You are on page 1of 10

Y. He et al.

/ Food Science and Human Wellness 11 (2022) 1-10 1

%QPVGPVUNKUVUCXCKNCDNGCV5EKGPEG&KTGEV

)RRG6FLHQFHDQG+XPDQ:HOOQHVV
LQWTPCNJQOGRCIGJVVRYYYMGCKRWDNKUJKPIEQOGPLQWTPCNUHQQFUEKGPEGCPFJWOCPYGNNPGUU

'HHGEVUQHFKGVCT[ſDGTQPJWOCPJGCNVJ
;CPI*Ga,b,1$KZKCPI9CPIb,1.KCPMWK9GPb,*(GPI\JQPI9CPIa,c,*,
*CPUQPI;Wb,c,*&QPIZKC%JGPb:KP5Wb%JK<JCPIb
a
Institute of Food Science and Technology, Chinese Academy of Agricultural Sciences (CAAS), Beijing 100193, China
b
Department of Food Science and Engineering, Jilin Agricultural University, Changchun 130118, China
c
Division of Soybean Processing, Soybean Research & Development Center, Chinese Agricultural Research System, Changchun 130118, China

#46+%.'+0(1 #$564#%6

Article history: 'LHWDU\¿EHU ') LVEHOLHYHGWRSURYLGHLPSRUWDQWKHDOWKEHQH¿WVDQGLWKDVEHFRPHDUHVHDUFKKRWVSRW') H[LVWV


4GEGKXGF5GRVGODGT LQPRVWQDWXUDOIRRGVLQFOXGLQJIUXLWV %í% YHJHWDEOHV %í% JUDLQV %í% 
4GEGKXGFKPTGXKUGFHQTO&GEGODGT DQGVRRQ')KDVGLIIHUHQWSK\VLFRFKHPLFDOSURSHUWLHVVXFKDVZDWHUKROGLQJFDSDFLW\ :+&  íJ/J 
#EEGRVGF,CPWCT[ ZDWHU VZHOOLQJ FDSDFLW\ :6&  í P//J  RLO KROGLQJ FDSDFLW\ 2+&  í J/J 
#XCKNCDNG1PNKPG#WIWUV JOXFRVHDGVRUSWLRQFDSDFLW\ *$&  íPPRO/J FKROHVWHURODGVRUSWLRQFDSDFLW\ &$&  íPJ/J 
DQGYLVFRVLW\ZKLFKPDNHLWH[KLELWVGLIIHUHQWSK\VLRORJLFDOIXQFWLRQVVXFKDVUHGXFLQJWKHULVNRIREHVLW\
Keywords: GLDEHWHVFDQFHUDQGLQWHVWLQDOGLVHDVHV 2QWKHEDVLVRIFRQVXOWLQJUHODWHGGDWDEDVHVWKHSK\VLFRFKHPLFDO
'LHWDU\¿EHU SURSHUWLHVRI')DQGLWVGHULYHGSK\VLRORJLFDOIXQFWLRQVDUHUHYLHZHG,WLVIRXQGWKDW')KDVHIIHFWRQWKH
3K\VLFRFKHPLFDOSURSHUWLHV SUHYHQWLRQDQGWUHDWPHQWRIREHVLW\GXHWRJRRG:+&:6&2+&DQG&$& *RRG*$&FDQUHOLHYHWKH
+XPDQKHDOWK V\PSWRPVRIGLDEHWHV7KHSK\VLFRFKHPLFDOSURSHUWLHVRI:+&DQG:6&FDQUHGXFHWKHULVNRIFDQFHU7KH
:+&:6&DQGYLVFRVLW\DUHEHQH¿FLDOWRLQWHVWLQDOKHDOWK
© 2021 BGKLKPI#ECFGO[QH(RQF5EKGPEGU2WDNKUJKPIUGTXKEGUD['NUGXKGT$8QPDGJCNHQH-G#K
%QOOWPKECVKQPU%Q.VF6JKUKUCPQRGPCEEGUUCTVKENGWPFGTVJG%%$;0%0&NKEGPUG
JVVR//ETGCVKXGEQOOQPUQTI/NKEGPUGU/D[PEPF// 

1. Introduction VRPHKHPLFHOOXORVHV>@')FRPHVIURPDZLGHUDQJHRIVRXUFHV
LQFOXGLQJ IUXLWV YHJHWDEOHV JUDLQV DV ZHOO DV RWKHUV 7DEOH  
'LHWDU\ ILEHU ')  LV WKH HGLEOH SDUWV RI SODQWV ZKLFK FDQ EH +LJKYLVFRVLW\6') VXFKDVSHFWLQDQGROLJRVDFFKDULGH LVKLJKHU
FRPSOHWHO\RUSDUWLDOO\IHUPHQWHGLQODUJHLQWHVWLQHZLWKXQGLJHVWHG LQ DSSOHV RUDQJHV SHUVLPPRQV SHDUV GULHG EHDQV FDXOLIORZHU
DQG XQDEVRUEDEOH LQ WKH KXPDQ VPDOO LQWHVWLQH ,W LV WKH VXP FDUURWV DQG SRWDWRHV HWF 3HFWLQ LV D QHWZRUN RI SRO\JDODFWXURQLF
RI FDUERK\GUDWHV DQG WKHLU DQDORJXHV LQFOXGLQJ QRQGLJHVWLEOH DFLG FRPSRVHG RI DJDODFWXURQLF DFLG UHVLGXHV OLQNHG E\ Į
SRO\VDFFKDULGHVROLJRVDFFKDULGHVDQGDVVRFLDWHGSODQWVXEVWDQFHV>@ JO\FRVLGLF ERQGV DQG KDV JRRG YLVFRVLW\ DQG DEVRUEDELOLW\ 6R
') FDQ EH GLYLGHG LQWR VROXEOH GLHWDU\ ILEHU 6')  DQG LQVROXEOH LW KDV WKH SURSHUW\ RI DGVRUE JOXFRVH FKROHVWHURO DQG KHDY\
GLHWDU\¿EHU ,') EDVHRQLWVVROXELOLW\LQKRWZDWHU6')LVQRQ PHWDOV >@ :LOODWV HW DO >@ VKRZHG WKDW WKH GLIIHUHQW GHJUHH RI
FHOOXORVLF SRO\VDFFKDULGH LQFOXGLQJ ROLJRVDFFKDULGHV DQG VRPH SRO\PHUL]DWLRQ DQG FRPSOH[ VLGH FKDLQ RI SHFWLQ DUH LPSRUWDQW LQ
LQGLJHVWLEOHSRO\VDFFKDULGHV HJLQXOLQDUDELFJXPJXPSHFWLQV DQWLFDQFHU DFWLYLW\ DQG RWKHU ELRORJLFDO DFWLYLW\ FKDUDFWHULVWLFV
JDODFWRPDQQDQDQG ȕJOXFDQV ,')LQFOXGHVFHOOXORVHOLJQLQDQG DQG LWV PHFKDQLVP QHHGV WR EH VWXGLHG 0RVW ,') VXFK DV OLJQLQ
FHOOXORVH DQG VHPLILEHUV LQ JUDLQV HJ ZKHDW VR\EHDQV PXQJ
1
 6JGCWVJQTUEQPVTKDWVGFGSWCNN[VQVJKUCTVKENG
EHDQV DQG RDWV  DUH VPDOO LQ YLVFRVLW\ EXW KDYH JRRG VZHOOLQJ
*
 %QTTGURQPFKPICWVJQTU SURSHUWLHV &HOOXORVH LV D SRO\PHULQVROXEOH SRO\VDFFKDULGH
E-mail addressYGPNKCPMWK"LNCWGFWEP .-9GP  FRQQHFWHGE\JOXFRVHWKURXJKȕ ĺ JO\FRVLGLFERQGVDQGEHORQJV
YCPIHGPI\JQPI"UKPCEQO (<9CPI [WJCPUQPI"EQO *5;W 
WR OLQHDU PROHFXOH 7KH DGMDFHQW FHOOXORVH FKDLQV KDYH D VWDEOH
2GGTTGXKGYWPFGTTGURQPUKDKNKV[QH-G#K%QOOWPKECVKQPU%Q.VF
¿EURXVVWUXFWXUHE\K\GURJHQERQGLQJZKLFKPDNHVFHOOXORVHKDV
ZDWHUKROGLQJFDSDFLW\ :+& ZDWHUVZHOOLQJFDSDFLW\ :6& RLO

Publishing services by Elsevier

JVVRFQKQTILHUJY
l$GKLKPI#ECFGO[QH(QQF5EKGPEGU2WDNKUJKPIUGTXKEGUD['NUGXKGT$8QPDGJCNHQH-G#K%QOOWPKECVKQPU%Q.VF6JKUKUCPQRGPCEEGUUCTVKENGWPFGTVJG%%
$;0%0&NKEGPUG JVVRETGCVKXGEQOOQPUQTINKEGPUGUD[PEPF 
2 Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10

KROGLQJ FDSDFLW\ 2+&  JOXFRVH DGVRUSWLRQ FDSDFLW\ *$&  DQG íJ DQGJUDLQV íJ  7DEOH 9HJHWDEOH')KDV
FKROHVWHURODGVRUSWLRQFDSDFLW\ &$& HWF>@ KLJK:+&GXHWRLWVLUUHJXODUORRVHDQGSRURXVVXUIDFHMXVWDV;LH
:LWK WKH UDSLG GHYHORSPHQW RI PRGHUQ VRFLHW\ PDQ\ SDWLHQWV HWDO>@VWXGLHGWKDW$QJHOLFDNHLVNHL¶VURRW')KDVKLJK:+&
ZLWKFKURQLFGLVHDVHVDUHGUDPDWLFDOO\LQFUHDVLQJLQFOXGLQJREHVLW\ =KHQJHWDO>@UHSRUWHGWKDWWKH')FRQWHQWLQFRFRQXWLV “
GLDEHWHV FDQFHU LQWHVWLQDO GLVHDVHV DQG RWKHU QRQFRPPXQLFDEOH  JZKLFKLVKLJKHUWKDQWKDWRIPRVWYHJHWDEOHV7KLVPD\EH
GLVHDVHV 7KH LQFLGHQFH RI GLVHDVH LV LQFUHDVLQJ \HDU E\ \HDU GXHWRWKH')VWUXFWXUHRIFRFRQXWLVORRVHUDQGPRUHSRURXVWKDQ
ZRUOGZLGHZKLFKIXUWKHUOHDGVWRDQLQFUHDVHLQPRUWDOLW\ZRUOGZLGH PRVWYHJHWDEOHVZKLFKLQFUHDVHVWKHVXUIDFHDUHDDQGIDFLOLWDWHVZDWHU
>@7KHRFFXUUHQFHRIFKURQLFGLVHDVHVQRWRQO\LQFUHDVHVWKH UHWHQWLRQ,QIRRGUHVHDUFKGHYHORSPHQWDQGLQGXVWULDOSURGXFWLRQ
IDPLO\DQGJOREDOHFRQRPLFEXUGHQEXWDOVRUHGXFHVWKHKDSSLQHVV SURFHVVLQJDQGPRGLILFDWLRQPHWKRGVDUHRIWHQXVHGWRFKDQJHWKH
LQGH[ (YLGHQFH VXJJHVWV WKDW ') FDQ SDUWLFLSDWH LQ KXPDQ VWUXFWXUH RI ') WR SURPRWH WKH GHVLUH IRU EHWWHU SK\VLFRFKHPLFDO
PHWDEROLVPWRSUHYHQWDQGWUHDWWKHVHGLVHDVHV )LJ >@:HL SURSHUWLHV')KDVDJRRG:+&ZKLFKLVRQHRIWKHPDLQUHDVRQV
HW DO >@ VKRZHG WKDW  J/GD\ LQFUHDVH LQ WRWDO ILEHU LQWDNH WKH IRULWVUROHLQLPSURYLQJKXPDQKHDOWK)LUVWO\')KDVDOXEULFDWLQJ
SUREDELOLW\RIVXIIHULQJIURPPHWDEROLFV\QGURPHLVUHGXFHGE\%
HIIHFWDIWHUDEVRUELQJZDWHUZKLFKFDQSURPRWHLQWHVWLQDOSHULVWDOVLV
,WKDVDOVREHHQIRXQGWKDWDPRQJWKH')FHUHDODQGIUXLW¿EHUFDQ
DQGLQWHVWLQDOPRWLOLW\>@6HFRQGO\')H[SDQGVWKHYROXPHRIIRRG
UHGXFHWKHULVNRIPHWDEROLFV\QGURPH')ZLWKGLIIHUHQWVWUXFWXUHV
DIWHUDEVRUELQJZDWHUZKLFKFDQUHGXFHIRRGLQWDNH')LVIHUPHQWHG
KDVGLIIHUHQWSK\VLFRFKHPLFDOSURSHUWLHVZKLFKSOD\DYLWDOUROHLQ
LQWKHLQWHVWLQHWRSURGXFH6&)$VZKLFKFDQSURPRWHWKHVHFUHWLRQ
UHJXODWLQJ WKH KHDOWK PHFKDQLVP RI KXPDQ ERG\ 6WXGLHV EHOLHYH
RI VDWLHW\ KRUPRQHV >@ 7KRVH ZLOO UHGXFH WKH ULVN RI REHVLW\
WKDW WKH SK\VLFRFKHPLFDO SURSHUWLHV RI ') VXFK DV :+& :6&
7KLUGO\ ') FDQ LQFUHDVH VWRRO YROXPH DIWHU DEVRUELQJ ZDWHU DQG
2+&*$&&$&DQGYLVLELOLW\DUHFORVHO\UHODWHGWRSK\VLRORJLFDO
VZHOOSURPRWHGHIHFDWLRQZKLOHWDNLQJFDUFLQRJHQVDZD\>@ $W
IXQFWLRQV7KHUHIRUHWKHUHODWLRQVKLSEHWZHHQWKHVHSK\VLFRFKHPLFDO
DQGSK\VLRORJLFDOIXQFWLRQVDUHGLVFXVVHGDQGWKHSURGXFWRI')LQ WKHVDPHWLPH')IHUPHQWDWLRQFDQUHJXODWHJXWPLFURELRWDVRDVWR
IXQFWLRQDOIRRGLQOLIHDUHFROOHFWHG7KHVHDUHH[SHFWHGWROD\WKH SUHYHQWGLVHDVHV>@
IRXQGDWLRQIRUWKHGHYHORSPHQWRI')KHDOWKIRRG
2.2 WSC
2. Physicochemical properties
7KHPHWKRGWRGHWHUPLQH:6&LVWRDGG')VDPSOH J DQG
2.1 WHC ZDWHULQWRWKHJUDGXDWHGF\OLQGHURYHUQLJKWDQGUHFRUGWKHH[SDQVLRQ
YROXPH >@')KDV:6&IRUWKHIROORZLQJUHDVRQV :DWHUKDV
:+&LVGHVFULEHDVPL[JRI') GU\EDVLV ZLWKZDWHUXQGHU VXUIDFH WHQVLRQ VWUHQJWK LW FDQ EH PDLQWDLQHG LQ WKH ') FDSLOODU\
VSHFLILFFRQGLWLRQVDQGFDOFXODWHWKHDPRXQWRIZDWHUUHWDLQHG J  VWUXFWXUH   +\GURJHQ ERQGV DQG GLSROH K\GURSKLOLF JURXSV ELQG
DIWHURYHUQLJKW>@7KH:+&KDVDFHUWDLQUHODWLRQVKLSZLWKWKH WRZDWHU 7KLVLVGLUHFWO\UHODWHGWRWKHFRQWDFWDUHDRI')7KH
VWUXFWXUH RI ') :KHQ WKH FRQWDFW DUHD DQG WKH K\GURSKLOLF JURXS :6&RI')UDQJHVIURPP//JWRP//JRIZKLFK:6&RI
RI')LQFUHDVHZDWHUFDQEHUHWDLQHGLQWKHK\GURSKLOLFSDUWRULQ YHJHWDEOHV')LVíJ/JWKHIUXLW')LVíJ/JDQG
WKHVSDFHRIQHWZRUN¿EHUVWUXFWXUHZKLFKZLOOLPSURYHWKH:+& WKHJUDLQ')LVíJ/J 7DEOH 7KH:6&RIFRFRQXW')LV
7KH:+&RI')UDQJHVIURPJ/JWRJ/JDPRQJWKHP WKHVWURQJHVWWRUHDFK “ J/JLWH[SRVHVPRUHK\GURJHQ
YHJHWDEOH')KDVWKHKLJKHVW:+& íJ IROORZHGE\IUXLWV ERQGVDQGGLSROHIRUPV>@;LHHWDO>@UHSRUWHGWKDWWKH:6&RI

