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Journal of Anxiety Disorders 22 (2008) 1120–1127

The role of trait anxiety in the recognition of emotional


facial expressions
Robbie M. Cooper *, Angela C. Rowe, Ian S. Penton-Voak
University of Bristol, Bristol, UK
Received 25 July 2007; received in revised form 22 November 2007; accepted 26 November 2007

Abstract
Previous work has suggested that elevated levels of trait anxiety are associated with an increased ability to accurately recognize
the facial expression of fear. However, to date, recognition has only been assessed after viewing periods of 10 s, despite the fact that
the process of emotion recognition from faces typically takes a fraction of this time. The current study required participants with
either high or low levels of non-clinical trait anxiety to make speeded emotional classification judgments to a series of facial
expressions drawn from seven emotional categories. Following previous work it was predicted that recognition of fearful facial
expressions would be more accurate in the high-trait anxious group compared with the low-trait anxious group. However, contrary
to this prediction, no anxiety-related differences in emotion perception were observed across all seven emotions. This suggests that
anxiety does not influence the perception of fear as has been previously proposed.
# 2007 Elsevier Ltd. All rights reserved.

Keywords: Anxiety; Emotion; Facial expressions; Recognition; Perception

1. Introduction production and recognition of emotional facial expres-


sions across different cultures.
The ability to interpret correctly the emotional facial Atypical processing of emotional facial expressions is
expressions of others is a skill fundamental to thought to be a feature of some neuropsychological and
successful human interaction. It has been suggested clinical disorders. For example, recognition of disgust is
that emotional facial expressions evolved as an external impaired in Huntington’s disease (Gray, Young, Barker,
representation of internal emotional states as a means Curtis, & Gibson, 1997). Furthermore, recognition of
for swift communication of these states between anger is thought to be impaired in patients with clinical
individuals (Blair, 2003). Darwin (1872) believed that depression (Mendlewicz, Linkowski, Bazelmans, &
facial expressions are an innate and universal compo- Philippot, 2005; although see Persad and Polivy (1993)
nent of non-verbal communication. In support of this who present evidence for a general processing deficit of
view, Ekman (1972) has shown comparable patterns of emotional facial expressions in depression). In contrast to
these processing deficits associated with some condi-
tions, recent evidence has emerged suggesting that
heightened (but non-clinical) levels of trait anxiety are
associated with an ability to more accurately recognize
* Corresponding author at: Department of Experimental Psychol-
ogy, 12a Priory Road, University of Bristol, Bristol, BS8 1TU, UK.
fearful facial expressions (Surcinelli, Codispoti,
Tel.: +44 117 928 8547; fax: +44 117 928 8588. Montebarocci, Rossi, & Baldaro, 2006) relative to lower
E-mail address: robbie.cooper@bris.ac.uk (R.M. Cooper). levels of trait anxiety. The current paper explores the

0887-6185/$ – see front matter # 2007 Elsevier Ltd. All rights reserved.
doi:10.1016/j.janxdis.2007.11.010
R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127 1121

