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E Learning e Education and Online Training 5th EAI International Conference eLEOT 2019 Kunming China August 18 19 2019 Proceedings Guanglu Sun
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Guanglu Sun
Jianhou Gan
Shuai Liu
Fei Lang
Zeguang Lu (Eds.)
299
e-Learning, e-Education,
and Online Training
5th EAI International Conference, eLEOT 2019
Kunming, China, August 18–19, 2019
Proceedings
123
Lecture Notes of the Institute
for Computer Sciences, Social Informatics
and Telecommunications Engineering 299
e-Learning, e-Education,
and Online Training
5th EAI International Conference, eLEOT 2019
Kunming, China, August 18–19, 2019
Proceedings
123
Editors
Guanglu Sun Jianhou Gan
Harbin University of Science Yunnan Normal University
and Technology Kunming, China
Harbin, China
Fei Lang
Shuai Liu Harbin University
Hunan Normal University Harbin, China
Hunan, China
Zeguang Lu
Institute of Data Science
National Academy of Guo Ding
Harbin, China
© ICST Institute for Computer Sciences, Social Informatics and Telecommunications Engineering 2019
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Preface
We are delighted to introduce the proceedings of the 5th edition of the 2019 European
Alliance for Innovation (EAI) International Conference on e-Learning, e-Education,
and Online Training (eLEOT 2019), held in the city of perpetual spring: Kunming,
China. This conference has brought together researchers, developers, and practitioners
from around the world who are leveraging and developing the research and applica-
tions of e-Learning, e-Education, and online training among the international scientific
community. The main purpose of this conference series is to address the most recent
and innovative trends in this broad area, ranging from distance education to collabo-
rative learning, from interactive learning environments to the modeling of STEM
(science, technology, mathematics, engineering) curricula. The program of this year’s
edition of eLEOT was especially interesting, as it covered the advancements in the ICT
sector, the most effective educational paradigms, the most prominent learning strate-
gies, and several other related topics, which were brought to the attention of our
community by engaged authors who sent their contributions from all around the world.
This year, the technical program of eLEOT 2019 consisted of 99 full papers,
including 5 invited papers in oral presentation sessions at the main conference tracks.
At least three Technical Program Committee (TPC) members were assigned for each
submitted paper during the rigorous, peer-review phase. At the end of the reviewing
process, 46 papers (acceptance rate: 47%) were accepted as they met the requirements
of scientific quality, novelty, and relevance set by the conference. The conference
tracks were: Track 1 – Blended Learning and Social Network; Track 2 – Best Practices
and Teaching/Educational Models; Track 3 – Assessment and Learning Analytics;
Track 4 – Massive Open Online Course; and Track 5 – New Technology Applied in
Education. Aside from the high-quality technical paper presentations, the technical
program also featured two keynote speeches. The two keynote speeches were Prof.
Zhijun Yang from Yunnan Institute of Educational Sciences, China, and Prof. Fei Lang
from Harbin University of Science and Technology, China.
The coordination with the Steering Committee chairs, Prof. Imrich Chlamtac and
Prof. Giovanni Vincenti, proved to be key to the success of this event, as well as the
precious contribution of the general chair, Prof. Guanglu Sun, the general co-chair,
Prof. Jian Liu, and the local chair, Prof. Tianwei Xu. We also wish to thank the
publications chairs, Prof. Fei Lang, Prof. Suxia Zhu, and Zeguang Lu, the TPC chairs,
Prof. Shuai Liu and Prof. Jianhou Gan, and all the other conference chairs for their
valuable support during all phases leading to the hosting of this conference. Special
thanks also to our EAI conference manager, Karolina Marcinova, and to our EAI
proceedings editor, Lucia Sedlarova, for their constant dedication and help during all
these months. We would also like to thank the city of Kunming for hosting the
conference and all the authors who attended the event for their participation. We hope
vi Preface
that the readers find several interesting research themes and insights into the topics in
this volume and that they are encouraged to participate in the future editions of eLEOT!