Obesity
Diabetes
h
Human health
Dietary fiber Cancer
Intestinal disease

Stomach Stimulate the secretion of gastric digestive juice,


enhance gastrointestinal motility

Regulate the gut


Resist the digestion and microbiota and promote
Small intestine the production of SCFAs
absorption of the human
small intestine Gut microbiota

Fully fermented fibers include β-glucan, pectin,


guar gum, acacia, seaweed gum and inulin, etc. Large intestine
Some fermented fibers include fiber crisp,
Excreted after absorbing
hemicellulose, lignin, vegetable wax and keratin, etc.
water and swelling

Fig. 1 /GVCDQNKUOQH&(KPJWOCPDQF[CPFFKUGCUGRTGXGPVKQP
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10 3

')IURPSXUSOHSRWDWRHVDUHUHODWHGWRSURFHVVLQJFRQGLWLRQV+LJK HQKDQFHWKH&$& PJ/J WKHUHE\UHGXFLQJEORRGFKROHVWHURO


SUHVVXUHKRPRJHQL]HG')KDVJRRG:6& P//J EHFDXVHWKH OHYHOV ')ZLWKJRRG&$&FDQSUHYHQWDQGWUHDWREHVLW\GLVHDVH')
DOWHUDWLRQRIFKHPLFDODQGVWUXFWXUDOQDWXUHVLQGXFHGE\KLJKSUHVVXUH FDQDEVRUEFKROHVWHURODQGWKH6&)$VSURGXFHGE\IHUPHQWDWLRQFDQ
WUHDWPHQWV*HQHUDOO\WKH:6&RI')LVEHQH¿FLDOWRLWVIXQFWLRQDO SDUWLFLSDWHLQERG\PHWDEROLVPDQGLPSURYHG\VOLSLGHPLDZKLFKDUH
SURSHUWLHVIRUWKHVDPHUHDVRQVDV:+& WKHUHDVRQVIRUSUHYHQWLQJREHVLW\>@

2.3 OHC 2.6 Viscosity

7KHPHWKRGWRGHWHUPLQH2+&LVWRPL[VDPSOH J ZLWKVR\EHDQ 9LVFRVLW\ LV WKH UHODWLRQVKLS EHWZHHQ IOXLG IORZ DQG WKH IRUFH
RLOOHDYHLWWRVWDQGIRUUHDFWLRQDQGFHQWULIXJHWRUHWDLQWKHDPRXQWRI GLUHFWHGRQWKHIOXLGDQGLWFDQEHDIIHFWHGE\PDQ\IDFWRUVVXFK
RLO>@7KH2+&RI')LVPDLQO\UHODWHGWRWKHWRWDOFKDUJHGHQVLW\ DV FKHPLFDO FRPSRVLWLRQ VWUXFWXUH SDUWLFOH VL]H VXUIDFH DUHD DQG
K\GURSKRELFLW\ VXUIDFH SURSHUWLHV DQG WUHDWPHQW FRQGLWLRQV 7KH SURFHVVLQJ FRQGLWLRQV *DOODKHU HW DO >@ XVHG ZKROH RDWPHDO RU
2+&RI')UDQJHVIURPJ/JWRJ/J,QJHQHUDOWKH2+&RI LQVWDQWRDWPHDOWRPDNHPXI¿QVDQGIHGWRUDWVDQGWKHYLVFRVLW\RI
YHJHWDEOH') íJ/J LVKLJKHUWKDQIUXLW') íJ/J  WKHLQWHVWLQDOFRQWHQWVVXSHUQDWDQWLVPHDVXUHG7KHUHVXOWVVKRZHG
DQG FHUHDO ') í J/J  7DEOH   $PRQJ WKHP ZDOQXW WKDWWKHYLVFRVLW\RIZKROHRDWPHDO DERXWF3 LVVLJQLILFDQWO\
SRZGHU KDV WKH ODUJHVW 2+& LW KDV PRUH SRURXV DQG KRQH\ FRPE KLJKHUWKDQHDWLQJLQVWDQWRDWPHDO DERXWF3 7KHUHDVRQLVWKDW
OLNH >@ <X HW DO >@ SURSRVHG WKDW WKH 2+& RI FDUURW ') LV PRUHYLVFRVLW\ILEHUVXFKDV ȕJOXFDQLQZKROHRDWVDQG ȕJOXFDQ
VLJQL¿FDQWO\LPSURYHGEHFDXVHWKHHQ]\PHWUHDWPHQWH[SRVHGPRUH KDV PRUH K\GURSKLOLF JURXSV ZKLFK SURPRWHV WKH LQFUHDVH RI WKH
K\GURSKLOLFJURXSVDQGWKHFHOOXODUQHWZRUNVWUXFWXUH=KDRHWDO>@ ZDWHU¿OPWKLFNQHVVRIWKHLQWHVWLQDOPXFRVDWKHUHE\LQFUHDVLQJWKH
VKRZHGWKDWWKH2+& J/J RIVXSHU¿QHJULQGLQJULFHEUDQ,') YLVFRVLW\RIVXSHUQDWDQWLQWHVWLQDOFRQWHQWV8OODKHWDO>@ IRXQGWKDW
LVVLJQL¿FDQWO\ORZHUWKDQWKDWRI¿QH,') J/J DQGFUXGH')
WKHYLVFRVLW\RIRNDUDVWDELOLW\LQFUHDVHGDIWHUWKHUPDOSUHWUHDWPHQW
J/J $VWKHSDUWLFOHVL]HRIVXSHUILQHJULQGLQJGHFUHDVHVWKH
7KHYLVFRVLW\RI')FDQSUHYHQWREHVLW\DQGGLDEHWHVE\GHOD\LQJIDW
KRQH\FRPE VWUXFWXUH RI ,') GLVDSSHDUV ZKLFK PDNHV WKH 2+&
IRUPDWLRQDQGJDVWULFHPSW\LQJDQGLQFUHDVLQJLQVXOLQVHQVLWLYLW\
GHFUHDVH7KH2+&RI')LVDOVRWKHEDVLVRILPSRUWDQWIXQFWLRQDO
SURSHUWLHV')FDQDEVRUEIDWDQGUHGXFHWKHLQWDNHRIH[FHVVFDORULHV
LQWKHERG\ZKLFKFDQSUHYHQWWKHRFFXUUHQFHRIREHVLW\>@ 3. DF and human health

3.1 DF and obesity


2.4 GAC
,QUHFHQW\HDUVREHVHSDWLHQWVKDYHEHHQLQFUHDVLQJ\HDUE\\HDU
)#%KUFGUETKDGCUVJGOKZ&( I CPFO.INWEQUGUQNWVKQP DOO RYHU WKH ZRUOG $FFRUGLQJ WR VWDWLVWLFV IURP  FRXQWULHV WKH
íOOQN/. KPCPGP\[OGTGCEVQTCPFOKZVJGCOQWPVQHINWEQUG JOREDO SUHYDOHQFH RI REHVLW\ KDV LQFUHDVHG IURP % LQ  WR
TGVCKPGFD[EGPVTKHWICVKQP=?)#%QH&(KUTGNCVGFVQKVURJ[UKECN %LQLQPHQ$WWKHVDPHWLPHWKHGDWDVXJJHVWVDQLQFUHDVH
RTQRGTVKGUDGECWUGINWEQUGECPCFJGTGVQVJGPGVYQTMUVTWEVWTGQH&( IURP%RIZRPHQWR%>@,QPRUHWKDQELOOLRQ
CPFTGFWEGVJGEQPVCEVYKVJJWOCPKPVGUVKPCNVTCEV=?6JG)#%QH DGXOWVDJHGZHUHRYHUZHLJKWRIZKLFKPRUHWKDQPLOOLRQZHUH
&(KPHQQFKUHTQOOOQN/IVQOOQN/ICPFUVWFKGUQP)#% REHVHDQGPRUHWKDQPLOOLRQFKLOGUHQDQGDGROHVFHQWVDJHGí
OCKPN[HQEWUQPXGIGVCDNG&( 6CDNG *WCPIGVCN=?HQWPFVJCV ZHUHRYHUZHLJKWRUREHVH7KHLQFUHDVLQJQXPEHURIREHVHSDWLHQWV
)#%QHICTNKEUVGOU&(KPETGCUGFCHVGTWNVTCUQPKEVTGCVOGPVYJKEJ SRVHV D WKUHDW WR KXPDQ KHDOWK ZRUOGZLGH LQFUHDVHV WKH JOREDO
KUFWGVQVJGKPETGCUGQHJQPG[EQODPGVYQTMUVTWEVWTG;WGVCN=? EXUGHQDQGEHFRPHVDFKDOOHQJHIRUKXPDQ>@ 5HGXFLQJREHVH
TGRQTVGF VJCV +&( GZVTCEVGF HTQO ECTTQV WUKPI C EQORNGZ GP\[OG SDWLHQWVKDVEHFRPHDQXUJHQWSUREOHP$GHTXDWHLQWDNHRI')KDV
OGVJQF OOQN/I CPF)#%GZJKDKVUCUKIPKHKECPVN[KORTQXGF EHHQVKRZQWRSOD\DQLPSRUWDQWUROHLQSUHYHQWLQJREHVLW\
YKVJVJGJKIJJ[FTQUVCVKERTGUUWTG OOQN/I 6JG)#%QH&(KU ') FDQ EORFN WKH IDW DEVRUSWLRQ DQG UHGXFH HQHUJ\ LQWDNH E\
TGNCVGFVQVJGINWEQUGOGVCDQNKUOHWPEVKQP6JKUKUDGECWUG&(ECP UHJXODWLQJ IRRG LQWDNH GLJHVWLRQ DEVRUSWLRQ DQG PHWDEROLVP 7KH
CDUQTDUWICTUCPFRTQFWEG5%(#UVJTQWIJHGTOGPVCVKQPKPVJGDQF[ PDLQZHLJKWORVVPHFKDQLVPRI')LVDVIROORZV 7KHYLVFRVLW\
YJKEJECPRTGXGPVVJGFKCDGVGU=? RI6')VXFKDVJXPDQGJXPDUDELFLVKLJKZKLFKFDQVORZGRZQ
WKH PLJUDWLRQ RI QXWULHQWV ') KDV JDODFWXURQLF DFLG VWUXFWXUH
2.5 CAC ZKLFK KDV D KLJK YLVFRVLW\ >@ $W WKH VDPH WLPH ') KDV JRRG
:+&2+&DQG&$&ZKLFKFDQHQKDQFHWKHDGVRUSWLRQRIIDW,Q
&$&LVH[SUHVVHGDVWKHPDVVRIFKROHVWHURODEVRUEHGE\JRI DGGLWLRQWKHQHWZRUNVWUXFWXUHRI,')PDNHVLWKDYHEHWWHU&$&>@
VDPSOH&$&RI')LVGLYLGHGLQWRSK\VLFDODQGFKHPLFDODGVRUSWLRQ 0DQGLPLND HW DO >@ GHVFULEHG WKDW EURFFROL ') FRQWDLQV VWLFN\
3K\VLFDO DGVRUSWLRQ LV PDLQO\ UHODWHG WR WKH SDUWLFOH VL]H VSHFLILF SHFWLQ ZKLFK UHGXFHV WKH LQWDNH RI GLHWDU\ IDW DQG HQKDQFHV WKH
VXUIDFH DUHD SRURVLW\ DQG UHDFWLRQ WHPSHUDWXUH RI ') >@ UHPRYDORIFKROHVWHURODQGELOHDFLGIURPHQWHURKHSDWLFFLUFXODWLRQ
&KHPLFDO DGVRUSWLRQ LV UHODWHG WR WKH ') FKDUJH DQG K\GURSKRELF WKHUHE\ UHGXFLQJ EORRG FKROHVWHURO OHYHOV /XR HW DO >@ IRXQG
JURXS &$& UDQJHV IURP  PJ/J WR  PJ/J DQG WKH PDMRU WKDW EDPERR VKRRW VKHOO FRQWDLQHG D ODUJH DPRXQW RI ,') ZKLFK
VWXGLHVRI&$&IRFXVRQYHJHWDEOHV') íPJ/J  7DEOH  FDQELQGRUDGVRUEELOHDFLGVDQGFKROHVWHUROOHDGLQJWRDUHGXFWLRQ
%DPERRVKRRWVKHOO')FDQELQGRUDGVRUEELOHDFLGVDQGFKROHVWHURO RI FKROHVWHURO LQ YDULRXV SRROV   ') KDV JRRG :6& ZKLFK FDQ
 PJ/J  WKHUHE\ UHGXFLQJ FKROHVWHURO >@ /LX HW DO >@ HQKDQFH VDWLHW\ DQG UHGXFH IRRG LQWDNH /DPEHUW HW DO >@ IRXQG
VKRZHG WKDW 6') LQ VR\EHDQ KXOOV KDV JRRG YLVFRVLW\ ZKLFK FDQ WKDWWKHLQWDNHRI')ZLOODEVRUEZDWHUDQGLQFUHDVHVDWLHW\ZKLFK
4
Table 1
5WOOCT[QH&(EQPVGPVCPFKVURJ[UKEQEJGOKECNRTQRGTVKGUHTQOFKHHGTGPVUQWTEGU
&(EQPVGPV % Dry) )#%
%NCUUKſECVKQP 5QWTEGU 9*% I/I 95% O./I 1*% I/I %#% OI/I 8KUEQUKV[ ER 'ZVTCEVKQPOGVJQFU 4GHGTGPEGU
6&( 5&(/+&( OOQN/I
)QNFGPDGTT[  / ± 6&( ± ±    %JGOKECN =?
$TGCFPWVHTWKV  / ± +&( ± ±    'P\[OGU =?
#RRNGRQOCEG   5&(      /KETQYCXGCUUKUVGF =?
/GZKECPCRRNGRQOCEG  / ± 6&( ± ±     =?
(TWKVU %KVTWURQOCEG  / ± +&( ± ±    ,GVITKPFKPI =?
)TCRGRQOCEG  / ± 6&( ± ±     =?

Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10


%QEQPWVſDTG  / ± 6&( ± ±    /WNVKRNGGP\[OGU =?
2CRC[CRGGN  ± 5&( ± ±    7NVTCUQWPFCUUKUVGF =?
1TCPIGRGGN ±  =?
2WTRNGƀGUJGFRQVCVQ  / ± 6&( ± ±    *KIJRTGUUWTG =?

;GNNQYƀGUJGFRQVCVQ   ± +&( ±    /WNVKRNGGP\[OGU =?


)CTNKEUVTCY  / ± +&( ± ± ± ±  /WNVKRNGGP\[OGU =?
8GIGVCDNGU %CTTQVRQOCEG  /  +&(      7NVTCUQWPFCUUKUVGF =?
$CODQQUJQQVUJGNN  ± 5&(  ± ±  ±  /WNVKRNGGP\[OGU =?
6QOCVQTGUKFWGſDGT  /   ±    %JGOKECN =?
0QFGQHNQVWUTQQV  ± 5&( ± ±    /WNVKRNGGP\[OGU =?
#NIC  Ō +&(  Ō    'PIN[UV =?
#PIGNKECMGKUMGK  ± +&( ± ±    /WNVKRNGGP\[OGU =?

4KEGDTCP  / ± +&( ± ±    %JGOKECN =?


5Q[JWNNUUQNWDNGſDGT  ± 5&(  ±  ±  %JGOKECN =?
$NCEMDGCPEQCV  /        =?
$CODCTCITQWPFPWVDNCEMG[G  ± +&( ±     'P\[OGU =?
%GTGCNU &GHCVVGFYCNPWVƀQWT   6&(      %JGOKECN =?
3KPIMGƀQWT  / ± 6&( ± ±    )TKPFKPI =?
%CEVWUTCEMGV  / ± 6&(  ±    %JGOKECN =?
1CVDTCP       220 =?

/CK\GUVTCY  / ± 6&( ± ±    %JGOKECN =?
1VJGTU
'URTGUUQEQEQCYCUVG  / ± 6&( ± ±    %JGOKECN =?
KPFKECVGUFKGVCT[ſDGTFKFPQVJCXGCPGHHGEVQPVJKURCTCOGVGT
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10 

FDQGHOD\QXWULHQWVDEVRUSWLRQDQGHIIHFWLYHO\UHGXFHHQHUJ\LQWDNH LQFUHDVHWKHVHQVLWLYLW\RILQVXOLQWRK\SRJO\FDHPLFHIIHFWV %REERLHW


  ') SURGXFHV 6&)$V WKURXJK LQWHVWLQDO PLFURELDO IHUPHQWDWLRQ DO>@QRWHGWKDWWKHYLVFRVLW\RILQJHVWHGJXDUJXPFDQUHGXFHWKH
ZKLOHSURPRWHVWKHVHFUHWLRQRIJOXFDJRQOLNHSHSWLGH */3 DQG SRVWSUDQGLDOK\SHUJO\FHPLD ')LVIHUPHQWHGE\WKHJXWPLFURELRWD
<<SHSWLGH 3<< */3LVUHOHDVHGIURPLQWHVWLQDO/FHOOVZKLFK WRSURGXFH6&)$VWKHUHE\VWLPXODWHWKHVHFUHWLRQRIVDWLHW\KRUPRQH
SURPRWHVLQVXOLQVHFUHWLRQDQGSDQFUHDWLFFHOOSUROLIHUDWLRQFRQWUROV */3DQG3<< WKRVHKRUPRQHVKHOSWRLQFUHDVHLQVXOLQVHFUHWLRQ
JO\FRJHQV\QWKHVLVLQPXVFOHFHOOVDQGHQKDQFHVVDWLHW\>@3<< DQG FRQWURO WKH EORRG JOXFRVH OHYHO =KDR HW DO >@ UHSRUWHG WKDW
LV DQ LQWHVWLQDO VHFUHWLRQ KRUPRQH ZLWK DQWLREHVLW\ HIIHFW ZKLFK DFHWLF DFLG DQG EXW\ULF DFLG FDQ LPSURYH JOXFRVH KRPHRVWDVLV E\
KDVWKHHIIHFWRIVXSSUHVVLQJDSSHWLWHDQGUHGXFLQJIRRGLQWDNH>@ LQGXFLQJ*/3DQG3<<LQWHVWLQDOSURGXFWLRQWKHUHE\VWLPXODWLQJ
&KDQJ HW DO >@ UHSRUWHG WKDW WKH ,') RI SHDU SRPDFH FDQ GLOXWH LQVXOLQVHFUHWLRQ
HQHUJ\LQFUHDVHHQHUJ\FRQVXPSWLRQDQGSURPRWH*/3DQG3<<
VHFUHWLRQDIWHULQWHVWLQDOIHUPHQWDWLRQWKHUHE\UHGXFLQJIDWFHOOVDQG 3.3 DF and cancer
HIIHFWLYHO\LPSURYHVREHVLW\
$OOLQDOOWKHLPSURYHPHQWRIREHVLW\E\')PDLQO\FRPHVIURP &DQFHULVRQHRIWKHPDLQSXEOLFSUREOHPVWKDWVHULRXVO\WKUHDWHQ
WKHSK\VLFRFKHPLFDOSURSHUWLHVVXFKDV:+&:6&2+&&$&DQG WKHKHDOWKRIKXPDQEHLQJV>@7KHUHIRUHWKHSUHYHQWLRQRIFDQFHU
YLVFRVLW\ZKLFKGHOD\WKHDEVRUSWLRQRIIDWIURPIRRG%RWK6')DQG LVXUJHQW7KHRFFXUUHQFHRIFDQFHULVFORVHO\UHODWHGWRHDWLQJKDELWV
,')KDYHWKHHIIHFWVRIUHGXFLQJREHVLW\ZKLFKLVEHWWHUUHTXLUHVDORW DQGKLJK')GLHWFDQHIIHFWLYHO\SUHYHQWWKHRFFXUUHQFHRIDYDULHW\RI
RIH[SHULPHQWDOUHVHDUFKHV FDQFHUV 7DEOH 
7KH PHFKDQLVP RI ') WR SUHYHQW FDQFHU LV PDLQO\ UHIOHFWHG
LQ WKH IROORZLQJ DVSHFWV   ') KDV :+& DQG 6:& ZKLFK FDQ
3.2 DF and diabetes
LQFUHDVH VWRRO YROXPH DFFHOHUDWH GHIHFDWLRQ WLPH DQG UHGXFH
,Q WKH SDVW IHZ GHFDGHV WKH LQFLGHQFH RI GLDEHWHV HVSHFLDOO\ WKH FRQFHQWUDWLRQ RI FDUFLQRJHQV LQ WKH LQWHVWLQH 'DKO HW DO >@
W\SHGLDEHWHVLVWKHKLJKHVWDFFRXQWLQJIRU%RIDOOGLVHDVHV,W UHSRUWHG WKDW ') FDQ UHGXFH WKH ULVN RI FRORUHFWDO FDQFHU EHFDXVH
KDVJURZQYHU\UDSLGO\DQGWKHQXPEHURISDWLHQWVZLWKGLDEHWHVKDV WKHLQFUHDVHGIHFDOEXONDQGGHFUHDVHGWUDQVLWWLPHWKHUHE\UHGXFLQJ
FRQWLQXHGWRLQFUHDVHJOREDOO\,WZDVPLOOLRQLQDQG WKH FDUFLQRJHQV FRQFHQWUDWLRQ RI FRORUHFWDO HSLWKHOLXP   ') FDQ
UHGXFH WKH FRQFHQWUDWLRQ RI FLUFXODWLQJ KRUPRQHV DQG LQFUHDVH
PLOOLRQLQ7KHJOREDOSUHYDOHQFHRIGLDEHWHVLVH[SHFWHGWREH
H[FUHWLRQ7KHSUHYHQWLRQPHFKDQLVPRIRYDULDQFDQFHUDQGEUHDVW
%E\>@7KHLQFUHDVHLQWKHQXPEHURIGLDEHWLFSDWLHQWV
FDQFHULVPDLQO\WRLQKLELWWKHVHFUHWLRQRIKRUPRQHVDQGUHGXFHWKH
DOVR LQFUHDVHV WKH JOREDO HFRQRPLF EXUGHQ 7KHVH LVVXHV PDNH LW
ELRDYDLODELOLW\RIKRUPRQHV+RHWDO>@VWXGLHVVKRZHGWKDW')
XUJHQWIRUXVWRSUHYHQWDQGWUHDWW\SHGLDEHWHVDVVRRQDVSRVVLEOH
LQIUXLWVDQGYHJHWDEOHVSUHYHQWVIHPDOHEUHDVWFDQFHUE\LQKLELWLQJ
6WXGLHVKDYHIRXQGWKDW')KDVDQLPSRUWDQWUROHLQWKHSUHYHQWLRQ
HVWURJHQ HQ]\PHV DQG UHGXFLQJ HVWURJHQ VHFUHWLRQ ,Q DGGLWLRQ
DQGWUHDWPHQWRIGLDEHWHV
;XHWDO>@UHSRUWHGWKDWLQWDNLQJ')FDQUHGXFHVWHURLGKRUPRQHV
7KHPHFKDQLVPRI')ORZHULQJEORRGJOXFRVHLVDVIROORZV 
DQGKHOSUHJXODWHLQVXOLQJURZWKIDFWRUWKHUHE\UHGXFLQJWKHULVNRI
7KHQHWZRUNVWUXFWXUHRI')FDQIRUPDSK\VLFDOEDUULHUDQGGHOD\ RYDULDQFDQFHU)RUHYHU\J/GD\')LQFUHDVHWKHULVNRIRYDULDQ
WKHGLIIXVLRQRIJOXFRVH)LEURXVSDUWLFOHVDFWDVDSK\VLFDOEDUULHU FDQFHU LV UHGXFHG E\ % UHODWLYH ULVN  % &, ± 
WRJOXFRVHPROHFXOHVDQGHQWUDSPHQWRIJOXFRVHLQWKH¿EHUIRUPLQJ   ') IHUPHQWDWLRQ SURGXFHV 6&)$V VXFK DV EXW\UDWH ZKLFK FDQ
QHWZRUN:KHQWKHVWUXFWXUHRI')EHFRPHVORRVHDQGSRURXVDQG LQKLELWKLVWRQHGHDFHW\ODVHDQGUHODWHGVLJQDOLQJSDWKZD\VLQFXOWXUHG
PRUHVXUIDFHIXQFWLRQDOJURXSVDUHH[SRVHGWKHLQWHUDFWLRQEHWZHHQ FDQFHU FHOOV DQG SURPRWH FDQFHU FHOO DSRSWRVLV >@ 6WXGLHV E\
')DQGJOXFRVHZLOOEHHQKDQFHGDQGWKH*$&ZLOOEHLPSURYHG .DZDNLWDHWDO>@VKRZHGWKDW')VHHPVWRKDYHDQWLLQÀDPPDWRU\
6ULFKDPURHQHWDO>@DQG$VKXWRVKHWDO>@UHSRUWHGWKDW')IURP HIIHFWVWKURXJKWKHSURGXFWLRQRI6&)$VE\JXWEDFWHULDZLWKDQWL
VXQIORZHUVHHGVWDPDULQGDQGFDVVLDVHHGVKDVDSRVLWLYHHIIHFWRQ SUROLIHUDWLRQDQGSURDSRSWRWLFSURSHUWLHVUHVXOWLQJLQUHGXFLQJWKH
UHWDUGLQJJOXFRVHGLIIXVLRQ,QDGGLWLRQ')H[WUDFWHGIURPSRWDWRHV ULVNRIFDQFHU
E\&KHQJHWDO>@KDVDVLJQL¿FDQWHIIHFWRQVORZLQJWKHGLIIXVLRQ
RIJOXFRVH,WLVGXHWRWKHSRURXVRI')DQGWKHH[SRVXUHRIVXUIDFH
3.4 DF and intestinal disease
IXQFWLRQDOJURXSVZKLFKHQKDQFHVWKHLQWHUDFWLRQEHWZHHQ')DQG
JOXFRVH 0LOOHW ,') FDQ GHOD\ JOXFRVH DEVRUSWLRQ DQG KHOS EHWWHU 7KH JDVWURLQWHVWLQDO V\VWHP LV WKH ODUJHVW EDUULHU WLVVXH RI WKH
FRQWUROVHUXPJOXFRVHOHYHOZKLFKLVPDLQO\FDXVHGE\WKHSK\VLFDO KXPDQERG\,QDKHDOWK\VWDWHWKHLQWHVWLQDOHQYLURQPHQWFDQSOD\
EDUULHURI,')>@ 9LVFRXV¿EHUVFDQGHOD\JDVWULFHPSW\LQJDQG D NH\ UROH LQ SURPRWLQJ IRRG GLJHVWLRQ DQG DIIHFWLQJ WKH PXFRVDO