relationship between recognition of emotional facial anxiety demonstrate attentional biases towards the
expressions and individual differences in trait anxiety. location of both angry (e.g., Bradley, Mogg, Falla, &
Early cognitive theories of emotional disorders (Beck, Hamilton, 1998) and fearful (Fox, 2002) expressions
1976; Bower, 1981) predict that anxiety should be relative to expressions that are happy or neutral.2
associated with biases favoring the processing of emotio- Furthermore, in an interpretation task, Richards et al.
nal stimuli across all domains of information processing (2002) presented ambiguous emotional facial expres-
(e.g., attention, memory, etc.). However, despite attempts sions that had been constructed by morphing together
to find such an information processing bias in memory two different emotional expressions (e.g., fear and
(e.g., Mogg, Mathews, & Weinman, 1987) there surprise). When these ambiguous expressions were
currently exists no consensus on the relationship between presented, high-anxious participants were more likely
memory for emotional stimuli and anxiety (Richards than low-anxious participants to identify them as
et al., 2002). Some studies have shown that high levels of fearful. Taken together, these studies demonstrate that
trait anxiety are associated with improved retrieval of at least some aspects of the processing of threat-related
threat-related information compared with neutral infor- emotional facial expressions (i.e., allocation of atten-
mation (e.g., Claeys, 1989; Eysenck & Byrne, 1994), tion; interpretation of ambiguous stimuli) are suscep-
while others have found no effects (e.g., Foa, McNally, & tible to individual differences in anxiety.
Murdock, 1989; Lang & Craske, 1997). Contrary to this, However, even in the study of attention there have
there is a large body of evidence suggesting anxiety is been mixed findings in terms of how anxiety influences
associated with a bias in attentional processing; a bias the way attentional resources are allocated to emotional
that favors the processing of threat-related information facial expressions. Rossignol, Phillippot, Douilliez,
(e.g., Eysenck, 1992; Mathews & Mackintosh, 1998; see Crommelinck, and Campanella (2005) gave partici-
Bar-Haim, Lamy, Pergamin, Bakermans-Kranenburg, & pants a visual oddball task where they had to detect a
van Ijzendoorn, 2007 for a recent meta-analysis). discrepant fearful or happy face amongst a series of
Furthermore, it has been argued that this bias in neutral faces. Participants who rated highly on self-
attentional processing towards threat stimuli is the most report measures of anxiety were quicker to identify
important cognitive factor in the etiology and main- discrepant faces than non-anxious participants. How-
tenance of anxiety (Mogg & Bradley, 1998). ever, fearful faces were detected just as quickly as
While such attentional bias is the most documented happy faces. Similarly, Fox et al. (2000) gave
of information processing biases in anxiety (Mathews, participants a visual search task in which they had to
Mackintosh, & Fulcher, 1997) it is not the only reported. search for either a positive or negative schematic face
Anxiety is also thought to influence the interpretation of amongst sets of distracter faces. They reported that
ambiguous stimuli. For example, when presented with negative faces were found more efficiently than positive
threat/neutral homophones and asked to make a faces but that this effect was not modulated by
classification judgment, high-anxious individuals are participants’ level of self-reported anxiety. Thus, some
more likely than low-anxious individuals to interpret the tasks demonstrate anxiety-related differences in the way
word as threat-related rather than neutral (Mathews, threat-related emotional facial expressions are pro-
Richards, & Eysenck, 1989). cessed while others do not, suggesting anxiety may only
Evidence for such anxiety-related biases in attention be influencing certain aspects of the way attention is
and interpretation are also displayed in relation to the allocated (Mathews & Mackintosh, 1998).
processing of emotional facial expressions. In the dot- More recently, and importantly for purposes of the
probe task for example,1 individuals with high levels of present study, research has examined how emotion
recognition (i.e., the ability to classify different
emotional facial expressions as belonging to different
1 discrete categories) might be influenced by anxiety.
In its typical form, the dot-probe task presents pairs of stimuli
(e.g., an angry face and a neutral face) for a brief period of time Surcinelli et al. (2006) gave a non-clinical sample of
(typically 500 ms). The allocation of spatial attention between the
stimuli is determined by response times to either identify or localize
the ‘probe’ which appears in the location previously occupied by one
2
of the stimuli. An attentional bias towards a stimulus category is Biases towards threat-related facial expressions have also been
assumed to be present if response times to probes occurring in the demonstrated in individuals not selected on the basis of anxiety level
location previously occupied by that stimulus type (e.g., angry faces) (e.g., Cooper & Langton, 2006) but it is the hyper-vigilance to these
are faster than when the probes appear in the location of the other stimuli that is thought to characterize anxiety (Mogg & Bradley,
stimulus type (e.g., neutral faces). 1998).
1122 R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127