Guanglu Sun
Jian Liu
Shuai Liu
Jianhou Gan
Organization
Steering Committee
Imrich Chlamtac University of Trento, Italy
Organizing Committee
General Chair
Guanglu Sun Harbin University of Science and Technology
General Co-chair
Jian Liu Yunnan Normal University
Local Chairs
Tianwei Xu Yunnan Normal University
Juxiang Zhou Yunnan Normal University
Workshops Chair
Zhijun Yang Scientific Research Institute of Education
Publications Chairs
Fei Lang Harbin University of Science and Technology
Suxia Zhu Harbin University of Science and Technology
Zeguang Lu National Academy of Guo Ding Institute of Data
Science
viii Organization
Web Chair
Jing Qiu Harbin University of Science and Technology
Panels Chairs
Wenxing Hong Xia Men University
Shuren Li Chinese Academy of Sciences
Demos Chair
Lin Tang Yunnan Normal University
Tutorials Chair
Mingjing Tang Yunnan Normal University
1 Introduction
The revised edition of the College English Syllabus states: “The purpose of college
English teaching is to train students to have a strong reading ability and a certain ability
to listen, speak, read, write and translate, so that they can communicate in English.” [1]
Under the requirements, the reading comprehension in CET-4 (abbreviate for College
English Test Band 4) test always has a high rating. This not only puts higher
requirements on the reading ability of candidates, but also brings severe challenges to
college English teaching of reading. It can be seen that English reading plays a central
role in English learning and is the main goal of English learning. English reading is
also a necessary content for various examinations or daily applications.
Fragmented English learning in mobile context has become more and more
important. Whether in public or in school libraries, many college students choose
mobile devices for English learning. Fragmented learning is based on digital learning
and is effectively combined with mobile computing technology to enable students to
© ICST Institute for Computer Sciences, Social Informatics and Telecommunications Engineering 2019
Published by Springer Nature Switzerland AG 2019. All Rights Reserved
G. Sun et al. (Eds.): eLEOT 2019, LNICST 299, pp. 3–18, 2019.
https://doi.org/10.1007/978-3-030-35095-6_1
4 J. Zhang et al.
learn English at any time and any place. On the one hand, it can be a reasonable use of
fragmentation time, which is a trend of learning at present; on the other hand, many
English learning APPs have been designed some very good functions, such as
punching sign, which can promote learner to develop good study habits.
English reading resources are rich, diverse in form, and highly shared. While
bringing convenience to people, it also brings about the problem of information
overload. In the face of massive information, learner cannot quickly get the information
that is truly valuable to him. Reading is a complex cognitive process, but the existing
English reading APPs only do the plans based on learning objectives and learning
needs when resources are pushed, or push English reading resources to learner
according to his learning preferences, but not fully considering the change of the
learner in the dynamic learning process. So it is difficult to constantly adjust the
resources of the next push according to the change of the learner. When the APP-based
learning method is adopted by more and more people, the problem in learning
adaptability of learner in this environment is becoming more and more prominent. It is
difficult for learner to achieve real online self-learning, and it is more difficult to obtain
adaptive learning resources [2]. How to help student find the learning resources quickly
and accurately which are suitable for his needs in the process of mobile learning? So
there is personalized recommendation. It enables information consumers to get infor-
mation they need quickly, moreover it can also provide predicted information to users
in a targeted manner, and strive to push the information that the target users want most.
The e-commerce APP and the English word APP have made a good exploration in this
respect, but there are almost few English reading APPs for auto-adaptive pushing in
English reading resources. Therefore, how to combine fragmented English reading with
auto-adaptive push in the fragmented learning of mobile context, construct an auto-
adaptive resources push model based on Android, and develop an auto-adaptive
resources push APP with fragmentation features? It has become one of the research
topics in the mobilization of English reading.
According to the learner’s learning model in the learning process (English reading
ability, cognitive style, learning target, learning situation and learning result) and the
characteristics of English reading resource model (question type, theme and difficulty),
Jianmin, Min and Bin [3] used machine-learning algorithms, also known as ID3
algorithm to construct a learning recommendation model for learning resources in
fragmented reading. In order to test the feasibility and results of auto-adaptive
resources recommendation model in fragmented English reading, this paper uses the
principles of software engineering to determine the learning needs of target learners.