Table 2
'HHGEVQH&(KPUQOGV[RGUQHECPEGT
6[RGUQHECPEGT #PCN[VKECNOGVJQFU 'ZRGTKOGPVCNOQFGN /GEJCPKUO 4GHGTGPEGU
1XCTKCPECPEGT /GVCCPCN[UKU %NKPKECNTGUGCTEJGUQXCTKCPECPEGT 4GFWEGEQPEGPVTCVKQPUQHEKTEWNCVKPIJQTOQPCN =?
2CPETGCVKEECPEGT /GVCCPCN[UKU %NKPKECNTGUGCTEJGURCPETGCVKEECPEGT GUVTQIGPJQTOQPCNRCPETGCVKEECTEKPQIGPGUKU =?
2TQUVCVGECPEGT 2TQURGEVKXGCPCN[UKU %NKPKECNTGUGCTEJGURTQUVCVGECPEGT GUVTQIGPUQTCPFTQIGPU =?
$TGCUVECPEGT %CUGEQPVTQNUVWF[ %NKPKECNTGUGCTEJGUYQOGPDTGCUVECPEGT =?
2TQOQVGVJGNCTIGCOQWPVQHUJQTVEJCKPHCVV[
%QNQTGEVCNECPEGT 1XGTXKGYQHGZRGTKOGPVCNFCVC %NKPKECNTGUGCTEJGUCPFCPKOCNGZRGTKOGPV =?
acids
*GCFCPFPGEMECPEGT %CUGEQPVTQNUVWFKGU %NKPKECNTGUGCTEJGUJGCFCPFPGEMECPEGT =?
 Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10

LPPXQH V\VWHP WKURXJK V\PELRWLF PLFURELRWD ,PEDODQFH RI WKH HVRSKDJHDOPRWLOLW\DQGUHGXFHVWKHIUHTXHQF\RIJDVWURHVRSKDJHDO


JXWPLFURELRWDPD\FDXVHGDPDJHWRWKHLQWHVWLQDOEDUULHUWKHUHE\ UHÀX[DQGKHDUWEXUQSHUZHHN>@
LQFUHDVLQJWKHVXVFHSWLELOLW\WRFHUWDLQGLVHDVHV>@ 7KHLQFLGHQFH ')KDVWKHHIIHFWRISURPRWLQJLQWHVWLQDOKHDOWKEHFDXVHLWFDQ
RILQWHVWLQDOGLVHDVHVLVFORVHO\UHODWHGWROLIHVW\OHDQGGLHW3HRSOH SURWHFWWKHLQWHVWLQDOEDUULHUDQGHQKDQFHWKHLQWHVWLQHOHQJWK 7KH6&)$V
ZKR FRQVXPH PRUH ') FDQ UHJXODWH WKH JXW PLFURELRWD SUHYHQW SURGXFHGE\')IHUPHQWDWLRQFDQLQKLELWLQÀDPPDWRU\IDFWRUVWKHUHE\
FRQVWLSDWLRQ DQG PDLQWDLQ WKH LQWHJULW\ RI WKH LQWHVWLQDO EDUULHU SURWHFWLQJWKHLQWHJULW\RIWKHLQWHVWLQDOEDUULHU &KDSPDQHWDO>@
WKHUHE\SURPRWLQJLQWHVWLQDOKHDOWK VWXGLHVUHSRUWHGWKDW LQFUHDVHGFRQFHQWUDWLRQRIEXW\ULFDFLGKDVDQ
') IHUPHQWDWLRQin vivo FDQ SURPRWH WKH JURZWK RI EHQHILFLDO HIIHFWRQDOOHYLDWLQJXOFHUDWLYHFROLWLV6&)$VIHUPHQWHGE\LQXOLQ
EDFWHULDDQGLQKLELWWKHJURZWKRIKDUPIXOEDFWHULD ZKLFKSOD\DYLWDO LQWKHFRORQFDQUHGXFHPXFRVDOSURLQIODPPDWRU\F\WRNLQHVZKLFK
UROHLQSUHYHQWLQJDQGWUHDWLQJGLVHDVHV.DQJHWDO>@ UHSRUWHGWKDW VKRZDORZHUGHJUHHRIPXFRVDOGDPDJHDQGDGHFUHDVHLQWKHVWLQJ
WKH .RQMDNX IORXU .)  VXSSOHPHQWDWLRQ VLJQLILFDQWO\ UHGXFHV WKH RIFU\SWGDPDJH>@=RKUHHWDO>@UHSRUWHGWKDWWKHLQWDNHRI
REHVHPLFHZHLJKW7KLVLVEHFDXVHWKH.)ERRVWVWKHDEXQGDQFHVRI FRRNHGSHD')LQFUHDVHVWKHDEXQGDQFHRI /DFKQRVSLUDFHDH LQWKH
LQWHVWLQH+LJKHUEXW\UDWHSURGXFLQJSURPRWHVWKH0XFDQG0XF
VRPHREHVLW\UHODWHGEHQH¿FLDOPLFURRUJDQLVPV VXFKDVMegasphaera
H[SUHVVLRQRIWKHPDLQFRPSRQHQWVLQWKHLQWHVWLQDOPXFXVOD\HUVR
elsdenii LQWKHJXWPLFURELRWDRIKLJKIDWGLHWPLFHZKLOHUHGXFHV
LWLVLPSRUWDQWIRUWKHSURWHFWLYHIXQFWLRQRIWKLVOD\HU7KHOHQJWKRI
WKRVHRIKDUPIXOPLFURRUJDQLVPV VXFKDVAlistipes, Alloprevotella,
WKHFRORQDIIHFWVWKHHIIHFWRI')IHUPHQWDWLRQZKLFKWKHLQFUHDVH
Bacteroides acidifaciens DQG Parabacteroides goldsteinii  ,Q
RIWKHFRORQOHQJWKFDQVKRUWHQWKHIHUPHQWDWLRQWLPHDQGLQFUHDVH
DGGLWLRQ ZH IRXQG WKDW VR\EHDQ ,') FDQ SUHYHQW REHVLW\ LQ PLFH
WKHGHJUHHRIIHUPHQWDWLRQ'HUWHWDO>@UHSRUWHGWKDWLQWHVWLQDO
RQ D KLJKIDW GLHW E\ UHJXODWLQJ WKH JXW PLFURELRWD 6R\EHDQ ,')
GLVHDVHVFDQVKRUWHQWKHOHQJWKRIWKHFRORQ2XUWHDPVWXGLHVIRXQG
LQWDNHFDQLQFUHDVHWKHSRWHQWLDOEHQH¿FLDOEDFWHULDLactobacillales,
WKDWWKHLQWDNHRIVR\EHDQ,')FDQLQFUHDVHWKHFRORQOHQJWKRIPLFH
Lactobacillus/DFKQRVSLUDFHDH_NK4A136_JURXSUHODWHGWRREHVLW\ VLJQL¿FDQWO\LPSURYHWKHLQWHVWLQDOGDPDJHRIKLJKIDWGLHWLQPLFH
7KH DEXQGDQFH RI Bacteroides_acidifaciens VLJQLILFDQWO\ UHGXFHG DQGPDNHWKHFRORQHSLWKHOLDOFHOOVRIPLFHLQWDFWDQGJREOHWFHOOV
WKHZHLJKWRIPLFHRQDKLJKIDWGLHW=KDQJHWDO>@VKRZHGWKDW LQWDFWDQGXQLIRUP
LQXOLQ WUHDWPHQW FDQ SURPRWH WKH DEXQGDQFH RI SURELRWLF EDFWHULD $ QXPEHU RI VWXGLHV KDYH IRXQG WKDW 6&)$V SURGXFHG E\ ')
Lactobacillus, /DFKQRVSLUDFHDH Phascolarctobacterium DQG WKURXJKWKHIHUPHQWDWLRQRIJXWPLFURELRWDFDQSUHYHQWDQGLPSURYH
BacteroidesLQGLDEHWLFUDWV)DUD]HWDO>@IRXQGWKDWKLJK')GLHW WKHRFFXUUHQFHRIREHVLW\GLDEHWHVFDQFHUDQGLQWHVWLQDOGLVHDVHV
FDQSURPRWHEHQH¿FLDOEDFWHULDUHVXOWLQJLQSURWHFWLQJHIIHFWRQFRORQ 7KHHIIHFWRI')RQSUHYHQWLRQDQGWUHDWPHQWGLVHDVHVLVUHODWHGZLWK
FDQFHU$OIDHWDO>@IRXQGWKDWUHVLVWDQWVWDUFKFDQLPSURYHWKHJXW WKH6&)$V )LJ ')LVIHUPHQWHGE\JXWPLFURELRWDWRSURGXFH
PLFURELRWDRIPLGGOHDJHGDQGHOGHUO\SHRSOHVWLPXODWHWKHJURZWKRI 6&)$VDQG6&)$VFDQSURPRWHWKHLQFUHDVHRI3<<*/3DQG
%DFWHURLGHWHVDQGFKDQJHWKHUDWLRRI)LUPLFXWHV/%DFWHURLGHWHVDQG LQVXOLQVHQVLWLYLW\FDQFHUFHOODSRSWRVLVDQGLQWHVWLQDOLQWHJULW\ZKLFK
WKHUHODWLYHDEXQGDQFHRIEXW\ULFDFLG LVFRQGXFLYHWRGLVHDVHSUHYHQWLRQDQGWUHDWPHQW
') KDV JRRG :+& DQG 6:& ZKLFK FDQ UHOLHYH IXQFWLRQDO
FRQVWLSDWLRQDQGSURPRWHJDVWURLQWHVWLQDOGLJHVWLRQ &KDRHWDO>@ 3.5 Other functions
VKRZHGWKDW')ZLWK:+&DQG6:&FDQLQFUHDVHGHIHFDWLRQDQG
KHOSWRUHOLHYHFRQVWLSDWLRQV\PSWRPVLQFKLOGUHQ,QSDWLHQWVZLWK +P TGEGPV [GCTU YKVJ VJG FGGRGPKPI QH TGUGCTEJGU UQOG QVJGT
QRQHURVLYHJDVWURHVRSKDJHDOUHÀX[GLVHDVHD')ULFKGLHWLPSURYHV HWPEVKQPU QH &( JCXG DGGP FKUEQXGTGF KPENWFKPI TGFWEKPI JGCX[

Gut microbiota
DF Fermentation Feaces
Satiety

SCFAs L cell
PYY
GLP -1

Inflammatory factors Cancer cell Obesity


Mucosal proinflammatory
cytokines Apoptosis
Proliferation Insulin sensitivity
Glucose homeostasis
Intestinal barrier
integrity Cancer Diabetes

Intestinal disease
Fig. 2 6JGGHHGEVQH&(QPRTGXGPVKQPCPFVTGCVOGPVFKUGCUGUKUTGNCVGFYKVJVJG5%(#U
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10 