high- and low-trait anxious participants an emotion was recorded in Surcinelli et al.’s (2006) study. Accuracy,
recognition task. Participants were presented with a as a measurement, gives only an absolute value for
series of images of emotional facial expressions, each whether or not the emotions on the faces were recognized
for 10 s. Once each image disappeared from the screen, correctly. It does not, for example, provide any
participants were required to select one of seven information as to whether elevated levels of anxiety
emotional labels (anger, sadness, happiness, fear, are associated with faster, more efficient, recognition of
surprise, disgust, and neutral) they thought best emotional expressions such as fear. There are many
described the face. Surcinelli et al. reported that studies of information processing in anxiety that show
recognition accuracy was equivalent across all emo- differences in choice reaction time as a function of
tional expressions with the exception of fear. Fearful anxiety level, but no such differences in task accuracy (for
faces were more accurately recognized by high-trait a summary see Eysenck, Derakshan, Santos, & Calvo,
anxious compared with low-trait anxious participants. 2007). Thus, reaction time to recognize the displayed
There are, however, two potential limitations to emotion might be a more sensitive measure to assess the
Surcinelli et al.’s (2006) study that question the specifics influence of anxiety on the processing of emotional facial
of their conclusion that trait anxiety is associated with an expressions compared with accuracy alone.
improvement in the ability to recognize fearful emotional The Surcinelli et al. (2006) study is, to our knowledge,
expressions. Firstly, although Surcinelli et al. assume the only one to date that has examined recognition of
they are measuring recognition of the emotional facial emotional facial expressions as a function of non-clinical
expressions in their experiment, the details of their trait anxiety.3 However, similar studies using word and
methodology are such that this assumption may not be picture stimuli have provided equivocal findings. Across
appropriate. The faces in their experiment were presented three experiments, and against their predictions, Math-
for 10 s before a response was allowed. Recognition rates ews and Milroy (1994) found no evidence that anxious
(i.e., the accuracy with which the emotional facial participants were any better at making a speeded valence
expressions are correctly identified) similar to those judgment concerning negative words (e.g., funeral,
reported by Surcinelli et al. can be achieved with stupid) than positive words (e.g., healthy, clever) when
presentation times of only 100 ms (Prkachin, 2003). compared with non-anxious participants. In contrast,
Thus, if the process of recognition takes only a fraction of Bradley and Lang (1999) required participants to perform
the time allowed to Surcinelli et al.’s participants, it may a very similar task to Mathews and Milroy, this time in
not be recognition per se that is being measured in their response to the display of pleasant (e.g., flowers) and
study. For example, deployment of visual attention to unpleasant (e.g., a snarling dog) images. They found
emotional facial expressions is thought to differ between anxious participants were faster at identifying the
high- and low-anxious individuals with high-anxious unpleasant images compared with non-anxious partici-
individuals being less able to disengage their attention pants but there was no difference in participants’ ability
from threat-related faces (Fox, Russo, Bowles, & Dutton, to identify the pleasant images.
2001). Across the 10-s presentation time in Surcinelli Given the equivocal nature of findings in this area with
et al.’s study, it could be that the high-anxious participants non-face stimuli, we look again at the relationship
spend more time attending to the fearful faces than the between anxiety and the recognition of emotional facial
low-anxious participants and this difference in attention expressions. The current study attempts to extend the
could account for the reported difference in recognition. work conducted by Surcinelli et al. (2006). A non-clinical
Furthermore, this relatively long presentation time of sample of high- and low-trait anxious participants were
10 s is a concern for reasons of ecological validity. In asked to perform a speeded emotional facial expression
evolutionary terms, a 10-s latency for recognition of the recognition task in which both response time and
emotional content of a conspecific’s facial expression accuracy were recorded. Following findings of Surcinelli
would render the system redundant in terms of signaling et al. it was predicted that participants with high self-
imminent danger. Thus, testing recognition rates reported levels of trait anxiety should be more accurate in
following a shorter exposure duration would provide a
more robust test of whether or not trait anxiety influences
the recognition of emotional facial expressions and not 3
some other cognitive process that might be influenced by Two other studies have shown that anxiety modulates the ERP
signal that arises from seeing emotional facial expressions (Rossignol,
anxiety in the 10 s that elapse before a response is made. Anselme, Vermeulen, Philippot, & Campanella, 2007; Rossignol
Secondly, and presumably because of the 10-s et al., 2005) but recognition of emotion was not directly measured
exposure duration, only accuracy, and not reaction time, in these studies.
R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127 1123

