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Sheila L. Videbeck
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Unit 1
Current Theories and Practice
1. Foundations of Psychiatric–Mental Health Nursing
2. Neurobiologic Theories and Psychopharmacology
3. Psychosocial Theories and Therapy
4. Treatment Settings and Therapeutic Programs
Unit 2
Building the Nurse–Client Relationship
5. Therapeutic Relationships
6. Therapeutic Communication
7. Client’s Response to Illness
8. Assessment
Unit 3
Current Social and Emotional Concerns
9. Legal and Ethical Issues
10. Grief and Loss
11. Anger, Hostility, and Aggression
12. Abuse and Violence
Unit 4
Nursing Practice for Psychiatric Disorders
13. Trauma and Stressor-Related Disorders
14. Anxiety and Anxiety Disorders
15. Obsessive–Compulsive and Related Disorders
16. Schizophrenia
17. Mood Disorders and Suicide
18. Personality Disorders
19. Addiction
20. Eating Disorders
21. Somatic Symptom Illnesses
22. Neurodevelopmental Disorders
2
23. Disruptive Behavior Disorders
24. Cognitive Disorders
Appendix A
Disorders of Sleep and Wakefulness
Appendix B
Sexual Dysfunctions and Gender Dysphoria
Appendix C
Drug Classification Under the Controlled Substances Act
Appendix D
Canadian Drug Trade Names
Appendix E
Mexican Drug Trade Names
3
4
5
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7th edition
Copyright © 2017 Wolters Kluwer
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mentioned copyright. To request permission, please contact Wolters Kluwer at Two Commerce Square, 2001 Market Street, Philadelphia, PA
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987654321
Printed in China
eISBN: 9781496355911
Library of Congress Cataloging-in-Publication Data
available upon request
This work is provided “as is,” and the publisher disclaims any and all warranties, express or implied, including any warranties as to accuracy,
comprehensiveness, or currency of the content of this work.
This work is no substitute for individual patient assessment based upon health care professionals’ examination of each patient and consideration
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the patient. The publisher does not provide medical advice or guidance, and this work is merely a reference tool. Health care professionals, and
not the publisher, are solely responsible for the use of this work, including all medical judgments and for any resulting diagnosis and treatments.
Given the continuous, rapid advances in medical science and health information, independent professional verification of medical diagnoses,
indications, appropriate pharmaceutical selections and dosages, and treatment options should be made and health care professionals should
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any changes in dosage schedule or contraindications, particularly if the medication to be administered is new, infrequently used, or has a narrow
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6
Josephine M. Britanico, MSN, RN, PNP, PhD(c)
Assistant Professor of Nursing
Borough of Manhattan Community College/CUNY
New York, New York
7
Barbara J. Goldberg, MS, RN, CNS
Assistant Professor
Onondaga Community College
Syracuse, New York
Lois Harder, RN
Senior Lecturer
West Virginia University
Morgantown, West Virginia
8
The seventh edition of Psychiatric–Mental Health Nursing maintains a strong student focus, presenting sound
nursing theory, therapeutic modalities, and clinical applications across the treatment continuum. The chapters
are short, and the writing style is direct in order to facilitate reading comprehension and student learning.
This text uses the nursing process framework and emphasizes therapeutic communication with examples
and pharmacology throughout. Interventions focus on all aspects of client care, including communication,
client and family education, and community resources, as well as their practical application in various clinical
settings.
In this edition, all DSM-5 content has been updated, as well as the Best Practice boxes, to highlight current
evidence-based practice. New special features include Concept Mastery Alerts, which clarify important
concepts that are essential to students’ learning, and Watch and Learn icons that alert students to important
video content available on . Cultural and Elder Considerations have special headings to help call
attention to this important content. The nursing process sections have a new design to help highlight this
content as well.
Unit 2: Building the Nurse–Client Relationship presents the basic elements essential to the practice of mental
health nursing. Chapters on therapeutic relationships and therapeutic communication prepare students to
begin working with clients both in mental health settings and in all other areas of nursing practice. The
chapter on the client’s response to illness provides a framework for understanding the individual client. An
entire chapter is devoted to assessment, emphasizing its importance in nursing.
Unit 3: Current Social and Emotional Concerns covers topics that are not exclusive to mental health settings.
These include legal and ethical issues; anger, aggression, and hostility; abuse and violence; and grief and loss.
Nurses in all practice settings find themselves confronted with issues related to these topics. Additionally,
many legal and ethical concerns are interwoven with issues of violence and loss.
Unit 4: Nursing Practice for Psychiatric Disorders covers all the major categories of mental disorders. This
unit has been reorganized to reflect current concepts in mental disorders. New chapters include trauma and
9
stressor-related disorders; obsessive–compulsive disorder and related disorders; somatic symptom disorders;
disruptive disorders; and neurodevelopmental disorders. Each chapter provides current information on
etiology, onset and clinical course, treatment, and nursing care. The chapters are compatible for use with any
medical classification system for mental disorders.