OGVCNVQZKEKV[VJGTKUMQHECTFKQXCUEWNCTCPFHGOCNGFKUGCUGUCPF &( JCU IQQF RJ[UKEQEJGOKECN CPF HWPEVKQPCN RTQRGTVKGU UQ


KORTQXKPICNNGTIKEU[ORVQOUUWEJCUTJKPKVKU KV RNC[U CP KORQTVCPV TQNG KP VJG FGXGNQROGPV QH HWPEVKQPCN HQQFU
&( KU C PCVWTCN EJGNCVKPI CIGPV YJKEJ ECP EQORNGZ YKVJ NGCF HQQFRCEMCIKPICPFVJGRTQVGEVKQPQHDGPGHKEKCNCEVKXGUWDUVCPEGU
KQPUVQHQTOCRTGEKRKVCVGUQKVECPGHHGEVKXGN[RTGXGPVVJGCDUQTRVKQP (WTVJGTOQTG &( HWPEVKQPCN HQQF JCU VJG GHHGEV QH NQYGTKPI DNQQF
QH NGCF KP VJG ICUVTQKPVGUVKPCN VTCEV &( ECP FGNC[ QT TGFWEG VJG HCVCPFDNQQFUWICTCPFIQQFHQQFRTQRGTVKGUQHDGXGTCIGUCPFLCO
CDUQTRVKQP QH JGCX[ OGVCNU CPF ECTEKPQIGPU KP VJG KPVGUVKPG 6JG RTQFWEVU DCMGT[ OGCV CPF QVJGT RTQFWEVU KPENWFKPI VCUVG EQNQT
NCZCVKXG GHHGEV QH &( VJG TGUKFGPEG VKOG QH HGEGU CPF JCTOHWN JCTFPGUU VGZVWTG IGN ECRCEKVKGU CPF OGEJCPKECN RTQRGTVKGU YJKEJ
UWDUVCPEGUKP VJG KPVGUVKPG KU UJQTVGPGF YJKEJ ITGCVN[ TGFWEGUVJG DGECWUG&(JCUVJGIQQFRJ[UKEQEJGOKECNRTQRGTVKGU 6CDNG 
JCTO QH JCTOHWN UWDUVCPEGU VQ VJG JWOCP DQF[ VJGTGD[ TGFWEKPI
VJGTKUMQHRCVJQNQIKECNEJCPIGUKPECPEGT=?5&(EQPVCKPUUQOG 4. Conclusion and future research
ECTDQZ[NUKFGEJCKPITQWRUYJKEJECPTGFWEGVJGVQZKEKV[QHJGCX[
OGVCNU+&(JCUCPGVYQTMUVTWEVWTGVJCVHQTOURJ[UKECNCFUQTRVKQP 6JGUG RJ[UKEQEJGOKECN RTQRGTVKGU QH &( KPENWFG 9*%
KPFKECVKPIVJCV+&(ECPJKPFGTVJGCDUQTRVKQPQHNGCFKQPUCPFTGFWEG íI/I 95% íO./I 1*% íI/I 
VJG TKUM QH NGCF RQKUQPKPI -JQVKOEJGPMQ GV CN =? UJQYGF VJCV )#% íOOQN/I %#% íOI/I CPFXKUEQUKV[
NQYGUVGTKHKGFRGEVKPJKIJGUVGTKHKGFRGEVKPCPFRGEVKPECNEKWOIGN YJKEJRNC[CXKVCNTQNGKPRTGXGPVKPIFKUGCUGU)QQF9*%95%
RCTVKENGUJCXGIQQFCFUQTRVKQPQHNGCFKQPU9CPIGVCN=?UVWFKGF 1*%CPF%#%JCXGCPGHHGEVQPQDGUKV[XKUEQUKV[CPF)#%JCXG
VJCVUQ[DGCP+&(GZJKDKVUJKIJGTNGCF ++ KQPCFUQTRVKQPECRCEKV[CPF VJGITGCVGUVKORCEVQPFKCDGVGUCFUQTRVKQPECRCEKV[JCUCPGHHGEVQP
CFUQTRVKQPQEEWTUQPVJGUWTHCEGQHſDGTRCTVKENGU ECPEGT9*%95%CPFXKUEQUKV[JCXGDGPGſVUHQTKPVGUVKPCNJGCNVJ
5QNKOCP GV CN =? HQWPF VJCV &( ECP TGFWEG VJG TKUM QH &(KUCNUQYKFGN[WUGFKPHWPEVKQPCNHQQFU+PVJKUTGXKGY&(HTQO
ECTFKQXCUEWNCT FKUGCUG 6JKU KU DGECWUG 5%(#U RTQFWEGF D[ HTWKVUJCXGVJGJKIJEQPVGPVYJKEJCTGGCU[VQQDVCKPCPFKPGZRGPUKXG
&( HGTOGPVCVKQP ECP KPETGCUG VJG FGITGG QH CEKFKHKECVKQP QH VJG 6JG&(HTQOITCKPUCPFDGCPUCNUQJCUVJGCDQXGCFXCPVCIGU6JQUG
GPXKTQPOGPV KP VJG EQNQP ECXKV[ YJKNG NQY R* ECP TGFWEG VJG ECPDGWUGFCUCIQQFUQWTEGQH&(KPVJGHWVWTGHQTVJGFGXGNQROGPV
UQNWDKNKV[QHHTGGDKNGCEKFUCPFVJGKTEQPXGTUKQPVQUGEQPFCT[DKNG QHJGCNVJHQQF
CEKFUCPFKPETGCUGDKNGGZETGVKQP=? *QYGXGT VJGTG CTG CNUQ UJQTVEQOKPIU QH &( UVWFKGU KP UQOG
0CICVCGVCN=?TGRQTVGFVJCVUQ[DGCP&(TGFWEGUOGPUVTWCN CURGEVU (KTUV QH CNN VJG FKHHGTGPV RQN[OGTK\CVKQP FGITGG QH &(
RCKP VJG TGCUQP KU VJCV &( KPVCMG ECP KPJKDKV GUVTQIGP GP\[OGU OC[ JCXG C RTGXGPVKXG GHHGEV QP FKHHGTGPV FKUGCUGU GURGEKCNN[ VJG
CPFTGFWEGVJGUGETGVKQPQHGUVTQIGPCPFUVGTQKFJQTOQPGUVJGTGD[ RQN[OGTK\CVKQPFGITGGQH&(HTQOFKHHGTGPVUQWTEGUKUCNUQFKHHGTGPV
TGFWEKPIF[UOGPQTTJGCCPFJGNRKPIVQTGFWEGVJGTKUMQHKNNPGUU 6JG RQN[OGTK\CVKQP FGITGG QH FKHHGTGPV &(U KU UVKNN WPENGCT QP
<JCPI GV CN =? HQWPF VJCV &( JCU C UKIPKHKECPV KPJKDKV VJG FKUGCUG RTGXGPVKQP OGEJCPKUO UQ VJG UVTWEVWTG QH FKHHGTGPV
CNNGTIKE TJKPKVKU EQORNKECVGF YKVJ CNNGTIKE KPHNCOOCVKQP QH RQN[OGTK\CVKQPFGITGGUQH&(HTQOFKHHGTGPVUQWTEGUKUPGEGUUCT[HQT
CUVJOC6JGTGUWNVUUJQYGFVJCVRTQRGTHKDGTKPVCMGECPTGIWNCVG HWTVJGTTGUGCTEJQPFKUGCUGRTGXGPVKQP5GEQPFN[VJGRTGXGPVKQPQH
IWV OKETQDKQVC CPF OCKPVCKP KOOWPG UVCDKNKV[ VQ RTGXGPV CPF FKUGCUGUD[&(KUPQVNKOKVGFVQVJQUGOGPVKQPGFKPVJKUTGXKGY+P
VTGCVCNNGTIKEFKUGCUGU RCTVKEWNCTVJGRTGXGPVKQPCPFVTGCVOGPVQHQVJGTFKUGCUGUD[TGIWNCVKPI

Table 3
6JGRTQFWEVUQH&(KPHWPEVKQPCNHQQF
%NCUUKſECVKQP 2TQFWEVU (KDGTV[RG &(5QWTEGU 2J[UKEQEJGOKECNRTQRGTVKGU (WPEVKQPCN 4GHGTGPEGU
+PETGCUGFſDGTKPVCMGHQTVKH[KPIKPITGFKGPV
&KGVCT[ſDGT[QIJWTV +&( 9JGCVQTQCVſDGTU 9*%95% =?
$GXGTCIGU DGVVGTVCUVGCPFƀCXQT
CPFLCOU $GXGTCIGUCPF %JKPGUGECDDCIG 8KUEQUKV[9*%1*%
5&( 2TGDKQVKECPFJ[RQIN[EGOKE =?
PWVTCEGWVKECNRTQFWEVU YCUVG )#%%#%

2GCFKGVCT[ſDGTECMGU DF 2GC 9*%95%1*% /QTGCVVTCEVKXGEQNQT =?


&CKN[TGSWKTGOGPVUQHCXGTCIGFKGVCT[ſDGTKPVCMGTGFWEGF
&KGVCT[ſDGTEQQMKGU 5&( )WCTIWO 9*%95% =?
JCTFPGUUUGPUQT[CEEGRVCDNG
$CMGT[RTQFWEVU
%QQMKGU DF 9JGCVDTCP 9*%1*%)#%%#% 6JGNQYGTVQVCNEJQNGUVGTQN 6% CPFVTKIN[EGTKFGU 6)  =?
'PJCPEGFVJGFKGVCT[ſDTGKPVCMGKORTQXGFINWVGPOQKUVWTG
#RRNGRQOCEGDTGCF DF #RRNG 9*%95% =?
CDUQTRVKQPYJGCVFQWIJTGUKUVCPEGNQYGTGZVGPUKDKNKV[
$GGHDWTIGTU DF 9JGCV 9*%95%1*%%#% 4GFWEGFJCTFPGUUCPFECNQTKGUKORTQXGFNKIJVPGUU =?
8KGPPCUCWUCIGU DF 5VCTHTWKV 9*%95%1*%%#% +ORTQXGFRJ[UKECNCPFUGPUQT[RTQRGTVKGU =?
/GCVRTQFWEVU $QNQIPCUCWUCIGU DF %KVTWU 9*%95%1*%%#% 4GFWEGFTGUKFWCNPKVTKVGEQPVGPV =?
.QYGTGZRTGUUKDNGOQKUVWTGCPFGPJCPEGFVJGFGXGNQROGPVQH
5CWUCIGU DF (TWKVRGGNU 9*%1*% =?
VJGTOQVQNGTCPVNCEVKECEKFDCEVGTKC
+ORTQXGFUOQQVJUWTHCEGUQHVPGUUVTCPURCTGPE[VGZVWTG
'FKDNGRCRGT +&( %JKPGUGECDDCIG 9*%95%1*% =?
UVTGPIVJCPFRCEMCIKPIRGTHQTOCPEG
5VCTEJDCUGFſNOU DF 1MCTC 9*%95% +ORTQXGFRGTHQTOCPEGTGFWEGFGNQPICVKQPCVDTGCM =?
1VJGTRTQFWEVU &KGVCT[ſDGTKUQNCVG DF %QEQPWVƀCMGU )#%%#% #OGNKQTCVGUINWEQUGCPFNKRKFNGXGNU =?
+PETGCUGF9*%GZRCPUKQPTCVKQ
(QQFIGN 5&( 6QOCVQRGGNU 9*%95% =?
CPFHTGG\GVJCYUVCDKNKV[
8KUEQUKV[9*%ˈ95% 'PJCPEKPIVJGPWVTKVKQPCNXCNWGCPF
(WPEVKQPCNHQQFKPITGFKGPVU DF 9KPG)TCRG2QOCEG =?
)#%%#% KORTQXKPIUVQTCDKNKV[QHVJGRTQFWEVU
 Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10