the recognition of fear compared to those with low levels term was assigned to a numeric key on the keyboard.
of trait anxiety. In addition, following the work of Half of the participants used one key assignment
Bradley and Lang (1999), response times to identify (1 = angry, 2 = disgust, 3 = fear, 4 = happy, 5 = neutral,
fearful emotional expressions should be faster in 6 = sad, 7 = surprise) while the other half used another
individuals with elevated levels of trait anxiety compared (1 = sad, 2 = surprise, 3 = neutral, 4 = fear, 5 = disgust,
with individuals with low levels of trait anxiety. 6 = angry, 7 = happy). These emotional terms and their
associated key assignments were displayed on screen
2. Method beneath the image throughout the experiment. Each
image remained on the screen until a response was
2.1. Participants detected or 4 s elapsed (whichever was sooner).
Experimental trials were preceded by six practice trials.
A total of 109 undergraduate students (58 female)
completed both the state and trait sections of the 3. Results
Spielberger State-Trait Anxiety Inventory (STAI;
Spielberger, Gorsuch, Lushene, Vagg, & Jacobs, Two participants were excluded due to accuracy
1983). Only participants who scored above the 75th being at, or below, chance level. Another participant
percentile or below the 25th percentile on the trait was excluded after boxplots identified their response
component of the STAI were selected, giving 54 times as outliers. This left 51 participants (26 high-
participants, 27 in each of the high-trait (M = 53.1, anxious). The two groups differed in trait-anxiety score,
S.D. = 5.0, 18 females) and low-trait (M = 29.3, t(49) = 21.0, p < .001, as well as state anxiety score,5
S.D. = 3.0, 9 females) anxiety groups. t(46) = 5.4, p < .001.

2.2. Materials
3.1. Accuracy6
Seventy images from the Ekman and Friesen (1976) Mean recognition accuracy scores are displayed in
pictures of facial affect series were selected. These Fig. 1. A 2 (trait anxiety group: high vs. low)  7
comprised seven images each from 10 individuals (5 (emotional facial expression) repeated measures analysis
female), each individual posing seven expressions of variance (ANOVA) revealed a main effect of emotion,
(anger, disgust, fear, happiness, neutral, sadness, and F(6,294) = 35.6, p < .001. Post-hot tests with Sidak
surprise). The images were cropped to show only the corrections revealed that happiness was recognized more
face (measuring 12.5 cm  16 cm) and were viewed accurately than all other emotional expressions (all
from approximately 60 cm. Image presentation was ps < .01), with the exception of surprise. Similarly,
controlled by E-prime software running on a Dell surprise was recognized more accurately than all other
Optiplex desktop PC and displayed on a 15-in. CRT expressions with the exception of happiness and neutral
monitor set to 85 Hz. (all ps < .01). In turn, neutral expressions were
recognized more accurately than all other expressions
with the exception of happiness and surprise (all
2.3. Procedure
ps < .01). Recognition accuracy for anger, disgust, fear,
and sadness did not differ significantly ( ps > .05).
Images were presented individually in a randomized
Importantly, the predicted trait anxiety level  emotional
order with each image displayed centrally on the screen.
facial expression interaction was not significant ( p > .6).
By means of a key-press, participants’ task was to quickly
and accurately identify which one of seven emotions was
being represented in the facial image.4 Each emotional 5
Three participants failed to complete the state component of the
STAI.
6
Recognition tasks such as this one can be subject to signal
4
The 70 images from the Ekman and Friesen (1976) set were detection analysis (e.g., Prkachin, 2003). This produces a new depen-
interspersed amongst 105 images from a new set of emotional facial dent variable (d0 or A0 ) which takes into account both accurate (hits)
expressions (Benton, Clark, Cooper, Penton-Voak, & Nikolov, 2007) and inaccurate (false alarms) responses, therefore giving a measure of
with the goal of comparing recognition accuracy across the two sets. sensitivity. To allow easier comparison with Surcinelli et al. (2006)
However, given that the Benton et al. set had not previously been this analysis is not presented here. However, consistent with the
validated, and to maintain similarity with Surcinelli et al. (2006), the analysis of the accuracy scores alone, A0 analysis did not reveal
data pertaining to these faces is not presented here. We return to this any anxiety-related modulation in the recognition of the various
issue in the discussion. emotional facial expressions, including fear.
1124 R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127

Fig. 1. Mean accuracy (%) to identify each emotional facial expression as a function of trait anxiety level (error bars represent 1 standard error of the
mean).

