)/)TC[/%%JKWGVCN%JGOKUVT[CPFCPCN[UKUQHUQNWDNG =? * %JGP % <JCQ , .K GV CN 'HHGEVU QH GZVTWUKQP QP UVTWEVWTCN CPF
FKGVCT[ſDGT(QQF6GEJ RJ[UKEQEJGOKECN RTQRGTVKGU QH UQNWDNG FKGVCT[ HKDGT HTQO PQFGU QH NQVWU
=? /# 'CUVYQQF 6JG RJ[UKQNQIKECN GHHGEVU QH FKGVCT[ HKDGT CP WRFCVG TQQV.96(QQF5EK6GEJPQN JVVRU//FQKQTI//
#PPW 4GX 0WVT JVVRU//FQKQTI//CPPWTGX LNYV
PW =? )/-JCP 0/ -JCP <7 -JCP GV CN 'HHGEV QH GZVTCEVKQP OGVJQFU
=? ,/.CVVKOGT/&*CWD'HHGEVUQHFKGVCT[ſDGTCPFKVUEQORQPGPVUQP QP UVTWEVWTCN RJ[UKQEJGOKECN CPF HWPEVKQPCN RTQRGTVKGU QH FKGVCT[ HKDGT
OGVCDQNKEJGCNVJ0WVTKGPVU JVVRU//FQKQTI// HTQOFGHCVVGFYCNPWVƀQWT(QQF5EK$KQVGEJPQN
PW JVVRU//FQKQTI//U
=? 9)69KNNCVU,2-PQZ,&/KMMGNUGP2GEVKPPGYKPUKIJVUKPVQCPQNF =? );W,$GK,<JCQGVCN/QFKſECVKQPQHECTTQV Daucus carota.KPP
RQN[OGTCTGUVCTVKPIVQIGN6TGPFU(QQF5EK6GEJ XCT5CVKXC*QHHO RQOCEGKPUQNWDNGFKGVCT[HKDGTYKVJEQORNGZGP\[OG
JVVRU//FQKQTI//LVKHU OGVJQFWNVTCſPGEQOOKPWVKQPCPFJKIJJ[FTQUVCVKERTGUUWTG(QQF%JGO
=? ,.&QWDNKGT2,9QQF4JGQNQIKECNRTQRGTVKGUQHCSWGQWUUQNWVKQPUQH JVVRU//FQKQTI//LHQQFEJGO
DGVCDINWECPHTQOQCVU Avena sativa. %GTGCN%JGO =? ) <JCQ 4 <JCPI . &QPI GV CN 2CTVKENG UK\G QH KPUQNWDNG FKGVCT[
HKDGT HTQO TKEG DTCP CHHGEVU KVU RJGPQNKE RTQHKNG DKQCEEGUUKDKNKV[ CPF
=? -/&CXKUQP0,6GORNG%GTGCNHKDGTHTWKVHKDGTCPFV[RGFKCDGVGU HWPEVKQPCN RTQRGTVKGU .96(QQF 5EK 6GEJPQN
GZRNCKPKPI VJG RCTCFQZ , &KCDGVGU %QORNKECV JVVRU//FQKQTI//LNYV
JVVRU//FQKQTI//LLFKCEQOR =? $/CVUWJKTQ.'.KNNQ%5¶GP\GVCN%JGOKECNEJCTCEVGTK\CVKQPQHVJG
=? #**KVG0WVTKVKQPCNGRKFGOKQNQI[QHEJTQPKEFKUGCUGCPFFGſPKPI³+HDOWK\ OWEKNCIGHTQOHTWKVUQH2SXQWLD¿FXVLQGLFD%CTDQJ[F2QN[O
Diet”)NQD(QQF5GE JVVRU//FQKQTI// JVVRU//FQKQTI//LECTDRQN
=? 22GGTCLKV0%JKGYEJCP5&GXCJCUVKP'HHGEVUQHRTGVTGCVOGPVOGVJQFUQP
=? &5 .WFYKI ($ *W . 6CRR[ GV CN &KGVCT[ ECTDQJ[FTCVGU TQNG JGCNVJTGNCVGFHWPEVKQPCNRTQRGTVKGUQHJKIJFKGVCT[ſDGTRQYFGTHTQONKOG
QH SWCNKV[ CPF SWCPVKV[ KP EJTQPKE FKUGCUG $/, TGUKFWGU (QQF %JGO JVVRU//FQKQTI//
