Professional Documents
Culture Documents
vii
ix
Forces of Evolution . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 99
5 Natural Selection . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Mutation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Gene Flow . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
100
101
101
Random Genetic Drift . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 101
Evolution in Action: Malaria and the Sickle-Cell Trait . . . . . . . . . . . . . . . . . 103
Check Your Understanding . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 105
Documenting Evolution: Population Genetics
and Hardy-Weinberg Equilibrium . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 107
Check Your Understanding . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 109
Lab Exercise 5.1: Sickle Cell and Balanced Polymorphism . . . . . . . . . . . . . 111
Lab Exercise 5.2: Documenting Evolution: Population Genetics . . . . . . . . 115
Lab Exercise 5.3: Population Genetics Problems. . . . . . . . . . . . . . . . . . . . . 123
xi
xii
xiii
Appendices. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 413
A Cutouts of Beetles, Lab Exercise 1.3 . . . . . . . . . . . . . . . . . . . . . . . . . 413
B Cutouts of DNA and RNA Bases, Amino Acids,
Ribosome and tRNA Molecules, Lab Exercise 2.2 . . . . . . . . . . . . 415
C Cutouts of Chromosomes, Lab Exercise 3.2 . . . . . . . . . . . . . . . . . . . 421
D Latin and Greek Roots for Selected Anatomical Terms. . . . . . . . . . 423
E Fossil Primates: Paleocene through Miocene
(New World primates not included). . . . . . . . . . . . . . . . . . . . . . . . 425
F Fossil Members of the Human Line (Hominids/Hominins) . . . . . . . 427
G Comparison of Homo rudolfensis
and Homo habilis Features . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 429
Glossary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 431
Photo Credits . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 441
Index. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 443
xiv
W
hat is physical anthropology? Physical, or biological, anthropology is a
unique scientific field. While anthropology in general is a social science,
physical anthropology is based upon biological principles using an
evolutionary perspective. Like the other sciences, physical anthropology depends
upon the scientific method to create explanatory theories. Physical anthropology
focuses on human evolutionary history and modern biological variation and
adaptations within the context of human cultures and behaviors. In order to
place humans in context as part of the natural world, physical anthropology
also deals with our closest living relatives, other members of the order Primates.
Physical Anthropologists
Many areas of study and types of professions fall within the realm of physical
anthropology. All such studies are undertaken following the steps of the scientific
method. Following are some examples of physical anthropologists:
1. Paleontologists study fossils to make inferences about human and nonhuman
primate evolutionary history.
Time 1 Time 2
Time 3
References
Andics, A., M. Gacsi,T. Farago, A. Kis, and A. Miklosi. “Voice-sensitive Gould, S. J. “Sex, drugs, disasters and the extinction of dinosaurs,” in
regions in the dog and human brain are revealed by comparative The Flamingo’s Smile. New York: WW Norton and Company,
fMRI.” Current Biology 24 (2014): 574–578. 1985.
Colbert, E. H., R. B. Cowles, and C. M. Bogert. “Temperature
tolerances in the American alligator and their bearing on the
habits, evolution, and extinction of the dinosaurs.” Bulletin
of the AMNH 86 (1946): 331–373.
Section Date
3 What is science?
4 Number the steps of the scientific method in correct order from one to five, and fill in the blanks.
8 What is a population?
11 Explain why variation within each species is important for natural selection.
12 What is wrong with the statement, “Evolution is just a theory”? How would you respond if someone said this to you?
14 What is wrong with the statement, “If humans evolved from apes, why are there still apes”? How would you respond
if someone asked you this question?
15 Why have humans changed more than chimpanzees and gorillas since diverging from our shared common ancestor?
Hypothesis 1: The rooster just feels like crowing at that time every day.