These results are in contrast with those of Surcinelli 3.2. Response time
et al. (2006) who found that participants who rated high
in trait anxiety were more accurate at recognizing The inter-participant means of median correct
fearful facial expressions compared with low-anxious response times are displayed in Fig. 2. A 2 (trait
participants. One possible explanation for the discre- anxiety group: high vs. low)  7 (emotional facial
pancy in the results of the present study and those of expression) mixed ANOVA revealed a main effect of
Surcinelli et al. is a difference in anxiety scores across emotion, F(6,294) = 32.9, p < .001. Post-hoc tests with
the two populations tested. In support of this Sidak corrections revealed that recognition of happy
interpretation, the high-anxious sample tested by and neutral faces was faster than all other expressions
Surcinelli et al. were, at least numerically, more (all ps < .007) but response time did not differ between
anxious than the current sample (mean = 57.1 and the two expressions ( p = .11). Latency to identify
53.1 respectively). In order to provide a further test of disgust, sadness, and surprise did not differ (all
the possibility that an increased ability to accurately ps < .15) and all three were recognized more quickly
recognize fearful facial expressions is only present in than both fear and anger (all ps < .003), the recognition
those participants who rate most highly in trait anxiety a time for which did not differ ( p > .11). Importantly, and
further analysis was conducted on 20 participants; those in keeping with the accuracy data, the predicted anxiety
with the 10 highest (M = 58.3, S.D. = 4.2) and lowest level  emotional facial expressions interaction for
(M = 26.0, S.D. = 1.6) trait anxiety scores. Recognition response time was not significant ( p > .5).
accuracy for fear faces was compared directly in these
two groups. This analysis revealed no evidence that 4. Discussion
high-trait anxious individuals recognize fearful facial
expressions more accurately than low-trait anxious The main goal of the present study was to investigate
individuals, t(18) = .65, p = .26 (one-tailed).7 how self-reported levels of trait anxiety influence the
ability to recognize emotional facial expressions,
particularly fear. It was predicted that participants with
high levels of trait anxiety would demonstrate greater
7
We also carried out the same analysis replacing trait anxiety scores
accuracy in recognizing the facial expression of fear
with state anxiety scores. Again, no anxiety-related differences were compared with participants with low levels of trait
observed and so these data are not reported here. anxiety. However, the main finding of the present study
R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127 1125

Fig. 2. Mean response time (ms) to identify each emotional facial expression as a function of trait anxiety level (error bars represent 1 standard error
of the mean).

is that participants’ trait anxiety level did not affect the the 10 participants who score most highly on trait
accuracy or speed with which fear, or any of the other anxiety with the lowest scoring 10 participants on
presented facial expressions (happy, sad, angry, a one-tailed test, there was no evidence to suggest
disgusted, surprised, and neutral), were recognized. recognition of fearful facial expressions is
This is despite overall recognition rates showing a very modulated by anxiety.
similar pattern to those observed by Surcinelli et al. (2) The current study also differs from Surcinelli
(2006). et al. (2006) in terms of the number of
The failure to observe any modulation of recognition participants tested and the number of stimuli
rates for emotional facial expressions as a function of presented to participants. However, the current
anxiety level contrasts sharply with the findings of study included greater numbers of each. There
Surcinelli et al. (2006) who reported that recognition were more participants (54 vs. 39) and more
accuracy of fearful facial expressions improves at exemplars of each emotional face stimulus (10
elevated levels of trait anxiety. There are, however, a vs. 7) than the Surcinelli et al. study. Increased
number of differences between the current study and numbers of participants and stimuli should lead
that of Surcinelli et al. that could account for this to increased power, and thus reduce the like-
discrepancy: We now discuss each in turn: lihood of a Type I error. Despite this, there was no
evidence to suggest recognition of fearful facial
(1) The trait anxiety level of the participants in expressions is modulated by anxiety.
Surcinelli et al.’s (2006) sample was marginally (3) One potential criticism of the current study is that
higher than that in the current study. This holds interspersed among the test images were 105
for both low-anxious (29.3 vs. 31.2) and high- other images of emotional facial expressions;
anxious (53.1 vs. 57.1) participants. Given that part of a set that were being validated. It could be
Surcinelli et al.’s results suggest that higher levels argued that the inclusion of these extra stimuli
of trait anxiety are associated with greater may have contributed to the lack of any anxiety-
recognition accuracy of fearful facial expressions related differences in the current data through,
it is possible that an increased level of trait perhaps, fatigue effects. However, while the
anxiety in the Surcinelli et al. sample might go current study did present more stimuli than those
some way to account for the discrepant findings. seen in Surcinelli et al.’s (2006) study, they were
However, even when the current study compared each displayed for approximately 2 s as opposed
1126 R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127