Through functional analysis, the relationship between the functional modules included
in the platform and the relationships between modules were determined. After that, a
database of fragmented English reading resources was built. Finally, using the C/S
architecture system, Android studio, Visual Studio, and SQL Server development tools,
the learning resource adaptive recommendation APP for fragmented English reading is
designed and implemented.
Design and Implementation of Adaptive Push APP Based on Android 5
Learner model
Cogni ve style
Resource model
*Reading ability
Ques on type
*Learning target
Difficulty
**Learning situa on
Theme
**Learning result
Update Resource
Learner portrait library
Learning scene data
Learn Reso
er urce
ID3 Push strategy
**Sequence of
knowledge points
to be learned
Data collec on
Note: (1) In the learning scene, the white circle represents learners, and the gray
circle represents the learning resources fragmented.
(2) “*” represents the relative stability of data. Items without “*” are more stable,
items with “*” take a longer time to update, and items with “**” update the fastest.
On the basis of literature review, it is found that in fragmented learning environ-
ment, learner models can be established according to learner’s reading ability, cogni-
tive style, learning target, learning situations, learning result and other information in
the four-level English reading. Then extract the feature information of English reading
Design and Implementation of Adaptive Push APP Based on Android 7
resources, and establish the connection between learner model and learning resources.
Thereafter, based on the feature information of the learner, the sequence of knowledge
points to be learned is generated. Based on this sequence, the push strategy is deter-
mined, the learning path is planned, and the learning resources are pushed.
The learner model shows that in order to realize the self-adaptive push of resources,
the system needs to understand 4 aspects, that are what level the learner is at, what
goals he want to achieve, where and how to learn, and how the effect is based on
resource push learning. Among them, the assessment of learner’s reading ability is
based on CET-4 syllabus. It is mainly based on the Chinese English Reading Ability
Rating Scale [10] and the seven factors mentioned in the article A Study on the
Structure and Development of College Students’ English Reading Ability [11]. And an
assessment scale for English reading ability is developed. Cognitive styles are mainly
field-dependent and field-independent. Students with different cognitive styles also
have different interests in reading, thinking patterns, learning strategies and the impact
of environment on them [9]. The vocabulary size and the accuracy of CET-4 are the
main learning target. Learning situation mainly considers the decibels of the noise level
of the current learner’s geographical location in mobile debris learning environment.
Learning result is mainly feedback through the correct rate of the problem of the learner
in the complex learning situation.
The question types, theme, and difficulty of reading materials have a greater impact
on learner. Different types of questions essentially examine different ways of thinking.
Based on different growth environment, learners have different familiarity with reading
materials on different subjects, and the results of the questions vary greatly. However,
subjects with different degrees of difficulty require learners to be at different levels of
learning. Understanding these differences in reading materials, the system can push
resources more specifically for learner. This study extracts the feature information of
English reading resources from the three-dimensional degree of topic type, subject
matter and difficulty level of reading resources, and establishes a three-dimensional
reading resource feature model. Question types are mainly considered from factual
details, inference problem, semantic comprehension problem, main idea and the atti-
tude of view problem. Theme is mainly considered from the humanities and social
sciences and natural sciences. The difficulty mainly lies in people’s experience of
careful reading of Level 4 to decide which is difficult or easy.
On this basis, by collecting learner characteristics and resource characteristics
information, the decision tree ID3 algorithm is used to establish the decision tree of
resource push, from which the rules of resource push can be found. ID3 will be used to
achieve the push of CET-4 careful reading resources in mobile fragmentation learning
situation and meet the individual learning needs of learner.
8 J. Zhang et al.
3 Systems Analysis
3.1 Requirement Analysis
Faced with the rapid expansion and rapid renewal of knowledge, people are always
eager to learn more knowledge to cope the rapidly changing times. Some developers
have caught people’s changes in this demand and developed a lot of learning APPs, but
most of them lack in theory of teaching design, and just concentrate a large amount of
resources. Seemingly very human, but it forced people to face with more multiple
choices when using these APPs. It making them even more confused and do not
knowing where to start. As a result, learning is like finding a needle in a haystack, and
nothing is achieved.