JVVRU//FQKQTI//DOLM LHQQFEJGO
=? , ;W / #JOGFPC (WPEVKQPCN EQORQPGPVU QH ITCRG RQOCEG VJGKT =? /5 /CHK /# $TKEM ' &KOCU GV CN )GPGVKE CTEJKVGEVWTG QH FKGVCT[
EQORQUKVKQP DKQNQIKECN RTQRGTVKGU CPF RQVGPVKCN CRRNKECVKQPU +PV , HKDGT CPF QNKIQUCEEJCTKFG EQPVGPV KP C OKFFNG #OGTKECP RCPGN QH
(QQF 5EK 6GEJ JVVRU//FQKQTI//L GFKDNG FT[ DGCP 2NCPV )GPQOG JVVRU//FQKQTI//
Z RNCPVIGPQOG
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10 9
=? .*WCPI:&KPI;<JCQGVCN/QFKſECVKQPQHKPUQNWDNGFKGVCT[ſDGT =? :*WC5:W/9CPIGVCN'HHGEVUQHJKIJURGGFJQOQIGPK\CVKQPCPF
HTQO ICTNKE UVTCY YKVJ WNVTCUQPKE VTGCVOGPV , (QQF 2TQEGUU 2TGU JKIJRTGUUWTGJQOQIGPK\CVKQPQPUVTWEVWTGQHVQOCVQTGUKFWGHKDGTU(QQF
JVVRU//FQKQTI//LHRR %JGO JVVRU//FQKQTI//LHQQFEJGO
=? . %JGPI : <JCPI ; *QPI %JCTCEVGTKUCVKQP QH RJ[UKEQEJGOKECN CPF =? *;CKEJ*)CTPC$$EJKTGVCN%JGOKECNEQORQUKVKQPCPFHWPEVKQPCN
HWPEVKQPCNRTQRGTVKGUQHUQNWDNGFKGVCT[ſDTGHTQORQVCVQRWNRQDVCKPGFD[ RTQRGTVKGUQHFKGVCT[HKDTGGZVTCEVGFD['PIN[UVCPF2TQUM[OGVJQFUHTQO
GP\[OGCUUKUVGFGZVTCEVKQP+PV,$KQN/CETQOQN VJG CNIC Ulva lactuca EQNNGEVGF KP 6WPKUKC #NICN 4GU
JVVRU//FQKQTI//LKLDKQOCE JVVRU//FQKQTI//LCNICN
=? .<JCQ(<JCPI:&KPIGVCN)WVDCEVGTKCUGNGEVKXGN[RTQOQVGFD[ =? <(GPI&9GK5#NCZKGVCN/QFKſGFUQNWDNGFKGVCT[ſDGTHTQODNCEM
FKGVCT[ſDGTUCNNGXKCVGV[RGFKCDGVGU5EKGPEG DGCP EQCVU YKVJ KVU TJGQNQIKECN CPF DKNG CEKF DKPFKPI RTQRGTVKGU (QQF
JVVRU//FQKQTI//UEKGPEGCCQ *[FTQEQNN JVVRU//FQKQTI//LHQQFJ[F
=? /2CTXKP$<CJTC-/5CLCFGVCN#RTQURGEVKXGUVWF[QHFKHHGTGPVV[RGUQH =? %(&KGFGTKEMU8#,KFGCPK2J[UKEQEJGOKECNCPFHWPEVKQPCNRTQRGTVKGUQHKPUQNWDNG
FKGVCT[ſDGTCPFTKUMQHECTFKQXCUEWNCTFKUGCUGVGJTCPNKRKFCPFINWEQUGUVWF[ FKGVCT[ſDGTKUQNCVGFHTQO$CODCTCITQWPFPWV Vigna subterranea =.?8GTFE ,
0WVTKGPVU JVVRU//FQKQTI//PW (QQF5EK JVVRU//FQKQTI//
=? +0CPFK#5GPIWRVC/)JQUJ'HHGEVUQHFKGVCT[ſDTGUGZVTCEVGFHTQO =? $&W(<JW$:W2J[UKEQEJGOKECNCPFCPVKQZKFCPVRTQRGTVKGUQHFKGVCT[