to 10 s in the Surcinelli et al. study meaning the Haim et al., 2007) the process of recognizing those
current experiment would have actually been expressions may not be similarly modulated by
shorter than that of Surcinelli et al. Furthermore, anxiety. This finding goes against early cognitive
experiments looking at the allocation of attention theories of emotional disorders (Beck, 1976; Bower,
in anxiety tend to have procedures that contain 1981) which predict that anxiety should be associated
hundreds of trials and these are sensitive to with biases favoring the processing of emotional
differences in anxiety (e.g., Bradley et al., 1998). stimuli across all domains of information processing.
Accordingly, while not being able to rule out this However, it is consistent with other evidence
potential criticism, it does seem unlikely that it suggesting processing domains such as memory are
can account for the difference in findings relatively unaffected by participants’ level of anxiety
between the two studies. (e.g., Lang and Craske, 1997). The data are also
(4) The biggest difference between the current study consistent with those obtained by Mathews and
and that of Surcinelli et al. (2006) (and the one we Milroy (1994) who found no anxiety-related differ-
believe best accounts for the difference in the ences in the classification of emotional words. We
findings of the two experiments) is the way in echo Mathews and Milroy’s conclusion that these data
which recognition was measured. Images in suggest that the processes required to make a
Surcinelli et al.’s experiment were presented for conscious assessment of a stimuli’s valence are
10 s with participants being asked to respond as distinct from those that lead to biases in attention
accurately as possible under no time pressure. In (e.g., Fox et al., 2001) and interpretation (e.g.,
contrast the images in the current study were Richards et al., 2002), and the former does not appear
presented until the participant responded (an to be not influenced by anxiety, at least not to the
average of 2059 ms across expressions) follow- same extent.
ing instructions to identify the faces as quickly However, the current results are somewhat incon-
and accurately as possible. Thus, while both the sistent with the results of Bradley and Lang (1999) who
current study and that of Surcinelli et al. claim to found that non-clinical high-anxious participants were
be measuring the recognition of emotional facial faster at making a valence decision to negative images
expressions, this may not be the case. The relative than were low-anxious participants. Unfortunately,
improvement in the ability of the high-anxious differences in methodology between the current study
group to accurately respond to the presence of and that of Bradley and Lang make it difficult to directly
fearful facial expressions in Surcinelli et al.’s compare these experiments.8
study may be due to the contribution of another What is clear is that this area would benefit from a
cognitive process that is influencing responses more detailed look at the influence of trait anxiety on the
later in the time course of stimulus presentation. recognition of emotional stimuli in general. The current
For example, in Surcinelli et al.’s study, it is experiment presents a clear challenge to the notion that
possible that the high-trait anxious group spent the recognition of emotional facial expressions is
longer attending to the fearful faces compared to modulated by elevated levels of trait anxiety. On the
the low-anxious group (c.f., Fox et al., 2001), and basis of the extant data overall, the evidence for a
it is this increased attention that might account general anxiety-related processing bias at the level of
for better performance on the subsequent stimulus recognition (be it word, picture, or face) is far
recognition task. A future experiment could from compelling.
directly test this idea by measuring eye move-
ments when performing Surcinelli et al.’s version Acknowledgement
of the task.
This work was funded by Grant RES 062-23-0052
The crucial point here is that the apparent from the ESRC.
improvement in recognition performance demon-
strated by the high-anxious group in Surcinelli
8
et al.’s (2006) experiment is not necessarily due to Bradley and Lang (1999) employed a two-alternative forced
anxiety directly influencing the recognition stage of choice task (i.e., decide whether the picture is positive or negative)
while the current study used a seven-alternative forced choice task
processing. Thus, while there is good evidence (i.e., choose one of seven possible emotions). It is possible that the
suggesting the allocation of attention to threat-related former task is more sensitive to picking up anxiety-related differences
facial expressions is modulated by anxiety (e.g., Bar- in emotional stimuli at shorter presentation times.
R.M. Cooper et al. / Journal of Anxiety Disorders 22 (2008) 1120–1127 1127

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