Therefore, in the fragmented learning situation, it is urgent to solve the problem of
auto-adaptive push of English reading resources. By collecting the learner’s charac-
teristics, resource characteristics, and the degree of influence of the environment
(volume) on the learner in the process of the using of the APP, we can determine the
learning demand based on the learning history data and adapt to push resources to help
learner get the information he need quickly when the learner log in again [12]. On the
other hand, the APP should also aim at the module of “Careful Reading” of CET-4,
with the guide of relevant theories of fragmented learning in mobile context, and
learner-centered, providing learner’s needs for autonomous learning, and provides the
possibility for learner to learn fragmented English reading in mobile environment.
Therefore, the APP should include five core modules: login registration, context
acquisition, learning resources recommendation, independent practice and learning
process.
In addition to functional requirements, the APP should also have non-functional
requirements. The APP should also be secure, simple to operate, and with appropriate
openness to meet the future needs of users.
4 System Design
4.1 System Architecture Design
The purpose of system construction is to better help decision makers to formulate
future recommendation strategies through effective data collection, so that the weak
links in learning can be targeted and intensively trained. The general architecture of the
adaptive push APP of English fragmented reading resources is shown in Fig. 2. The
general framework of the system is divided into five levels, including the base layer,
data layer, service layer, application layer and user layer.
1. Base layer
The base layer is the lowest layer of the system architecture, mainly based on the
campus network (local area network) to support the network service of the auto-
auto-adaptive learning resource recommendation system to meet the requirements
of network service of the system.
Design and Implementation of Adaptive Push APP Based on Android 11
2. Data layer
The data layer is the second layer of the system framework, which is mainly for
operating and managing system data, including resource data, management data and
behavioral data. The resource data includes English reading resources, cognitive
style test resources and English reading ability scale test resources. Management
data includes student personal information, software information, etc. Behavioral
data includes learner’s behavior data, self-designed learning plan and the semester
test, etc. The evaluation data includes exercise results. The data layer is mainly for
collecting, processing, exchanging and storing the above data.
3. Service layer
The service layer is the middle layer of the general architecture of the system, which
is used to connect the data layer and the service layer. The service layer is mainly
used to support the system to provide application services for users at the appli-
cation layer. The ASP.NET WebAPI framework is mainly used to quickly provide
API for HTTP clients to create Web services.
4. Application layer
The application layer presents an adaptive learning service that the auto-adaptive
learning resource recommendation system ultimately provides for learner. It mainly
embodies the functions of login and registration, learning resources recommenda-
tion, independent practice, situation acquisition, and learning record provided by the
system. Among them, login and registration are used for authentication and reading
ability, and cognitive style test. Learning resource recommendation is mainly used
for the system to recommend learning resources for learner. Autonomous exercises
are mainly used for learner to learn according to their own learning needs, selecting
the learning module and autonomous setting plans to learn. The situational acqui-
sition is mainly to obtain the environmental volume of the current user’s geo-
graphical location, thereby judging the influence of the annoyance of the
geographical volume on the learning. The learning record mainly records the cur-
rent learner’s basic information and learning behavior.
5. User layer
The user layer is the mainly layer which the system directly contacts the users, and
is the outermost layer of the system architecture. In the fragmented English reading,
most of the user of the learning resources auto-adaptive push system is learners who
are preparing for CET-4, and this system aims to provide learning services for them.