FGHCVVGF UGUCOG JWUM TKEG DTCP HNCZUGGF QP J[RGTEJQNGUVGTQOKE TCVU ſDGTUHTQO3KPIMG JWNNNGUUDCTNG[ ƀQWTCUCHHGEVGFD[WNVTCſPGITKPFKPI
$KQCEV%CTDQJ[FT&KGV(KDTG JVVRU//FQKQTI// $KQCEV%CTDQJ[FT&KGV(KDTG JVVRU//FQKQTI//
LDEFH LDEFH
=? 6 /CPFKOKMC ) 2CVWTK %'& )W\OCP GV CN 'HHGEVU QH FKGVCT[ =? %4/CTYC#5QWJKT65CDW7UGQHITGGPEJGOKUVT[OGVJQFUKPVJG
DTQEEQNKHKDTGCPFEQTPQKNQPUGTWONKRKFUHCGECNDKNGCEKFGZETGVKQPCPF GZVTCEVKQP QH FKGVCT[ HKDGTU HTQO ECEVWU TCEMGVU Opuntia ficus indica
JGRCVKE IGPG GZRTGUUKQP KP TCVU (QQF %JGO UVTWEVWTCN CPF OKETQUVTWEVWTCN UVWFKGU +PV , $NQN /CETQOQN
JVVRU//FQKQTI//LHQQFEJGO JVVRU//FQKQTI//LKLDKQOCE
=? : .WQ 3 9CPI $ <JGPI GV CN *[FTCVKQP RTQRGTVKGU CPF DKPFKPI =? ;/CNMMK-#WVKQ1*CPPKPGP1CVDTCPEQPEGPVTCVGURJ[UKECNRTQRGTVKGU
ECRCEKVKGUQHFKGVCT[ſDGTUHTQODCODQQUJQQVUJGNNCPFKVUJ[RQNKRKFGOKE QH DGVCINWECP CPF J[RQEJQNGUVGTQNGOKE GHHGEVU KP TCVU %GTGCN %JGO
G H H G E V U K P O K E G ( Q Q F % J G O 6 Q Z K E Q N
JVVRU//FQKQTI//LHEV =? ,5.X:;.KW:2<JCPI%JGOKECNEQORQUKVKQPCPFHWPEVKQPCNEJCTCEVGTKUVKEU
=? %.KW:..KP<9CPGVCN6JGRJ[UKEQEJGOKECNRTQRGTVKGU in vitro QHFKGVCT[ſDGTTKEJRQYFGTQDVCKPGFHTQOEQTGQHOCK\GUVTCY(QQF%JGO
DKPFKPI ECRCEKVKGU CPF in vivo J[RQEJQNGUVGTQNGOKE CEVKXKV[ QH UQNWDNG JVVRU//FQKQTI//LHQQFEJGO
FKGVCT[ſDGTGZVTCEVGFHTQOUQ[JWNNU(QQF(WPEV =? 0*WUUCKP$#2#IWU#</&CNKGVCN&GVGTOKPCVKQPQHHWPEVKQPCN
JVVRU//FQKQTI//%(1( RTQRGTVKGUQHEQEQCYCUVGHTQOEQPEGPVTCVGFEQEQCFTKPM,(QQF/GCU%JCTCEV
=? 5%JCPI:%WK/)WQGVCN+PUQNWDNGFKGVCT[ſDGTHTQORGCTRQOCEG JVVRU//FQKQTI//U
ECPRTGXGPVJKIJHCVFKGVKPFWEGFQDGUKV[KPTCVUOCKPN[D[KORTQXKPIVJG =? 0%& 4KUM (CEVQT %QNNCDQTCVKQP 0%&4KU% 6TGPFU KP CFWNV DQF[
UVTWEVWTG QH VJG IWV OKETQDKQVC , /KETQDKQN $KQVGEJPQN OCUUKPFGZKPEQWPVTKGUHTQOVQCRQQNGFCPCN[UKUQH
ŌJVVRU//FQKQTI//LOD RQRWNCVKQPDCUGF OGCUWTGOGPV UVWFKGU YKVJ OKNNKQP RCTVKEKRCPVU
=? && )CNNCJGT -, 9QQF %/ )CNNCJGT GV CN +PVGUVKPCN EQPVGPVU 6JG .CPEGV JVVRU//FQKQTI//5
UWRGTPCVCPVXKUEQUKV[QHTCVUHGFQCVDCUGFOWHſPUCPFEGTGCNRTQFWEVU%GTGCN :