PEDAGOGICAL FEATURES
Psychiatric–Mental Health Nursing incorporates several pedagogical features designed to facilitate student
learning:
SPECIAL FEATURES
• Clinical Vignettes, provided for each major disorder discussed in the text, “paint a picture” of a client
dealing with the disorder to enhance understanding.
• Nursing Care Plans demonstrate a sample plan of care for a client with a specific disorder.
• Drug Alerts highlight essential points about psychotropic drugs.
• Warning boxes are the FDA black box drug warnings for specific medications.
• Cultural Considerations sections highlight diversity in client care.
• Elder Considerations sections highlight the key considerations for a growing older adult population.
• Therapeutic dialogues give specific examples of the nurse–client interaction to promote therapeutic
communication skills.
• Client/Family Education boxes provide information that helps strengthen students’ roles as educators.
• Nursing Interventions provide a summary of key interventions for the specific disorder.
• DSM-5 Diagnostic Criteria boxes include specific diagnostic information for the disorder.
• Best Practices boxes highlight current evidence-based practice and future directions for research on a wide
variety of practice issues.
• Self-Awareness features encourage students to reflect on themselves, their emotions, and their attitudes as a
way to foster both personal and professional development.
• Concept Mastery Alerts clarify important concepts that are essential to students’ learning and practice.
• Watch and Learn icons alert the reader to important resources available on to enhance student
understanding of the topic.
10
ANCILLARY PACKAGE FOR THE SEVENTH EDITION
Instructor Resources
The Instructor Resources are available online at http://thepoint.lww.com/Videbeck7e for instructors who
adopt Psychiatric–Mental Health Nursing. Information and activities that will help you engage your students
throughout the semester include:
• PowerPoint Slides
• Image Bank
• Test Generator
• Pre-Lecture Quizzes
• Discussion Topics
• Written, Group, Clinical, and Web Assignments
• Guided Lecture Notes
• Case Studies
Student Resources
Students who purchase a new copy of Psychiatric–Mental Health Nursing gain access to the following learning
tools on using the access code in the front of their book:
• , highlighting films depicting individuals with mental health disorders, provide students the
opportunity to approach nursing care related to mental health and illness in a novel way.
• NCLEX-Style Review Questions help students review important concepts and practice for the NCLEX
examination.
• Journal Articles offer access to current research available in Wolters Kluwer journals.
• Online video series, Lippincott Theory to Practice Video Series includes videos of true-to-life clients
displaying mental health disorders, allowing students to gain experience and a deeper understanding of
these patients.
• Internet Resources provide relevant weblinks to further explore chapter content.
11
before moving him/her on to higher levels of learning. This will be a different experience for each student
based on the number of questions he/she answers and whether he/she answers them correctly. Each question
is also “normed” by all students in PrepU around the country—how every student answers a specific question
generates the difficulty level of each question in the system. This adaptive experience allows students to
practice at their own pace and study much more effectively.
Personalized Reports
Students get individual feedback about their performance, and instructors can track class statistics to gauge the
level of understanding. Both get a window into performance to help identify areas for remediation. Instructors
can access the average mastery level of the class, students’ strengths and weaknesses, and how often students
use PrepU. Students can see their own progress charts showing strengths and weaknesses—so they can
continue quizzing in areas where they are weaker.
Mobile Optimized
Students can study anytime, anywhere with PrepU, as it is mobile optimized. More convenience equals more
quizzing and more practice for students!
There is a PrepU resource available with this book! For more information, visit
http://thepoint.lww.com/PrepU.
This leading content is also incorporated into Lippincott CoursePoint, a dynamic learning solution that
integrates this book’s curriculum, adaptive learning tools, real-time data reporting, and the latest evidence-
based practice content into one powerful student learning solution. Lippincott CoursePoint improves the
nursing students’ critical thinking and clinical reasoning skills to prepare them for practice. Learn more at
www.NursingEducationSuccess.com/CoursePoint.
12
Many years of teaching and practice have shaped my teaching efforts and this textbook.
Students provide feedback and ask ever-changing questions that guide me to keep this text useful, easy to
read and understand, and focused on student learning. Students also help keep me up to date, so the text can
stay relevant to their needs. I continue to work with students in simulation lab experiences as nursing
education evolves with advances in technology.
I want to thank the people at Wolters Kluwer for their valuable assistance in making this textbook a reality.
Their contributions to its success are greatly appreciated. I thank Natasha McIntyre, Dan Reilly, Zach
Shapiro, Helen Kogut, and Cynthia Rudy for a job well done once again.
My friends continue to listen, support, and encourage my efforts in all endeavors. My brother and his
family provide love and support in this endeavor, as well as in the journey of life. I am truly fortunate and
grateful.