5 Key Technology
The framework is built by ASP.NET WebAPI of Visual Studio. SQL Server is used to
store the database. The server and the database are connected through the corresponding
interface. And the server is deployed on the computer through the specific IP address
and port number in the campus network by using the convenience of the campus
network. Communication principle: after the client sends a http request to the server, the
server parses the data, and then indirectly accesses the database and returns the data from
the server to the client. After the client obtains the data in the JSON format from the
server, the program will parse these data quickly. After the analysis is completed, the
results will be displayed to users through the interface of the mobile intelligent terminal,
as shown in Fig. 4.
cognitive style test and English reading ability test during the registration process. If he
is an old user, it will through the ID3 decision tree algorithm for comprehensive
recommendation based on the learner’s characteristics and learning behavior (cognitive
style, reading ability, learning goal, learning plan, and learning result) to find matching
questions in the database. Then encapsulate the relevant information of the title,
options, corresponding articles, answers, etc. into an object PractisePackage, and use
the JSON format to transfer to the front-end APP. The client uses the WebView control
to display the returned data, which is equivalent to a browser is embedded in the client
to load and display the web page. And the sliding answer effect answers of the selected
questions, options, related articles which parsed by client, is achieved by ViewPager
+fragment. The learning resources recommendation interface is shown in Fig. 5.
The ID3 algorithm first acquires learner’s characteristics, resources features and
classification properties. Then stores them in a two-dimensional array, and preprocesses
the discrete data to normalize it. According to ID3 algorithm, the information gain of
each attribute is calculated by the formula. Compare the information gain of these
attributes, and use the attribute with the largest information gain as the test attribute of
the S set, that is, the root node of the decision tree [14]. The flow chart of the algorithm
design is shown in Fig. 6.
Design and Implementation of Adaptive Push APP Based on Android 15
Fig. 148.—Aplustrum
aplustre L. Mauritius,
showing the partly
internal shell (S); F,
foot; LL, cephalic
lappets; TT, double
set of tentacles. (After
Quoy and Gaimard.)
Fig. 149.—Sigaretus
laevigatus Lam.,
showing shell partially
immersed in the foot;
F, anterior
prolongation of the
foot. (After Souleyet.)
(3) Internal; e.g. Philine, Gastropteron, Pleurobranchus, Aplysia,
Limax, Arion, Hyalimax, Parmacella, Lamellaria, Cryptochiton, and,
among bivalves, Chlamydoconcha.
(4) Absent; e.g. all Nudibranchiata and Aplacophora, many
Cephalopoda, a few land Mollusca, e.g. all Onchidiidae, Philomycus,
and Vaginula.
The Univalve Shell.—In univalve Mollusca the normal form of the
shell is an elongated cone twisted into a spiral form round an axis,
the spiral ascending to the left. Probably the original form of the shell
was a simple cone, which covered the vital parts like a tent. As these
parts tended to increase in size, their position on the dorsal side of
the animal caused them gradually to fall over, drawing the shell with
them. The result of these two forces combined, the increasing size of
the visceral hump, and its tendency to pull the shell over with it,
probably resulted in the conversion of the conical into the spiral shell,
which gradually came to envelop the whole animal. Where the
visceral hump, instead of increasing in size, became flattened, the
conical shape of the shell may have been modified into a simple
elliptical plate (e.g. Limax), the nucleus representing the apex of the
cone. In extreme cases even this plate dwindles to a few calcareous
granules, or disappears altogether (Arion, Vaginula).
Varieties of the Spiral.—Almost every conceivable modification
of the spiral occurs, from the type represented by Gena, Haliotis,
Sigaretus, and Lamellaria, in which the spire is practically confined to
the few apical whorls, with the body-whorl inordinately large in
proportion, to a multispiral form like Terebra, with about twenty
whorls, very gradually increasing in size.
Fig. 151.—Examples of
shells with
disconnected whorls;
A, Cyathopoma cornu
Mf., Philippines; B,
Cylindrella hystrix
Wright, Cuba. (Both ×
4.)
Fig. 152.—Example of a
shell whose apical
whorls alone are
coiled, and the
remainder produced in
a regular curve.
(Cyclosurus Mariei
Morel., Mayotte.)
In some cases the regularly spiral form is kept, but the whorls are
completely disconnected; e.g. some Scalaria, Spirula; among fossil
Cephalopoda, Gyroceras, Crioceras, and Ancyloceras; and, among
recent land Mollusca, Cylindrella hystrix and Cyathopoma cornu (Fig.