%JGO JVVRU//FQKQTI//%%*'/ =? '5 /QWUVCHC 2 (TQIWGN (TQO QDGUKV[ IGPGVKEU VQ VJG HWVWTG QH
=? + 7NNCJ 6 ;KP 5 :KQPI GV CN 'HHGEVU QH VJGTOCN RTGVTGCVOGPV QP RGTUQPCNK\GFQDGUKV[VJGTCR[0CV4GX'PFQETKPQN
RJ[UKEQEJGOKECNRTQRGTVKGUQHPCPQUK\GFQMCTC UQ[DGCPTGUKFWG KPUQNWDNG JVVRU//FQKQTI//PTGPFQ
FKGVCT[ſDGTRTGRCTGFD[YGVOGFKCOKNNKPI,(QQF'PI =? ($ /QTGN 3 &CK , 0K GV CN Į)CNCEVQQNKIQUCEEJCTKFGU FQUG
JVVRU//FQKQTI//LLHQQFGPI FGRGPFGPVN[TGFWEGCRRGVKVGCPFFGETGCUGKPƀCOOCVKQPKPQXGTYGKIJVCFWNVU
=? 5//QMJVCT*/5YCKNCO'5'ODCD[2J[UKEQEJGOKECNRTQRGTVKGU ,0WVT JVVRU//FQKQTI//LP
PWVTKVKQPCNXCNWGCPFVGEJPQHWPEVKQPCNRTQRGTVKGUQHIQNFGPDGTT[ Physalis =? ( +UMGP 5 -NCWU / 1UVGTJQHH GV CN 'HHGEVU QH NQPIVGTO UQNWDNG XU
peruviana YCUVGRQYFGTEQPEKUGVKVNGEQORQUKVKQPQHIQNFGPDGTT[LWKEGYCUVG KPUQNWDNGFKGVCT[ſDGTKPVCMGQPJKIJHCVFKGVKPFWEGFQDGUKV[KP%$./,
(QQF%JGO JVVRU//FQKQTI//LHQQFEJGO OKEG , 0WVT $KQEJGO JVVRU//FQKQTI//
=? 8 5WT[CPVK 6 -WUWOCPKPIUKJ ;5 4WOKPIV[CU 2J[UKEQEJGOKECN LLPWVDKQ
RTQRGTVKGUQHFKGVCT[ſDGTUHTQOArtocarpus camansKHTWKV+QR%QPHGTGPEG =? 5%JCPI:%WK/)WQGVCN+PUQNWDNGFKGVCT[ſDGTHTQORGCTRQOCEG
JVVRU//FQKQTI//:/193/1/ ECPRTGXGPVJKIJHCVFKGVKPFWEGFQDGUKV[KPTCVUOCKPN[D[KORTQXKPIVJG
=? .+ :WGNK *' :KWNK .8 ;WCPRKPI GV CN 'ZVTCEVKQP CPF HWPEVKQPCN UVTWEVWTG QH VJG IWV OKETQDKQVC , /KETQDKQN $KQVGEJPQN
RTQRGTVKGUQHYCVGTUQNWDNGFKGVCT[ſDGTHTQOCRRNGRQOCEG,(QQF2TQEGUU JVVRU//FQKQTI//LOD
'PI JVVRU//FQKQTI//LHRG =? ( .W ; .KW $ .K 1MCTC FKGVCT[ HKDGT CPF J[RQIN[EGOKE GHHGEV QH
=? # %GTFC6CRKC / FG . 2ÃTG\%JCDGNC , 2ÃTG\NXCTG\ GV CN QMCTC HQQFU $KQCEV %CTDQJ[FT &KGV (KDTG
8CNQTK\CVKQP QH RQOCEG RQYFGT QDVCKPGF HTQO PCVKXG /GZKECP CRRNG JVVRU//FQKQTI//LDEFH
(Malus domestica XCT TC[CFC EJGOKECN VGEJPQHWPEVKQPCN CPF =? #5TKEJCOTQGP8%JCXCUKVIn vitro TGVCTFCVKQPQHINWEQUGFKHHWUKQPYKVJ
CPVKQZKFCPV RTQRGTVKGU 2NCPV (QQF *WO 0WVT IWO GZVTCEVGF HTQO OCNXC PWV UGGFU RTQFWEGF KP 6JCKNCPF (QQF %JGO