VJGIWVOKETQDKQVCTGSWKTGUCNQVQHGZRGTKOGPVCNTGUGCTEJGU6JKTFVJG =? 6* <JCPI ; ;CPI ; .KCPI GV CN $GPGHKEKCN GHHGEV QH KPVGUVKPCN
KPVCMGQH&(CPFVJGV[RGQH&(YKNNJCXGCPKORCEVQPVJGRTGXGPVKQP HGTOGPVCVKQP QH PCVWTCN RQN[UCEEJCTKFGU 0WVTKGPVU      
JVVRU//FQKQTI//PW
CPF VTGCVOGPV QH FKUGCUGU +V KU PGEGUUCT[ VQ UVWF[ VJG FQUCIG CPF
=? $9GK;.KW:.KPGVCN&KGVCT[HKDGTKPVCMGCPFTKUMQHOGVCDQNKE
V[RG 5&(QT+&( QH&(KPVJGRTGXGPVKQPCPFVTGCVOGPVQHFKUGCUGU U[PFTQOGCOGVCCPCN[UKUQHQDUGTXCVKQPCNUVWFKGU%NKP0WVT    
VQFGVGTOKPGVJGQRVKOCNKPVCMGQHFKHHGTGPVV[RGU#HVGTVJGCDQXG JVVRU//FQKQTI//LENPW
RTQDNGOUCTGUQNXGF&(YKNNDGOQTGEQPFWEKXGVQVJGRTGXGPVKQP =? ,#4QDGTVUQP(&&/QPTGFQP2&[UUGNGTGVCN*[FTCVKQPRTQRGTVKGUQH
CPFCNNGXKCVKQPQHJWOCPFKUGCUGUYJKNGRTQOQVKPIVJGFGXGNQROGPV FKGVCT[ſDTGCPFTGUKUVCPVUVCTEJC'WTQRGCPEQNNCDQTCVKXGUVWF[.96(QQF
5EK6GEJPQN    JVVRU//FQKQTI//HUVN
QHJWOCPJGCNVJ
=? ( :KG ; 9CPI , 9W GV CN +PUQNWDNG FKGVCT[ HKDGTU HTQO #PIGNKEC
MGKUMGKD[RTQFWEV CPF VJGKT HWPEVKQPCN CPF OQTRJQNQIKECN RTQRGTVKGU
%QPƀKEVQHKPVGTGUVUVCVGOGPV 5VCTEJ5V¼TMG  JVVRU//FQKQTI//UVCT
=? ; <JGPI ; .K 2J[UKEQEJGOKECN CPF HWPEVKQPCN RTQRGTVKGU QH EQEQPWV
6JGTGKUPQEQPƀKEVQHKPVGTGUV (Cocos nucifera .  ECMG FKGVCT[ HKDGTU GHHGEVU QH EGNNWNCUG J[FTQN[UKU
CEKFVTGCVOGPVCPFRCTVKENGUK\GFKUVTKDWVKQP(QQF%JGO    
JVVRU//FQKQTI//LHQQFEJGO
Acknowledgments =? *% %JCQ /9 .CK /5 -QPI GV CN %WVQHH XQNWOG QH FKGVCT[ HKDGT
VQ COGNKQTCVG EQPUVKRCVKQP KP EJKNFTGP , 2GFKCVT      
6JKUYQTMYCUUWRRQTVGFD[%JKPC#ITKEWNVWTG4GUGCTEJ5[UVGO JVVRU//FQKQTI//LLRGFU
QH /1( CPF /#4# %#45  #PF YG CNUQ VJCPM VJG 0CVKQPCN =? ,'.CODGTV,#2CTPGNN,/6WPPKENKHHGGVCN%QPUWOKPI[GNNQYRGC
HKDGT TGFWEGU XQNWPVCT[ GPGTI[ KPVCMG CPF DQF[ HCV KP QXGTYGKIJV/QDGUG
'PIKPGGTKPI .CDQTCVQT[ HQT 9JGCV CPF %QTP &GGR 2TQEGUUKPI
CFWNVU KP C YGGM TCPFQOK\GF EQPVTQNNGF VTKCN %NKP 0WVT   
QH %JKPC CPF VJG -G[ .CDQTCVQT[ HQT #ITKEWNVWTCN 2TQFWEV &GGR JVVRU//FQKQTI//LENPW
2TQEGUUKPIHQTUWRRQTV =? 9,&CJN/.5VGYCTV2QUKVKQPQHVJGCECFGO[QHPWVTKVKQPCPFFKGVGVKEU
JGCNVJ KORNKECVKQPU QH FKGVCT[ HKDGT , #ECF 0WVT &KGV     
References JVVRU//FQKQTI//LLCFC
=? 5 /QTQ\QX 8 +UCMQX / -QPQXCNQXC (KDGTGPTKEJGF FKGV JGNRU VQ
=? #OGTKECP#UUQEKCVKQPQH%GTGCN%JGOKUVU(KDGT%QOOKVVGG&KGVCT[HKDGT EQPVTQNU[ORVQOUCPFKORTQXGUGUQRJCIGCNOQVKNKV[KPRCVKGPVUYKVJPQP
FGſPKVKQPEQOOKVVGGFGſPKVKQPQHFKGVCT[ſDGTTGRQTVQHVJGFKGVCT[ſDGT GTQUKXGICUVTQGUQRJCIGCNTGƀWZFKUGCUG9QTNF,)CUVTQGPVGTQ  
FGſPKVKQPEQOOKVVGGVQVJGDQCTFQHFKTGEVQTUQHVJG#OGTKECP#UUQEKCVKQP JVVRU//FQKQTI//YLIXK
QH%GTGCN%JGOKUVU%GTGCN(QQFU9QTNF  Ō =? :<JQW.<JCPI9,,KGVCN8CTKCVKQPKPFKGVCT[UCNVKPVCMGKPFWEGU
=? '5¶PEJG\<CRCVC/8KWFC/CTVQUD,(GTP¶PFG\.ÎRG\GVCN4GUKUVCPV EQQTFKPCVGFF[PCOKEUQHOQPQE[VGUWDUGVUCPFOQPQE[VGRNCVGNGVCIITGICVGU
UVCTEJ TGUKUVCPV UVCTEJ CU HWPEVKQPCN KPITGFKGPV HWPEVKQPCN KPITGFKGPV KPJWOCPUKORNKECVKQPUKPGPFQTICPKPƀCOOCVKQP2NQU1PG    
2QN[UCEEJCTKFGU   JVVRU//FQKQTI//LQWTPCNRQPG
=? /'NNGWEJ&$GFKIKCP14QKUGWZGVCN&KGVCT[ſDTGCPFſDTGTKEJD[ =? . -WPKCM 4* /CTEJGUUCWNV 5VWF[ QH VJG ETQUUNKPMKPI TGCEVKQP
RTQFWEVUQHHQQFRTQEGUUKPIEJCTCEVGTKUCVKQPVGEJPQNQIKECNHWPEVKQPCNKV[ DGVYGGP GRKEJNQTQJ[FTKP CPF UVCTEJ 5VCTEJ5V“TMG    
CPFEQOOGTEKCNCRRNKECVKQPUCTGXKGY(QQF%JGO     JVVRU//FQKQTI//UVCT
JVVRU//FQKQTI//LHQQFEJGO =? (:KG/.K:.CPGVCN/QFKHKECVKQPQHFKGVCT[HKDGTUHTQORWTRNG
=? /,)KFNG[ )';CMWDQX(WPEVKQPCN ECVGIQTKUCVKQPQHFKGVCT[HKDTG KP HNGUJGF RQVCVQGU Heimeiren  YKVJ JKIJ J[FTQUVCVKE RTGUUWTG CPF JKIJ
HQQFUDG[QPFµVROXEOH¶XU‘KPUQNWDNG¶6TGPFU(QQF5EK6GEJ   RTGUUWTGJQOQIGPK\CVKQPRTQEGUUKPICEQORCTCVKXGUVWF[+PPQX(QQF5EK
JVVRU//FQKQTI//LVKHU 'OGTI  JVVRU//FQKQTI//LKHUGV
=? #1NUQP)/)TC[/%%JKWGVCN%JGOKUVT[CPFCPCN[UKUQHUQNWDNG =? * %JGP % <JCQ , .K GV CN 'HHGEVU QH GZVTWUKQP QP UVTWEVWTCN CPF
FKGVCT[ſDGT(QQF6GEJ     RJ[UKEQEJGOKECN RTQRGTVKGU QH UQNWDNG FKGVCT[ HKDGT HTQO PQFGU QH NQVWU
=? /# 'CUVYQQF 6JG RJ[UKQNQIKECN GHHGEVU QH FKGVCT[ HKDGT CP WRFCVG TQQV.96(QQF5EK6GEJPQN  JVVRU//FQKQTI//
#PPW 4GX 0WVT     JVVRU//FQKQTI//CPPWTGX LNYV
PW =? )/-JCP 0/ -JCP <7 -JCP GV CN 'HHGEV QH GZVTCEVKQP OGVJQFU
=? ,/.CVVKOGT/&*CWD'HHGEVUQHFKGVCT[ſDGTCPFKVUEQORQPGPVUQP QP UVTWEVWTCN RJ[UKQEJGOKECN CPF HWPEVKQPCN RTQRGTVKGU QH FKGVCT[ HKDGT
OGVCDQNKEJGCNVJ0WVTKGPVU    JVVRU//FQKQTI// HTQOFGHCVVGFYCNPWVƀQWT(QQF5EK$KQVGEJPQN    
PW JVVRU//FQKQTI//U
=? 9)69KNNCVU,2-PQZ,&/KMMGNUGP2GEVKPPGYKPUKIJVUKPVQCPQNF =? );W,$GK,<JCQGVCN/QFKſECVKQPQHECTTQV Daucus carota.KPP
RQN[OGTCTGUVCTVKPIVQIGN6TGPFU(QQF5EK6GEJ     XCT5CVKXC*QHHO RQOCEGKPUQNWDNGFKGVCT[HKDGTYKVJEQORNGZGP\[OG
JVVRU//FQKQTI//LVKHU OGVJQFWNVTCſPGEQOOKPWVKQPCPFJKIJJ[FTQUVCVKERTGUUWTG(QQF%JGO
=? ,.&QWDNKGT2,9QQF4JGQNQIKECNRTQRGTVKGUQHCSWGQWUUQNWVKQPUQH     JVVRU//FQKQTI//LHQQFEJGO
 DGVCDINWECPHTQOQCVU Avena sativa. %GTGCN%JGO     =? ) <JCQ 4 <JCPI . &QPI GV CN 2CTVKENG UK\G QH KPUQNWDNG FKGVCT[
 HKDGT HTQO TKEG DTCP CHHGEVU KVU RJGPQNKE RTQHKNG DKQCEEGUUKDKNKV[ CPF
=? -/&CXKUQP0,6GORNG%GTGCNHKDGTHTWKVHKDGTCPFV[RGFKCDGVGU HWPEVKQPCN RTQRGTVKGU .96(QQF 5EK 6GEJPQN    
GZRNCKPKPI VJG RCTCFQZ , &KCDGVGU %QORNKECV       JVVRU//FQKQTI//LNYV
JVVRU//FQKQTI//LLFKCEQOR =? $/CVUWJKTQ.'.KNNQ%5¶GP\GVCN%JGOKECNEJCTCEVGTK\CVKQPQHVJG
=? #**KVG0WVTKVKQPCNGRKFGOKQNQI[QHEJTQPKEFKUGCUGCPFFGſPKPI³+HDOWK\ OWEKNCIGHTQOHTWKVUQH2SXQWLD¿FXVLQGLFD%CTDQJ[F2QN[O    
Diet”)NQD(QQF5GE    JVVRU//FQKQTI// JVVRU//FQKQTI//LECTDRQN
 =? 22GGTCLKV0%JKGYEJCP5&GXCJCUVKP'HHGEVUQHRTGVTGCVOGPVOGVJQFUQP
=? &5 .WFYKI ($ *W . 6CRR[ GV CN &KGVCT[ ECTDQJ[FTCVGU TQNG JGCNVJTGNCVGFHWPEVKQPCNRTQRGTVKGUQHJKIJFKGVCT[ſDGTRQYFGTHTQONKOG
QH SWCNKV[ CPF SWCPVKV[ KP EJTQPKE FKUGCUG $/,     TGUKFWGU (QQF %JGO       JVVRU//FQKQTI//
JVVRU//FQKQTI//DOLM LHQQFEJGO
=? , ;W / #JOGFPC (WPEVKQPCN EQORQPGPVU QH ITCRG RQOCEG VJGKT =? /5 /CHK /# $TKEM ' &KOCU GV CN )GPGVKE CTEJKVGEVWTG QH FKGVCT[
EQORQUKVKQP DKQNQIKECN RTQRGTVKGU CPF RQVGPVKCN CRRNKECVKQPU +PV , HKDGT CPF QNKIQUCEEJCTKFG EQPVGPV KP C OKFFNG #OGTKECP RCPGN QH
(QQF 5EK 6GEJ       JVVRU//FQKQTI//L GFKDNG FT[ DGCP 2NCPV )GPQOG      JVVRU//FQKQTI//
Z RNCPVIGPQOG
Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10 9
=? .*WCPI:&KPI;<JCQGVCN/QFKſECVKQPQHKPUQNWDNGFKGVCT[ſDGT =? :*WC5:W/9CPIGVCN'HHGEVUQHJKIJURGGFJQOQIGPK\CVKQPCPF
HTQO ICTNKE UVTCY YKVJ WNVTCUQPKE VTGCVOGPV , (QQF 2TQEGUU 2TGU    JKIJRTGUUWTGJQOQIGPK\CVKQPQPUVTWEVWTGQHVQOCVQTGUKFWGHKDGTU(QQF
 JVVRU//FQKQTI//LHRR %JGO  JVVRU//FQKQTI//LHQQFEJGO
=? . %JGPI : <JCPI ; *QPI %JCTCEVGTKUCVKQP QH RJ[UKEQEJGOKECN CPF =? *;CKEJ*)CTPC$$EJKTGVCN%JGOKECNEQORQUKVKQPCPFHWPEVKQPCN