Hypothesis 3: The rooster has an “internal clock” based on its genetically determined circadian rhythm
(natural biological processes occurring on a 24-hour cycle).
a What question are you likely to ask about this phenomenon that would lead you to developing a hypothesis?
b Formulate a testable hypothesis to investigate this phenomenon. It may be in the form of a question or stated
as a prediction.
d Design a simple experiment or set of observations that would allow you to test your (testable) hypothesis.
3 Working with your lab partner(s), think of some phenomenon that you find interesting and would like to investigate.
b What is a question that might lead to developing a hypothesis about this phenomenon?
d Design a simple experiment or set of observations that would allow you to test your (testable) hypothesis.
4 Explain what is scientific about the study of evolution. (Hint: Think about the definition of evolution and how you
would apply the steps of the scientific method.)
"Both the English and Dutch languages will be used and taught
in public schools where parents of the children desire it, and
allowed in Courts of Law.
"I also beg to inform your Honour that the new Government will
take into immediate consideration the possibility of assisting
by loan the occupants of farms who will take the oath of
allegiance to repair any injury sustained by destruction of
buildings or loss of stock during the war, and that no special
war tax will be imposed on farmers to defray the expense of
the war.
{513}
On the 16th of March the following reply came from the Boer
Commandant: "I have the honour to acknowledge receipt of your
Excellency's letter stating what steps your Excellency's
Government is prepared to take in the event of a general and
total cessation of hostilities. I have advised my Government
of your Excellency's said letter; but, after the mutual
exchange of views at our interview at Middelburg on 28th
February last, it will certainly not surprise your Excellency
to know that I do not feel disposed to recommend that the
terms of the said letter shall have the earnest consideration
of my Government. I may add also that my Government and my
chief officers here entirely agree to my views." This ended
the negotiations.
{514}
"He did not particularly admire the Boers. To his mind they
had too much of the conservative element in them; but, judging
between the Afrikanders and the English who went to South
Africa, whilst fully recognizing that among the latter there
were many respectable men, he thought, taking them
collectively, the Boers were the better men. If we wanted to
maintain our rule in South Africa the Boers were the safest
men with whom to be on good terms. What were the Boers ready
to do? As he read the correspondence, they were ready to enter
the area of the British Empire, but only upon terms. Surely
our problem was to find terms honourable to us and to them,
which would lead to South Africa becoming one of those great
commonwealths connected with the Empire such as existed in
Australia and Canada. He suggested that, in the first place,
we should offer a full and absolute amnesty. He urged that the
Orange State and the Transvaal should as soon as possible be
made self-governing colonies. The Orange State was regarded by
every Englishman who had written about it as a model State. As
to the Transvaal, he admitted there was a difficulty, but he
would suggest that the main area of the country should be
separated from the Rand. The Rand might be administered by a
governor, a military governor if they liked, while in the rest
of the country the Dutch would have a majority. If this course
were adopted, instead of our giving some sort of pecuniary aid
to the Transvaalers, they might be paid a reasonable rent for
the Rand district, of which they would be deprived. … They on
that side of the House would be perfectly ready to agree to
the establishment of a provisional government, military or
civil—he should himself prefer Lord Kitchener to Sir Alfred
Milner—to carry on the country while they were arranging for
the colony to be self-governing. They were accustomed to be
told that Sir Alfred Milner was a sort of divine pro-consul.
He believed Sir Alfred Milner to be a most honourable man, and
very intelligent in many walks of life; but the truth was that
he began life as an Oxford don and then became an official in
the Treasury, facts which militated against his success in
practical politics. He believed that a man like Lord Dufferin
would do more for the cause of peace in South Africa than all
our soldiers."
"A long time has elapsed since I have attempted to send to you
any general review of South African affairs. The reason is
twofold. In the first place, I am occupied every day that
passes from morning till night by business, all of which is
urgent, and the amount and variety of which you are doubtless
able to judge from the communications on a great variety of
subjects, which are constantly passing between us. In the next
place, I have always hoped that some definite point would be
reached at which it might be possible to sum up that chapter
of our history which contained the war, and to forecast the
work of administrative reconstruction which must succeed it.