JVVRU//FQKQTI//U JVVRU//FQKQTI//LHQQFEJGO
=? $6CQ;(C[KP.*CPI2JGPQNKERTQſNGCPFKPXKVTQCPVKQZKFCPVECRCEKV[ =? /#UJWVQUJ,5JKXGUJIn vitro RQUVRTCPFKCNINWEQUGNQYGTKPIGHHGEVUQHFKGVCT[
QHKPUQNWDNGFKGVCT[ſDGTRQYFGTUHTQOEKVTWU Citrus junos5KGDGZ6CPCMC HKDGTUKUQNCVGFHTQOTamarindus indicaCPFCassia fistulaUGGFU#OGTKECP,
RQOCEGCUCHHGEVGFD[WNVTCſPGITKPFKPI,#IT(QQF%JGO (QQF6GEJ JVVRU//FQKQTI//CLHV
JVVRU//FQKQTI//LHD =? . %JGPI : <JCPI ; *QPI %JCTCEVGTKUCVKQP QH RJ[UKEQEJGOKECN CPF
=? ,;W/#JOGFPC(WPEVKQPCNEQORQPGPVUQHITCRGRQOCEGVJGKTEQORQUKVKQP HWPEVKQPCNRTQRGTVKGUQHUQNWDNGFKGVCT[ſDTGHTQORQVCVQRWNRQDVCKPGFD[
DKQNQIKECNRTQRGTVKGUCPFRQVGPVKCNCRRNKECVKQPU+PV,(QQF5EK6GEJ GP\[OGCUUKUVGFGZVTCEVKQP+PV,$KQN/CETQOQN
JVVRU//FQKQTI//LZ JVVRU//FQKQTI//LKLDKQOCE
=? 9 <JCPI ) <GPI ; 2CP GV CN 2TQRGTVKGU QH UQNWDNG FKGVCT[ HKDGT =? /.$CPIQWTC,0UQT#VKPFCPC-<JWGVCN2QVGPVKCNJ[RQIN[ECGOKE
RQN[UCEEJCTKFGHTQORCRC[CRGGNQDVCKPGFVJTQWIJCNMCNKPGQTWNVTCUQWPF GHHGEVU QH KPUQNWDNG HKDTGU KUQNCVGF HTQO HQZVCKN OKNNGVU =Setaria italic
CUUKUVGF CNMCNKPG GZVTCEVKQP %CTDQJ[F 2QN[O . 2 $GCWXQKU? +PV , (QQF 5EK 6GEJ
JVVRU//FQKQTI//LECTDRQN JVVRU//FQKQTI//LZ
=? ;.KW%(CP/6KCPGVCN'HHGEVQHFT[KPIOGVJQFUQPRJ[UKEQEJGOKECN =? 4/(GPI;0<QPI5/%CQGVCN%WTTGPVECPEGTUKVWCVKQPKP%JKPC
RTQRGTVKGUCPF in vitroJ[RQIN[EGOKEGHHGEVUQHQTCPIGRGGNFKGVCT[ſDGT,(QQF IQQFQTDCFPGYUHTQOVJG)NQDCN%CPEGT5VCVKUVKEU!%CPEGT%QOOWP
2TQEGUU2TGUGTX GJVVRU//FQKQTI//LHRR JVVRU//FQKQTI//U
10 Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10
=? 5% *Q 5< %JGPI %: <JCPI GV CN 'HHGEV QH FKGVCT[ HKDGT KPVCMG ſDGT,(QQF5EK6GEJ JVVRU//FQKQTI//U
QP DTGCUV ECPEGT TKUM CEEQTFKPI VQ GUVTQIGP CPF RTQIGUVGTQPG TGEGRVQT
UVCVWU 'WT , %NKP 0WVT JVVRU//FQKQTI// =? )# 5QNKOCP &KGVCT[ HKDGT CVJGTQUENGTQUKU CPF ECTFKQXCUEWNCT FKUGCUG
GLEP 0WVTKGPVU JVVRU//FQKQTI//PW
=? * :W ; &KPI : :KP GV CN &KGVCT[ HKDGT KPVCMG KU CUUQEKCVGF YKVJ C =? ,9#PFGTUQP2$CKTF4*&CXKUGVCN*GCNVJDGPGſVUQHFKGVCT[ſDGT0WVT