151). Sometimes only the last whorl becomes disconnected from the
others, as in Rhiostoma (see Fig. 180, p. 266), Teinostoma, and in
the fossil Ophidioceras and Macroscaphites. Sometimes, again, not
more than one or two whorls at the apex are spirally coiled, and the
rest of the shell is simply produced or coiled in an exceedingly
irregular manner, e.g. Cyclosurus, Lituites, Orygoceras, Siliquaria
(Fig. 153), Vermetus. In Coecum (Fig. 170, p. 260) the spiral part is
entirely lost, and the shell becomes simply a cylinder. In a few cases
the last whorl is coiled irregularly backwards, and is brought up to
the apex, so that the animal in crawling must carry the shell with the
spire downwards, as in Anostoma (Fig. 154), Opisthostoma (Fig.
208, p. 309), Strophostoma, and Hypselostoma (Fig. 202 A, p. 302).
Fig. 153.—Siliquaria anguina Lam.,
showing scalariform coil of upper
whorls and irregular extension of
the lower.
Fig. 156.—Fulgur
perversum L., Florida. ½.
Fig. 161.—Cornucopia-
shaped monstrosity of
Helix aspersa, from
Ilfracombe. (British
Museum.)
Composition of the Shell.—The shell is mainly composed of
pure carbonate of lime, with a very slight proportion of phosphate of
lime, and an organic base allied to chitin, known as conchiolin. The
proportion of carbonate of lime is known to vary from about 99 p.c. in
Strombus to about 89 p.c. in Turritella. Nearly 1 p.c. of phosphate of
lime has been obtained from the shell of Helix nemoralis, and nearly
2 p.c. from that of Ostrea virginica. The conchiolin forms a sort of
membranous framework for the shell; it soon disappears in dead
specimens, leaving the shell much more brittle than it was when
alive. Carbonate of magnesia has also been detected, to the extent
of ·12 p.c. in Telescopium and ·48 p.c. in Neptunea antiqua. A trace
of silica has also occasionally been found.
When the shell exhibits a crystalline formation, the carbonate of
lime may take the form either of calcite or aragonite. The calcite
crystals are rhombohedral, optically uniaxal, and cleave easily, while
the aragonite cleave badly, belong to the rhombic system, and are
harder and denser, and optically biaxal. Both classes of crystal may
occur in the same shell.
Two main views have been held with regard to the formation and
structure of the shell—(1) that of Bowerbank and Carpenter, that the
shell is an organic formation, growing by interstitial deposit, in the
same manner as the teeth and bones of the higher animals; (2) that
of Réaumur, Eisig, and most modern writers, that the shell is of the
nature of an excretion, deposited like a cuticle on the outside of the
skin, being formed simply of a number of calcareous particles held
together by a kind of ‘animal glue.’ Leydig’s view is that the shell of
the Monotocardia is a secretion of the epithelium, but that in the
Pulmonata it originates within the skin itself, and afterwards
becomes free.[335]
According to Carpenter, when a fragment of any recent shell is
decalcified by being placed in dilute acid, a definite animal basis
remains, often so fine as to be no more than a membranous film, but
sometimes consisting of an aggregation of ‘cells’ with perfectly
definite forms. He accordingly divides all shell structure into cellular
and membranous, according to the characteristics of the animal
basis. Cellular structure is comparatively rare; it occurs most notably
in Pinna, where the shell is composed of a vast multitude of tolerably
regular hexagonal prisms (Fig. 162 B). Membranous structure
comprises all forms of shell which do not present a cellular tissue.
Carpenter held that the membrane itself was at one time a
constituent part of the mantle of the mollusc, the carbonate of lime
being secreted in minute ‘cells’ on its surface, and afterwards
spreading over the subjacent membrane through the bursting of the
cells.
The iridescence of nacreous shells is due to a peculiar lineation of
their surface, which can be readily detected by a lens. According to
Brewster, the iridescence is due to the alternation of layers of
granular carbonate of lime and of a very thin organic membrane, the
layers very slightly undulating. Carpenter, on the other hand, holds
that it depends upon the disposition of a single membranous layer in
folds or plaits, which lie more or less obliquely to the general surface,
so that their edges show as lines. The nacreous type of shell occurs
largely among those Mollusca which, from other details in their
organisation, are known to represent very ancient forms (e.g.