But I am reluctantly forced to the conclusion that there will
be no such dividing line. I have not the slightest doubt of
the ultimate result, but I foresee that the work will be
slower, more difficult, more harassing, and more expensive
than was at one time anticipated. At any rate, it is idle to
wait much longer in the hope of being able to describe a clear
and clean-cut situation. Despite the many other calls upon my
time, and despite the confused character of the present
position, I think it better to attempt to describe, however
roughly and inadequately, the state of things as it exists
to-day.
"It is no use denying that the last half-year has been one of
retrogression. Seven months ago this Colony was perfectly
quiet, at least as far as the Orange River. The southern half
of the Orange River Colony was rapidly settling down, and even
a considerable portion of the Transvaal, notably the
south-western districts, seemed to have definitely accepted
British authority, and to rejoice at the opportunity of a
return to orderly government, and the pursuits of peace.
To-day the scene is completely altered. It would be
superfluous to dwell on the increased losses to the country
caused by the prolongation of the struggle, and by the form
which it has recently assumed. The fact that the enemy are now
broken up into a great number of small forces, raiding in
every direction, and that our troops are similarly broken up
in pursuit of them, makes the area of actual fighting, and
consequently of destruction, much wider than it would be in
the case of a conflict between equal numbers operating in
large masses.
{515}
Moreover, the fight is now mainly over supplies. The Boers
live entirely on the country through which they pass, not only
taking all the food they can lay hands upon on the farms,
grain, forage, horses, cattle, &c., but looting the small
village stores for clothes, boots, coffee, sugar, &c., of all
which they are in great need. Our forces, on their side, are
compelled to denude the country of everything moveable, in
order to frustrate these tactics of the enemy. No doubt a
considerable amount of the stock taken by us is not wholly
lost, but simply removed to the refugee camps, which are now
being established at many points along the railway lines. But
even under these circumstances, the loss is great, through
animals dying on the route, or failing to find sufficient
grass to live upon when collected in large numbers at the
camps. Indeed, the loss of crops and stock is a far more
serious matter than the destruction of farm buildings, of
which so much has been heard. I say this not at all as an
advocate of such destruction. I am glad to think that the
measure is now seldom if ever resorted to. At the same time,
the destruction of even a considerable number of farms, having
regard to the very rough and inexpensive character of the
majority of these structures in the Orange River Colony and
Transvaal, is a comparatively small item in the total damage
caused by the war to the agricultural community.
{516}
"As the guerrilla warfare swept back over the whole of the
western Transvaal, and practically the whole of the Orange
River Colony, its effect upon the Cape Colony also became very
marked. There was a time, about the middle of last year, when
the bulk of the Dutch population in the Cape Colony, even
those who had been most bitter against us at the outset,
seemed disposed to accept the 'fait accompli,' and were
prepared to acquiesce in the union of all South Africa under
the British flag. Some of them even began to see certain
advantages in such a consummation. The irreconcilable line
taken in the Cape Parliament, during its recent Session from
July to October, was a desperate effort to counteract this
tendency. But I doubt whether it would have succeeded to the
moderate extent to which it has, had it not been for the
recrudescence of the war on the borders of the Colony, and the
embittered character which it assumed. Every act of harshness,
however necessary, on the part of our troops, was exaggerated
and made the most of, though what principally inflamed the
minds of the people were alleged instances of needless cruelty
which never occurred. Never in my life have I read of, much
less experienced, such a carnival of mendacity as that which
accompanied the pro-Boer agitation in this Colony at the end
of last year. And these libels still continue to make
themselves felt. …
The same Blue Book made known the fact that, on the 3d of
April, Sir Alfred Milner applied for and obtained leave of
absence for three months from his duties in South Africa.
also,
CONSTITUTION (GRONDWET) OF THE SOUTH AFRICAN
REPUBLIC.
SOUTH AUSTRALIA.