TGFWEGFTKUMQHQXCTKCPECPEGTCFQUGTGURQPUGOGVCCPCN[UKU0WVT4GU 4GX JVVRU//FQKQTI//LZ
JVVRU//FQKQTI//LPWVTGU =? %0CICVC-*KTQMCYC05JKOK\WGVCN#UUQEKCVKQPUQHOGPUVTWCNRCKP
=? , %JGP . 8KVGVVC +PHNCOOCVKQPOQFWNCVKPI GHHGEV QH DWV[TCVG KP VJG YKVJKPVCMGUQHUQ[HCVCPFFKGVCT[ſDGTKP,CRCPGUGYQOGP'WT,%NKP
RTGXGPVKQP QH EQNQP ECPEGT D[ FKGVCT[ HKDGT %NKP %QNQTGEVCN %CPE 0WVT JVVRU//FQKQTI//ULGLEP
JVVRU//FQKQTI//LENEE =? <<JCPI.5JK92CPIGVCN&KGVCT[HKDGTKPVCMGTGIWNCVGUKPVGUVKPCN
=? &-CYCMKVC;%#.GG(6WTCVKGVCN&KGVCT[HKDGTKPVCMGCPFJGCF OKETQHNQTC CPF KPJKDKVU QXCNDWOKPKPFWEGF CNNGTIKE CKTYC[ KPHNCOOCVKQP
CPFPGEMECPEGTTKUMCRQQNGFCPCN[UKUKPVJGKPVGTPCVKQPCNJGCFCPFPGEM KPCOQWUGOQFGN2NQU1PG JVVRU//FQKQTI//
ECPEGTGRKFGOKQNQI[EQPUQTVKWO+PV,%CPEGT LQWTPCNRQPG
JVVRU//FQKQTI//KLE =? 0 6QOKE $ &QLPQX , /KQEKPQXKE GV CN 'PTKEJOGPV QH [QIJWTV YKVJ
=? $ <JGPI * 5JGP * *CP GV CN &KGVCT[ HKDGT KPVCMG CPF TGFWEGF KPUQNWDNGFKGVCT[ſDGTHTQOVTKVKECNGŌCUGPUQT[RGTURGEVKXG.96(QQF5EK
TKUM QH QXCTKCP ECPEGT C OGVCCPCN[UKU 0WVT , 6GEJPQN JVVRU//FQKQTI//LNYV
JVVRU//FQKQTI//U =? 5; 2CTM -; ;QQP 'P\[OCVKE RTQFWEVKQP QH UQNWDNG FKGVCT[ HKDGT
=? %*9CPI%3KCQ4%9CPIGVCN&KGVCT[ſDGTKPVCMGCPFRCPETGCVKE HTQO VJG EGNNWNQUG HTCEVKQP QH %JKPGUG ECDDCIG YCUVG CPF RQVGPVKCN
ECPEGTTKUMCOGVCCPCN[UKUQHGRKFGOKQNQIKEUVWFKGU5EK4GR WUG CU C HWPEVKQPCN HQQF UQWTEG (QQF 5EK $KQ
JVVRU//FQKQTI//UTGR JVVRU//FQKQTI//U
=? / &GUEJCUCWZ % 2QWEJKGW / *KU GV CN &KGVCT[ VQVCN CPF KPUQNWDNG =? . $GNIJKVJ(GPFTK ( %JCCTK ( -CNNGN GV CN 2GC CPF DTQCF DGCP
HKDGT KPVCMGU CTG KPXGTUGN[ CUUQEKCVGF YKVJ RTQUVCVG ECPEGT TKUM , 0WVT RQFU CU C PCVWTCN UQWTEG QH FKGVCT[ HKDGT VJG KORCEV QP VGZVWTG CPF
JVVRU//FQKQTI//LP UGPUQT[ RTQRGTVKGU QH ECMG , (QQF 5EK
=? # 6CLCFFKPK # 2QWT\CPF < 5CPCCV GV CN &KGVCT[ TGUKUVCPV UVCTEJ JVVRU//FQKQTI//
EQPVCKPGFHQQFUCPFDTGCUVECPEGTTKUMCECUGEQPVTQNUVWF[KPPQTVJYGUVQH =? &/WFIKN5$CTCM$5-JCVMCT%QQMKGVGZVWTGURTGCFTCVKQCPFUGPUQT[