Nucula, Avicula, Trigonia, Nautilus). It is also the least permanent,
and thus in some strata we find that only casts of the nacreous shells
remain, while those of different constitution are preserved entire.
Porcellanous shells (of which the great majority of Gasteropoda
are instances) usually consist of three layers, each of which is
composed of a number of adjacent plates, like cards on edge. The
inclination of the plates in the different layers varies, but that of the
plates in the inner and outer layer is frequently the same, thus if the
plates are transverse in the middle stratum, they are longitudinal in
the inner and outer strata, and, if longitudinal in the middle, they are
transverse in the other two. Not uncommonly (Fig. 163 B) other
layers occur. In bivalves the disposition and nature of the layers is
much more varied.
Fig. 162.—A, Section of shell of Unio: a, periostracal layer; b, prismatic layer; c,
nacreous layer. B, Horizontal section of shell of Pinna, showing the hexagonal
prisms.
Fig. 165.—Neritina
longispina Récl., Mauritius.
(Operculum removed.)
The various details of sculpture on the exterior surface of the
shell, the striae, ribs, nodules, imbrications, spines, and other forms
of ornamentation are all the product of similar and corresponding
irregularities in the mantle margin, and have all been originally
situated at the edge of the lip. Spines, e.g. those of Murex and
Pteroceras, are first formed as a hollow thorn, cleft down its lower
side, and are afterwards filled in with solid matter as the mantle edge
withdraws. What purpose is served by the extreme elaboration of
these spiny processes in some cases, can hardly be considered as
satisfactorily ascertained. Possibly they are a form of sculptural
development which is, in the main, protective, and secures to its
owners immunity from the attacks of predatory fishes.
‘Attached’ genera (e.g. Chama, Spondylus) when living on smooth
surfaces have a flat shell, but when affixed to coral and other uneven
surfaces they become very irregular in shape. The sculpture of the
base on which they rest is often reproduced in these ‘attached’
shells, not only on the lower, but also on the upper valve, the
growing edge of which rests on the uneven surface of the base.
Oysters attached to the branches of the mangrove frequently display
a central convex rib, modelled on the shape of the branch, from
which the plaits of sculpture radiate, while specimens fixed to the
smooth trunk have no such rib. Crepidula, a genus which is in the
habit of attaching itself to other shells, varies in sculpture according
to that of its host. Sometimes the fact may be detected that a
specimen has lived on a ribbed shell when young, and on a smooth
one when old, or vice versâ. A new genus was actually founded by
Brown for a Capulus which had acquired ribs through adhesion to a
Pecten. A specimen of Hinnites giganteus in the British Museum
must at one period of its growth have adhered to a surface on which
was a Serpula, the impression of which is plainly reproduced on the
upper valve of the Hinnites.[339]
Fig. 166.—A specimen of Anomia
ephippium L., Weymouth,
taken upon Pecten maximus,
the sculpture of which is
reproduced on the upper
valve of the Anomia, and
even on a young Anomia
attached to the larger
specimen.
Growth of the Shell.—Nothing very definite is known with regard
to the rate of growth of the shell in marine Mollusca. Under
favourable conditions, however, certain species are known to
increase very rapidly, especially if the food supply be abundant, and
if there is no inconvenient crowding of individuals. Petit de la
Saussaye mentions[340] the case of a ship which sailed from
Marseilles for the west coast of Africa, after being fitted with an
entirely new bottom. On arriving at its destination, the vessel spent
68 days in the Gambia River, and took 86 days on its homeward
voyage. On being cleaned immediately on its return to Marseilles, an
Avicula 78 mm. and an Ostrea 95 mm. long (both being species
peculiar to W. Africa) were taken from its keel. These specimens had
therefore attained this growth in at most 154 days, for at the period
of their first attachment they are known to be exceedingly minute. P.
Fischer relates[341] that in 1862 a buoy, newly cleaned and painted,
was placed in the basin at Arcachon. In less than a year after, it was
found to be covered with thousands of very large Mytilus edulis, 100