HWPEVKQPCNRTQRGTVKGUQHUQNWDNGFKGVCT[ſDTGHTQORQVCVQRWNRQDVCKPGFD[ RTQRGTVKGUQHFKGVCT[HKDTGGZVTCEVGFD['PIN[UVCPF2TQUM[OGVJQFUHTQO
GP\[OGCUUKUVGFGZVTCEVKQP+PV,$KQN/CETQOQN   VJG CNIC Ulva lactuca EQNNGEVGF KP 6WPKUKC #NICN 4GU    
JVVRU//FQKQTI//LKLDKQOCE JVVRU//FQKQTI//LCNICN
=? .<JCQ(<JCPI:&KPIGVCN)WVDCEVGTKCUGNGEVKXGN[RTQOQVGFD[ =? <(GPI&9GK5#NCZKGVCN/QFKſGFUQNWDNGFKGVCT[ſDGTHTQODNCEM
FKGVCT[ſDGTUCNNGXKCVGV[RGFKCDGVGU5EKGPEG     DGCP EQCVU YKVJ KVU TJGQNQIKECN CPF DKNG CEKF DKPFKPI RTQRGTVKGU (QQF
JVVRU//FQKQTI//UEKGPEGCCQ *[FTQEQNN  JVVRU//FQKQTI//LHQQFJ[F
=? /2CTXKP$<CJTC-/5CLCFGVCN#RTQURGEVKXGUVWF[QHFKHHGTGPVV[RGUQH =? %(&KGFGTKEMU8#,KFGCPK2J[UKEQEJGOKECNCPFHWPEVKQPCNRTQRGTVKGUQHKPUQNWDNG
FKGVCT[ſDGTCPFTKUMQHECTFKQXCUEWNCTFKUGCUGVGJTCPNKRKFCPFINWEQUGUVWF[ FKGVCT[ſDGTKUQNCVGFHTQO$CODCTCITQWPFPWV Vigna subterranea =.?8GTFE ,
0WVTKGPVU    JVVRU//FQKQTI//PW (QQF5EK    JVVRU//FQKQTI//
=? +0CPFK#5GPIWRVC/)JQUJ'HHGEVUQHFKGVCT[ſDTGUGZVTCEVGFHTQO =? $&W(<JW$:W2J[UKEQEJGOKECNCPFCPVKQZKFCPVRTQRGTVKGUQHFKGVCT[
FGHCVVGF UGUCOG JWUM TKEG DTCP  HNCZUGGF QP J[RGTEJQNGUVGTQOKE TCVU ſDGTUHTQO3KPIMG JWNNNGUUDCTNG[ ƀQWTCUCHHGEVGFD[WNVTCſPGITKPFKPI
$KQCEV%CTDQJ[FT&KGV(KDTG  JVVRU//FQKQTI// $KQCEV%CTDQJ[FT&KGV(KDTG  JVVRU//FQKQTI//
LDEFH LDEFH
=? 6 /CPFKOKMC ) 2CVWTK %'& )W\OCP GV CN 'HHGEVU QH FKGVCT[ =? %4/CTYC#5QWJKT65CDW7UGQHITGGPEJGOKUVT[OGVJQFUKPVJG
DTQEEQNKHKDTGCPFEQTPQKNQPUGTWONKRKFUHCGECNDKNGCEKFGZETGVKQPCPF GZVTCEVKQP QH FKGVCT[ HKDGTU HTQO ECEVWU TCEMGVU Opuntia ficus indica 
JGRCVKE IGPG GZRTGUUKQP KP TCVU (QQF %JGO       UVTWEVWTCN CPF OKETQUVTWEVWTCN UVWFKGU +PV , $NQN /CETQOQN  
JVVRU//FQKQTI//LHQQFEJGO JVVRU//FQKQTI//LKLDKQOCE
=? : .WQ 3 9CPI $ <JGPI GV CN *[FTCVKQP RTQRGTVKGU CPF DKPFKPI =? ;/CNMMK-#WVKQ1*CPPKPGP1CVDTCPEQPEGPVTCVGURJ[UKECNRTQRGTVKGU
ECRCEKVKGUQHFKGVCT[ſDGTUHTQODCODQQUJQQVUJGNNCPFKVUJ[RQNKRKFGOKE QH DGVCINWECP CPF J[RQEJQNGUVGTQNGOKE GHHGEVU KP TCVU %GTGCN %JGO
G H H G E V U  K P  O K E G   ( Q Q F  % J G O   6 Q Z K E Q N                          
JVVRU//FQKQTI//LHEV =? ,5.X:;.KW:2<JCPI%JGOKECNEQORQUKVKQPCPFHWPEVKQPCNEJCTCEVGTKUVKEU
=? %.KW:..KP<9CPGVCN6JGRJ[UKEQEJGOKECNRTQRGTVKGU in vitro QHFKGVCT[ſDGTTKEJRQYFGTQDVCKPGFHTQOEQTGQHOCK\GUVTCY(QQF%JGO  
DKPFKPI ECRCEKVKGU CPF in vivo J[RQEJQNGUVGTQNGOKE CEVKXKV[ QH UQNWDNG  JVVRU//FQKQTI//LHQQFEJGO
FKGVCT[ſDGTGZVTCEVGFHTQOUQ[JWNNU(QQF(WPEV     =? 0*WUUCKP$#2#IWU#</&CNKGVCN&GVGTOKPCVKQPQHHWPEVKQPCN
JVVRU//FQKQTI//%(1( RTQRGTVKGUQHEQEQCYCUVGHTQOEQPEGPVTCVGFEQEQCFTKPM,(QQF/GCU%JCTCEV
=? 5%JCPI:%WK/)WQGVCN+PUQNWDNGFKGVCT[ſDGTHTQORGCTRQOCEG     JVVRU//FQKQTI//U
ECPRTGXGPVJKIJHCVFKGVKPFWEGFQDGUKV[KPTCVUOCKPN[D[KORTQXKPIVJG =? 0%& 4KUM (CEVQT %QNNCDQTCVKQP 0%&4KU%  6TGPFU KP CFWNV DQF[
UVTWEVWTG QH VJG IWV OKETQDKQVC , /KETQDKQN $KQVGEJPQN      OCUUKPFGZKPEQWPVTKGUHTQOVQCRQQNGFCPCN[UKUQH
ŌJVVRU//FQKQTI//LOD RQRWNCVKQPDCUGF OGCUWTGOGPV UVWFKGU YKVJ  OKNNKQP RCTVKEKRCPVU
=? && )CNNCJGT -, 9QQF %/ )CNNCJGT GV CN +PVGUVKPCN EQPVGPVU 6JG .CPEGV        JVVRU//FQKQTI//5
UWRGTPCVCPVXKUEQUKV[QHTCVUHGFQCVDCUGFOWHſPUCPFEGTGCNRTQFWEVU%GTGCN   :
%JGO    JVVRU//FQKQTI//%%*'/ =? '5 /QWUVCHC 2 (TQIWGN (TQO QDGUKV[ IGPGVKEU VQ VJG HWVWTG QH
=? + 7NNCJ 6 ;KP 5 :KQPI GV CN 'HHGEVU QH VJGTOCN RTGVTGCVOGPV QP RGTUQPCNK\GFQDGUKV[VJGTCR[0CV4GX'PFQETKPQN    
RJ[UKEQEJGOKECNRTQRGTVKGUQHPCPQUK\GFQMCTC UQ[DGCPTGUKFWG KPUQNWDNG JVVRU//FQKQTI//PTGPFQ
FKGVCT[ſDGTRTGRCTGFD[YGVOGFKCOKNNKPI,(QQF'PI   =? ($ /QTGN 3 &CK , 0K GV CN Į)CNCEVQQNKIQUCEEJCTKFGU FQUG
JVVRU//FQKQTI//LLHQQFGPI FGRGPFGPVN[TGFWEGCRRGVKVGCPFFGETGCUGKPƀCOOCVKQPKPQXGTYGKIJVCFWNVU
=? 5//QMJVCT*/5YCKNCO'5'ODCD[2J[UKEQEJGOKECNRTQRGTVKGU ,0WVT    JVVRU//FQKQTI//LP
PWVTKVKQPCNXCNWGCPFVGEJPQHWPEVKQPCNRTQRGTVKGUQHIQNFGPDGTT[ Physalis =? ( +UMGP 5 -NCWU / 1UVGTJQHH GV CN 'HHGEVU QH NQPIVGTO UQNWDNG XU
peruviana YCUVGRQYFGTEQPEKUGVKVNGEQORQUKVKQPQHIQNFGPDGTT[LWKEGYCUVG KPUQNWDNGFKGVCT[ſDGTKPVCMGQPJKIJHCVFKGVKPFWEGFQDGUKV[KP%$./,
(QQF%JGO  JVVRU//FQKQTI//LHQQFEJGO OKEG , 0WVT $KQEJGO       JVVRU//FQKQTI//
=? 8 5WT[CPVK 6 -WUWOCPKPIUKJ ;5 4WOKPIV[CU 2J[UKEQEJGOKECN LLPWVDKQ
RTQRGTVKGUQHFKGVCT[ſDGTUHTQOArtocarpus camansKHTWKV+QR%QPHGTGPEG =? 5%JCPI:%WK/)WQGVCN+PUQNWDNGFKGVCT[ſDGTHTQORGCTRQOCEG
  JVVRU//FQKQTI//:/193/1/ ECPRTGXGPVJKIJHCVFKGVKPFWEGFQDGUKV[KPTCVUOCKPN[D[KORTQXKPIVJG
=? .+ :WGNK *' :KWNK .8 ;WCPRKPI GV CN 'ZVTCEVKQP CPF HWPEVKQPCN UVTWEVWTG QH VJG IWV OKETQDKQVC , /KETQDKQN $KQVGEJPQN     
RTQRGTVKGUQHYCVGTUQNWDNGFKGVCT[ſDGTHTQOCRRNGRQOCEG,(QQF2TQEGUU JVVRU//FQKQTI//LOD
'PI    JVVRU//FQKQTI//LHRG =? ( .W ; .KW $ .K 1MCTC FKGVCT[ HKDGT CPF J[RQIN[EGOKE GHHGEV QH
=? # %GTFC6CRKC / FG . 2ÃTG\%JCDGNC ,  2ÃTG\NXCTG\ GV CN QMCTC HQQFU $KQCEV %CTDQJ[FT &KGV (KDTG      
8CNQTK\CVKQP QH RQOCEG RQYFGT QDVCKPGF HTQO PCVKXG /GZKECP CRRNG JVVRU//FQKQTI//LDEFH
(Malus domestica XCT TC[CFC  EJGOKECN VGEJPQHWPEVKQPCN CPF =? #5TKEJCOTQGP8%JCXCUKVIn vitro TGVCTFCVKQPQHINWEQUGFKHHWUKQPYKVJ
CPVKQZKFCPV RTQRGTVKGU 2NCPV (QQF *WO 0WVT       IWO GZVTCEVGF HTQO OCNXC PWV UGGFU RTQFWEGF KP 6JCKNCPF (QQF %JGO
JVVRU//FQKQTI//U     JVVRU//FQKQTI//LHQQFEJGO
=? $6CQ;(C[KP.*CPI2JGPQNKERTQſNGCPFKPXKVTQCPVKQZKFCPVECRCEKV[ =? /#UJWVQUJ,5JKXGUJIn vitro RQUVRTCPFKCNINWEQUGNQYGTKPIGHHGEVUQHFKGVCT[
QHKPUQNWDNGFKGVCT[ſDGTRQYFGTUHTQOEKVTWU Citrus junos5KGDGZ6CPCMC  HKDGTUKUQNCVGFHTQOTamarindus indicaCPFCassia fistulaUGGFU#OGTKECP,
RQOCEGCUCHHGEVGFD[WNVTCſPGITKPFKPI,#IT(QQF%JGO     (QQF6GEJ    JVVRU//FQKQTI//CLHV
JVVRU//FQKQTI//LHD =? . %JGPI : <JCPI ; *QPI %JCTCEVGTKUCVKQP QH RJ[UKEQEJGOKECN CPF
=? ,;W/#JOGFPC(WPEVKQPCNEQORQPGPVUQHITCRGRQOCEGVJGKTEQORQUKVKQP HWPEVKQPCNRTQRGTVKGUQHUQNWDNGFKGVCT[ſDTGHTQORQVCVQRWNRQDVCKPGFD[
DKQNQIKECNRTQRGTVKGUCPFRQVGPVKCNCRRNKECVKQPU+PV,(QQF5EK6GEJ   GP\[OGCUUKUVGFGZVTCEVKQP+PV,$KQN/CETQOQN  
JVVRU//FQKQTI//LZ JVVRU//FQKQTI//LKLDKQOCE
=? 9 <JCPI ) <GPI ; 2CP GV CN 2TQRGTVKGU QH UQNWDNG FKGVCT[ HKDGT =? /.$CPIQWTC,0UQT#VKPFCPC-<JWGVCN2QVGPVKCNJ[RQIN[ECGOKE
RQN[UCEEJCTKFGHTQORCRC[CRGGNQDVCKPGFVJTQWIJCNMCNKPGQTWNVTCUQWPF GHHGEVU QH KPUQNWDNG HKDTGU KUQNCVGF HTQO HQZVCKN OKNNGVU =Setaria italic
CUUKUVGF CNMCNKPG GZVTCEVKQP %CTDQJ[F 2QN[O     .  2 $GCWXQKU? +PV , (QQF 5EK 6GEJ      
JVVRU//FQKQTI//LECTDRQN JVVRU//FQKQTI//LZ
=? ;.KW%(CP/6KCPGVCN'HHGEVQHFT[KPIOGVJQFUQPRJ[UKEQEJGOKECN =? 4/(GPI;0<QPI5/%CQGVCN%WTTGPVECPEGTUKVWCVKQPKP%JKPC
RTQRGTVKGUCPF in vitroJ[RQIN[EGOKEGHHGEVUQHQTCPIGRGGNFKGVCT[ſDGT,(QQF IQQFQTDCFPGYUHTQOVJG)NQDCN%CPEGT5VCVKUVKEU!%CPEGT%QOOWP
2TQEGUU2TGUGTX    GJVVRU//FQKQTI//LHRR     JVVRU//FQKQTI//U
10 Y. He et al. / Food Science and Human Wellness 11 (2022) 1-10