+TCP#UKCP2CE,%CPEGT2 CEEGRVCDKNKV[QHEQQMKGUCUCHWPEVKQPQHUQNWDNGFKGVCT[ſDGTDCMKPIVKOG
=? $.2QQN<QDGN,5CWGT1XGTXKGYQHGZRGTKOGPVCNFCVCQPTGFWEVKQPQH CPF FKHHGTGPV YCVGT NGXGNU .96(QQF 5EK 6GEJ
EQNQTGEVCNECPEGTTKUMD[KPWNKPV[RGHTWEVCPU,0WVT 5 JVVRU//FQKQTI//LNYV
JVVRU//FQKQTI//LP/5 =? ;*QPI9<,WP:,KCPGVCN&GXGNQROGPVQHVJGFKGVCT[ſDGTHWPEVKQPCN
=? ;$-CPI;.K;&WGVCN-QPLCMWƀQWTTGFWEGUQDGUKV[KPOKEGD[ HQQFCPFUVWFKGUQPKVUVQZKEQNQIKECNCPFRJ[UKQNQIKERTQRGTVKGU(QQF%JGO
OQFWNCVKPIVJGEQORQUKVKQPQHVJGIWVOKETQDKQVC+PV,1DGUKV[ 6QZKE JVVRU//FQKQTI//LHEV
JVVRU//FQKQTI//UZ =? 3.W*.KW39CPIGVCN5GPUQT[CPFRJ[UKECNSWCNKV[EJCTCEVGTKUVKEU
=? 3 <JCPI * ;W : :KCQ GV CN +PWNKPV[RG HTWEVCP KORTQXGU FKCDGVKE QHDTGCFHQTVKſGFYKVJCRRNGRQOCEGWUKPIHW\\[OCVJGOCVKECNOQFGN+PV,
RJGPQV[RG CPF IWV OKETQDKQVC RTQHKNGU KPTCVU 2GGT, (QQF5EK6GEJ JVVRU//FQKQTI//KLHU
JVVRU//FQKQTI//RGGTL =? .6%CTXCNJQ/#2KTGU,%$CNFKPGVCN2CTVKCNTGRNCEGOGPVQHOGCV
=? $(CTC\'2JKNNKR20CKNNKYGVCN&KGVCT[ſDGTVTGCVOGPVEQTTGEVUVJG CPFHCVYKVJJ[FTCVGFYJGCVHKDGTKPDGGHDWTIGTUFGETGCUGUECNQTKEXCNWG
EQORQUKVKQPQHIWVOKETQDKQVCRTQOQVGU5%(#RTQFWEVKQPCPFUWRRTGUUGU YKVJQWV TGFWEKPI VJGHGGNKPIQH UCVKGV[CHVGT EQPUWORVKQP/GCV 5EK
EQNQP ECTEKPQIGPGUKU )GPGU JVVRU//FQKQTI// JVVRU//FQKQTI//LOGCVUEK
IGPGU =? 8.#PIGNGU25#TCEGNK4++UTCGNGVCN%JGOKECNRJ[UKECNCPFUGPUQT[
=? /,#NHC&5VTCPI256CRRKCGVCN#TCPFQOK\GFVTKCNVQFGVGTOKPG RTQRGTVKGU QH 8KGPPC UCWUCIGU HQTOWNCVGF YKVJ C UVCTHTWKV FKGVCT[ HKDGT
VJG KORCEV QH C FKIGUVKQP TGUKUVCPV UVCTEJ EQORQUKVKQP QP VJG IWV EQPEGPVTCVG,(QQF5EK6GEJ JVVRU//FQKQTI//
OKETQDKQOGKPQNFGTCPFOKFCIGCFWNVU%NWP0WVT U
JVVRU//FQKQTI//LENPW =? ,/(GTPCPFG\)KPGU,(GTPCPFG\.QRG\'5C[CU$CTDGTCGVCN'HHGEV
=? /#5%JCROCP/()TCJP/#$Q[NG$WV[TCVGQZKFCVKQPKUKORCKTGF QHUVQTCIGEQPFKVKQPUQPSWCNKV[EJCTCEVGTKUVKEUQHDQNQIPCUCWUCIGUOCFG
KP VJG EQNQPKE OWEQUC QH UWHHGTGTU QH SWKGUEGPV WNEGTCVKXG EQNKVKU )WV YKVJEKVTWUſDGT,(QQF5EK JVVRU//FQKQTI//
JVVRU//FQKQTI//IWV LVDZ
=? &, 4QUG /6 &GOGQ # -GUJCXCT\KCP +PHNWGPEG QH FKGVCT[ HKDGT QP =? ,&ÈC\8GNC#6QVQUCWU/.2ÃTG\%JCDGNC+PVGITCVKQPQHCITQKPFWUVTKCN
KPƀCOOCVQT[DQYGNFKUGCUGCPFEQNQPECPEGTKORQTVCPEGQHHGTOGPVCVKQP EQRTQFWEVUCUHWPEVKQPCNHQQFKPITGFKGPVUECEVWURGCT 2SXQWLD¿FXVLQGLFD)
RCVVGTP 0WVT 4GX JVVRU//FQKQTI//L ƀQWTCPFRKPGCRRNG Ananas comosus RGGNƀQWTCUſDGTUQWTEGKPEQQMGF
VDZ UCWUCIGUKPQEWNCVGFYKVJNCEVKECEKFDCEVGTKC,(QQF2TQEGUU2TGUGTX
=? * <QJTG ( ,CPGNNG + *[WP GV CN &KGVCT[ RGC HKDGT UWRRNGOGPVCVKQP JVVRU//FQKQTI//LHRR
KORTQXGU IN[EGOKC CPF KPFWEGU EJCPIGU KP VJG EQORQUKVKQP QH IWV =? 69CPI</C#PQXGNKPUQNWDNGFKGVCT[HKDGTDCUGFGFKDNGRCRGTHTQO
OKETQDKQVCUGTWOUJQTVEJCKPHCVV[CEKFRTQſNGCPFGZRTGUUKQPQHOWEKPUKP %JKPGUGECDDCIG%GNNWNQUG JVVRU//FQKQTI//
INWEQUGKPVQNGTCPVTCVU0WVTKGPVU JVVRU//FQKQTI// U
PW =? <3 (W / 9W :; *CP GV CN 'HHGEV QH QMCTC FKGVCT[ HKDGT QP VJG
=? %>V/2JKNNKRUQP.*QNOGVCN2TGXGPVKXGCPFVJGTCRGWVKEGHHGEVUQH RTQRGTVKGUQHUVCTEJDCUGFHKNOU5VCTEJ5VTMG
PKVTKVGUWRRNGOGPVCVKQPKPGZRGTKOGPVCNKPƀCOOCVQT[DQYGNFKUGCUG4GFQZ JVVRU//FQKQTI//UVCT
$KQN JVVRU//FQKQTI//LTGFQZ =? 626TKPKFCF#%/CNNKNNKP&*8CNFG\GVCN&KGVCT[ſDGTHTQOEQEQPWV
=? # 1\GT 4GOQXCN QH 2D ++ KQPU HTQO CSWGQWU UQNWVKQPU D[ UWNRJWTKE HNQWT C HWPEVKQPCN HQQF +PPQX (QQF 5EK 'OGTI
CEKFVTGCVGF YJGCV DTCP , *C\CTF /CVGT JVVRU//FQKQTI//LKHUGV
JVVRU//FQKQTI//LLJC\OCV =? ;0KW3:KC/)WGVCN+PVGTRGPGVTCVKPIPGVYQTMIGNUEQORQUGFQH
=? /-JQVKOEJGPMQ8-QXCNGX;-JQVKOEJGPMQ'SWKNKDTKWOUVWFKGUQH IGNCVKPCPFUQNWDNGFKGVCT[HKDGTUHTQOVQOCVQRGGNU(QQF*[FTQEQNN
UQTRVKQPQHNGCF ++ KQPUD[FKHHGTGPVRGEVKPEQORQWPFU,*C\CTF/CVGT JVVRU//FQKQTI//LHQQFJ[F
JVVRU//FQKQTI//LLJC\OCV =? 9)2QOCEG2%QORQWPF&(KDGT&GXGNQROGPVQHCPVKQZKFCPVFKGVCT[
=? * 9CPI 6 *WCPI <% 6Q GV CN 6JG CFUQTRVKQP QH NGCF ++ KQPU D[ HKDGTU HTQO YKPG ITCRG RQOCEG CPF VJGKT CRRNKECVKQPU CU HWPEVKQPCN HQQF
F[PCOKEJKIJRTGUUWTGOKETQƀWKFK\CVKQPVTGCVGFKPUQNWDNGUQ[DGCPFKGVCT[ KPITGFKGPVU9KPG)TCRG2QOCEG