=? 5% *Q 5< %JGPI %: <JCPI GV CN 'HHGEV QH FKGVCT[ HKDGT KPVCMG ſDGT,(QQF5EK6GEJ    JVVRU//FQKQTI//U
QP DTGCUV ECPEGT TKUM CEEQTFKPI VQ GUVTQIGP CPF RTQIGUVGTQPG TGEGRVQT 
UVCVWU 'WT , %NKP 0WVT       JVVRU//FQKQTI// =? )# 5QNKOCP &KGVCT[ HKDGT CVJGTQUENGTQUKU CPF ECTFKQXCUEWNCT FKUGCUG
GLEP 0WVTKGPVU    JVVRU//FQKQTI//PW
=? * :W ; &KPI : :KP GV CN &KGVCT[ HKDGT KPVCMG KU CUUQEKCVGF YKVJ C =? ,9#PFGTUQP2$CKTF4*&CXKUGVCN*GCNVJDGPGſVUQHFKGVCT[ſDGT0WVT
TGFWEGFTKUMQHQXCTKCPECPEGTCFQUGTGURQPUGOGVCCPCN[UKU0WVT4GU 4GX  JVVRU//FQKQTI//LZ
  JVVRU//FQKQTI//LPWVTGU =? %0CICVC-*KTQMCYC05JKOK\WGVCN#UUQEKCVKQPUQHOGPUVTWCNRCKP
=? , %JGP . 8KVGVVC +PHNCOOCVKQPOQFWNCVKPI GHHGEV QH DWV[TCVG KP VJG YKVJKPVCMGUQHUQ[HCVCPFFKGVCT[ſDGTKP,CRCPGUGYQOGP'WT,%NKP
RTGXGPVKQP QH EQNQP ECPEGT D[ FKGVCT[ HKDGT %NKP %QNQTGEVCN %CPE 0WVT  JVVRU//FQKQTI//ULGLEP
    JVVRU//FQKQTI//LENEE =? <<JCPI.5JK92CPIGVCN&KGVCT[HKDGTKPVCMGTGIWNCVGUKPVGUVKPCN
=? &-CYCMKVC;%#.GG(6WTCVKGVCN&KGVCT[HKDGTKPVCMGCPFJGCF OKETQHNQTC CPF KPJKDKVU QXCNDWOKPKPFWEGF CNNGTIKE CKTYC[ KPHNCOOCVKQP
CPFPGEMECPEGTTKUMCRQQNGFCPCN[UKUKPVJGKPVGTPCVKQPCNJGCFCPFPGEM KPCOQWUGOQFGN2NQU1PG    JVVRU//FQKQTI//
ECPEGTGRKFGOKQNQI[EQPUQTVKWO+PV,%CPEGT     LQWTPCNRQPG
JVVRU//FQKQTI//KLE =? 0 6QOKE $ &QLPQX , /KQEKPQXKE GV CN 'PTKEJOGPV QH [QIJWTV YKVJ
=? $ <JGPI * 5JGP * *CP GV CN &KGVCT[ HKDGT KPVCMG CPF TGFWEGF KPUQNWDNGFKGVCT[ſDGTHTQOVTKVKECNGŌCUGPUQT[RGTURGEVKXG.96(QQF5EK
TKUM QH QXCTKCP ECPEGT C OGVCCPCN[UKU 0WVT ,       6GEJPQN  JVVRU//FQKQTI//LNYV
JVVRU//FQKQTI//U =? 5; 2CTM -; ;QQP 'P\[OCVKE RTQFWEVKQP QH UQNWDNG FKGVCT[ HKDGT
=? %*9CPI%3KCQ4%9CPIGVCN&KGVCT[ſDGTKPVCMGCPFRCPETGCVKE HTQO VJG EGNNWNQUG HTCEVKQP QH %JKPGUG ECDDCIG YCUVG CPF RQVGPVKCN
ECPEGTTKUMCOGVCCPCN[UKUQHGRKFGOKQNQIKEUVWFKGU5EK4GR     WUG CU C HWPEVKQPCN HQQF UQWTEG (QQF 5EK $KQ      
JVVRU//FQKQTI//UTGR JVVRU//FQKQTI//U
=? / &GUEJCUCWZ % 2QWEJKGW / *KU GV CN &KGVCT[ VQVCN CPF KPUQNWDNG =? . $GNIJKVJ(GPFTK ( %JCCTK ( -CNNGN GV CN 2GC CPF DTQCF DGCP
HKDGT KPVCMGU CTG KPXGTUGN[ CUUQEKCVGF YKVJ RTQUVCVG ECPEGT TKUM , 0WVT RQFU CU C PCVWTCN UQWTEG QH FKGVCT[ HKDGT VJG KORCEV QP VGZVWTG CPF
    JVVRU//FQKQTI//LP UGPUQT[ RTQRGTVKGU QH ECMG , (QQF 5EK      
=? # 6CLCFFKPK # 2QWT\CPF < 5CPCCV GV CN &KGVCT[ TGUKUVCPV UVCTEJ JVVRU//FQKQTI//
EQPVCKPGFHQQFUCPFDTGCUVECPEGTTKUMCECUGEQPVTQNUVWF[KPPQTVJYGUVQH =? &/WFIKN5$CTCM$5-JCVMCT%QQMKGVGZVWTGURTGCFTCVKQCPFUGPUQT[
+TCP#UKCP2CE,%CPEGT2     CEEGRVCDKNKV[QHEQQMKGUCUCHWPEVKQPQHUQNWDNGFKGVCT[ſDGTDCMKPIVKOG
=? $.2QQN<QDGN,5CWGT1XGTXKGYQHGZRGTKOGPVCNFCVCQPTGFWEVKQPQH CPF FKHHGTGPV YCVGT NGXGNU .96(QQF 5EK 6GEJ    
EQNQTGEVCNECPEGTTKUMD[KPWNKPV[RGHTWEVCPU,0WVT    5 JVVRU//FQKQTI//LNYV
JVVRU//FQKQTI//LP/5 =? ;*QPI9<,WP:,KCPGVCN&GXGNQROGPVQHVJGFKGVCT[ſDGTHWPEVKQPCN
=? ;$-CPI;.K;&WGVCN-QPLCMWƀQWTTGFWEGUQDGUKV[KPOKEGD[ HQQFCPFUVWFKGUQPKVUVQZKEQNQIKECNCPFRJ[UKQNQIKERTQRGTVKGU(QQF%JGO
OQFWNCVKPIVJGEQORQUKVKQPQHVJGIWVOKETQDKQVC+PV,1DGUKV[     6QZKE    JVVRU//FQKQTI//LHEV
JVVRU//FQKQTI//UZ =? 3.W*.KW39CPIGVCN5GPUQT[CPFRJ[UKECNSWCNKV[EJCTCEVGTKUVKEU
=? 3 <JCPI * ;W : :KCQ GV CN +PWNKPV[RG HTWEVCP KORTQXGU FKCDGVKE QHDTGCFHQTVKſGFYKVJCRRNGRQOCEGWUKPIHW\\[OCVJGOCVKECNOQFGN+PV,
RJGPQV[RG CPF IWV OKETQDKQVC RTQHKNGU KPTCVU 2GGT,      (QQF5EK6GEJ    JVVRU//FQKQTI//KLHU
JVVRU//FQKQTI//RGGTL =? .6%CTXCNJQ/#2KTGU,%$CNFKPGVCN2CTVKCNTGRNCEGOGPVQHOGCV
=? $(CTC\'2JKNNKR20CKNNKYGVCN&KGVCT[ſDGTVTGCVOGPVEQTTGEVUVJG CPFHCVYKVJJ[FTCVGFYJGCVHKDGTKPDGGHDWTIGTUFGETGCUGUECNQTKEXCNWG
EQORQUKVKQPQHIWVOKETQDKQVCRTQOQVGU5%(#RTQFWEVKQPCPFUWRRTGUUGU YKVJQWV TGFWEKPI VJGHGGNKPIQH UCVKGV[CHVGT EQPUWORVKQP/GCV 5EK
EQNQP ECTEKPQIGPGUKU )GPGU       JVVRU//FQKQTI//  JVVRU//FQKQTI//LOGCVUEK
IGPGU =? 8.#PIGNGU25#TCEGNK4++UTCGNGVCN%JGOKECNRJ[UKECNCPFUGPUQT[
=? /,#NHC&5VTCPI256CRRKCGVCN#TCPFQOK\GFVTKCNVQFGVGTOKPG RTQRGTVKGU QH 8KGPPC UCWUCIGU HQTOWNCVGF YKVJ C UVCTHTWKV FKGVCT[ HKDGT
VJG KORCEV QH C FKIGUVKQP TGUKUVCPV UVCTEJ EQORQUKVKQP QP VJG IWV EQPEGPVTCVG,(QQF5EK6GEJ  JVVRU//FQKQTI//
OKETQDKQOGKPQNFGTCPFOKFCIGCFWNVU%NWP0WVT     U
JVVRU//FQKQTI//LENPW =? ,/(GTPCPFG\)KPGU,(GTPCPFG\.QRG\'5C[CU$CTDGTCGVCN'HHGEV
=? /#5%JCROCP/()TCJP/#$Q[NG$WV[TCVGQZKFCVKQPKUKORCKTGF QHUVQTCIGEQPFKVKQPUQPSWCNKV[EJCTCEVGTKUVKEUQHDQNQIPCUCWUCIGUOCFG
KP VJG EQNQPKE OWEQUC QH UWHHGTGTU QH SWKGUEGPV WNEGTCVKXG EQNKVKU )WV YKVJEKVTWUſDGT,(QQF5EK    JVVRU//FQKQTI//
  JVVRU//FQKQTI//IWV LVDZ
=? &, 4QUG /6 &GOGQ # -GUJCXCT\KCP +PHNWGPEG QH FKGVCT[ HKDGT QP =? ,&ÈC\8GNC#6QVQUCWU/.2ÃTG\%JCDGNC+PVGITCVKQPQHCITQKPFWUVTKCN
KPƀCOOCVQT[DQYGNFKUGCUGCPFEQNQPECPEGTKORQTVCPEGQHHGTOGPVCVKQP EQRTQFWEVUCUHWPEVKQPCNHQQFKPITGFKGPVUECEVWURGCT 2SXQWLD¿FXVLQGLFD)
RCVVGTP 0WVT 4GX       JVVRU//FQKQTI//L ƀQWTCPFRKPGCRRNG Ananas comosus RGGNƀQWTCUſDGTUQWTEGKPEQQMGF
VDZ UCWUCIGUKPQEWNCVGFYKVJNCEVKECEKFDCEVGTKC,(QQF2TQEGUU2TGUGTX  
=? * <QJTG ( ,CPGNNG + *[WP GV CN &KGVCT[ RGC HKDGT UWRRNGOGPVCVKQP  JVVRU//FQKQTI//LHRR
KORTQXGU IN[EGOKC CPF KPFWEGU EJCPIGU KP VJG EQORQUKVKQP QH IWV =? 69CPI</C#PQXGNKPUQNWDNGFKGVCT[HKDGTDCUGFGFKDNGRCRGTHTQO
OKETQDKQVCUGTWOUJQTVEJCKPHCVV[CEKFRTQſNGCPFGZRTGUUKQPQHOWEKPUKP %JKPGUGECDDCIG%GNNWNQUG    JVVRU//FQKQTI//
INWEQUGKPVQNGTCPVTCVU0WVTKGPVU    JVVRU//FQKQTI// U
PW =? <3 (W / 9W :; *CP GV CN 'HHGEV QH QMCTC FKGVCT[ HKDGT QP VJG
=? %&GTV/2JKNNKRUQP.*QNOGVCN2TGXGPVKXGCPFVJGTCRGWVKEGHHGEVUQH RTQRGTVKGUQHUVCTEJDCUGFHKNOU5VCTEJ5VTMG    
PKVTKVGUWRRNGOGPVCVKQPKPGZRGTKOGPVCNKPƀCOOCVQT[DQYGNFKUGCUG4GFQZ JVVRU//FQKQTI//UVCT
$KQN  JVVRU//FQKQTI//LTGFQZ =? 626TKPKFCF#%/CNNKNNKP&*8CNFG\GVCN&KGVCT[ſDGTHTQOEQEQPWV
=? # 1\GT 4GOQXCN QH 2D ++  KQPU HTQO CSWGQWU UQNWVKQPU D[ UWNRJWTKE HNQWT C HWPEVKQPCN HQQF +PPQX (QQF 5EK 'OGTI      
CEKFVTGCVGF YJGCV DTCP , *C\CTF /CVGT       JVVRU//FQKQTI//LKHUGV
JVVRU//FQKQTI//LLJC\OCV =? ;0KW3:KC/)WGVCN+PVGTRGPGVTCVKPIPGVYQTMIGNUEQORQUGFQH
=? /-JQVKOEJGPMQ8-QXCNGX;-JQVKOEJGPMQ'SWKNKDTKWOUVWFKGUQH IGNCVKPCPFUQNWDNGFKGVCT[HKDGTUHTQOVQOCVQRGGNU(QQF*[FTQEQNN
UQTRVKQPQHNGCF ++ KQPUD[FKHHGTGPVRGEVKPEQORQWPFU,*C\CTF/CVGT  JVVRU//FQKQTI//LHQQFJ[F
  JVVRU//FQKQTI//LLJC\OCV =? 9)2QOCEG2%QORQWPF&(KDGT&GXGNQROGPVQHCPVKQZKFCPVFKGVCT[
=? * 9CPI 6 *WCPI <% 6Q GV CN 6JG CFUQTRVKQP QH NGCF ++  KQPU D[ HKDGTU HTQO YKPG ITCRG RQOCEG CPF VJGKT CRRNKECVKQPU CU HWPEVKQPCN HQQF
F[PCOKEJKIJRTGUUWTGOKETQƀWKFK\CVKQPVTGCVGFKPUQNWDNGUQ[DGCPFKGVCT[ KPITGFKGPVU9KPG)TCRG2QOCEG  

You might also like