13
Unit 1
Current Theories and Practice
1. Foundations of Psychiatric–Mental Health Nursing
Mental Health and Mental Illness
Diagnostic and Statistical Manual of Mental Disorders
Historical Perspectives of the Treatment of Mental Illness
Mental Illness in the 21st Century
Cultural Considerations
Psychiatric Nursing Practice
2. Neurobiologic Theories and Psychopharmacology
The Nervous System and How it Works
Brain Imaging Techniques
Neurobiologic Causes of Mental Illness
The Nurse’s Role in Research and Education
Psychopharmacology
Cultural Considerations
3. Psychosocial Theories and Therapy
Psychosocial Theories
Cultural Considerations
Treatment Modalities
The Nurse and Psychosocial Interventions
4. Treatment Settings and Therapeutic Programs
Treatment Settings
Psychiatric Rehabilitation and Recovery
Special Populations of Clients with Mental Illness
Interdisciplinary Team
Psychosocial Nursing in Public Health and Home Care
Unit 2
Building the Nurse–Client Relationship
14
5. Therapeutic Relationships
Components of a Therapeutic Relationship
Types of Relationships
Establishing the Therapeutic Relationship
Avoiding Behaviors that Diminish the Therapeutic Relationship
Roles of the Nurse in a Therapeutic Relationship
6. Therapeutic Communication
What is Therapeutic Communication?
Verbal Communication Skills
Nonverbal Communication Skills
Understanding the Meaning of Communication
Understanding Context
Understanding Spirituality
Cultural Considerations
The Therapeutic Communication Session
Assertive Communication
Community-Based Care
7. Client’s Response to Illness
Individual Factors
Interpersonal Factors
Cultural Factors
8. Assessment
Factors Influencing Assessment
How to Conduct the Interview
Content of the Assessment
Assessment of Suicide or Harm Toward Others
Data Analysis
Unit 3
Current Social and Emotional Concerns
15
Disenfranchised Grief
Complicated Grieving
Application of the Nursing Process
Unit 4
Nursing Practice for Psychiatric Disorders
13. Trauma and Stressor-Related Disorders
Posttraumatic Stress Disorder
Etiology
Cultural Considerations
Treatment
Elder Considerations
Community-Based Care
Mental Health Promotion
Application of the Nursing Process
16
Onset and Clinical Course
Related Disorders
Etiology
Cultural Considerations
Treatment
Elder Considerations
Community-Based Care
Mental Health Promotion
Panic Disorder
Application of the Nursing Process: Panic Disorder
Phobias
Generalized Anxiety Disorder
16. Schizophrenia
Clinical Course
Related Disorders
Etiology
Cultural Considerations
Treatment
Application of the Nursing Process
Elder Considerations
Community-Based Care
Mental Health Promotion
17
Suicide
Elder Considerations
Community-Based Care
Mental Health Promotion
19. Addiction
Types of Substance Abuse
Onset and Clinical Course
Related Disorders
Etiology
Cultural Considerations
Types of Substances and Treatment
Treatment and Prognosis
Application of the Nursing Process
Elder Considerations
Community-Based Care
Mental Health Promotion
18
Substance Abuse in Health Professionals
19
Community-Based Care
Mental Health Promotion
Appendix A
Disorders of Sleep and Wakefulness
Appendix B
Sexual Dysfunctions and Gender Dysphoria
Appendix C
Drug Classification Under the Controlled Substances Act
Appendix D
Canadian Drug Trade Names
Appendix E
Mexican Drug Trade Names
20
Key Terms
• asylum
• boarding
• case management
• deinstitutionalization
• Diagnostic and Statistical Manual of Mental Disorders (DSM-5)
• managed care
• managed care organizations
• mental health
• mental illness
• phenomena of concern
• psychotropic drugs
• self-awareness
• standards of care
• utilization review firms
Learning Objectives
After reading this chapter, you should be able to:
2. Discuss the purpose and use of the American Psychiatric Association’s Diagnostic and Statistical Manual of
21
Mental Disorders (DSM-5).
5. Discuss the American Nurses Association (ANA) standards of practice for psychiatric–mental health
nursing.
AS YOU BEGIN THE STUDY OF psychiatric–mental health nursing, you may be excited, uncertain, and even
somewhat anxious. The field of mental health often seems a little unfamiliar or mysterious, making it hard to
imagine what the experience will be like or what nurses do in this area. This chapter addresses these concerns
and others by providing an overview of the history of mental illness, advances in treatment, current issues in
mental health, and the role of the psychiatric nurse.
Mental Health
The World Health Organization defines health as a state of complete physical, mental, and social wellness,
not merely the absence of disease or infirmity. This definition emphasizes health as a positive state of well-
being. People in a state of emotional, physical, and social well-being fulfill life responsibilities, function
effectively in daily life, and are satisfied with their interpersonal relationships and themselves.
No single universal definition of mental health exists. Generally, a person’s behavior can provide clues to his
or her mental health. Because each person can have a different view or interpretation of behavior (depending
on his or her values and beliefs), the determination of mental health may be difficult. In most cases, mental
health is a state of emotional, psychological, and social wellness evidenced by satisfying interpersonal
relationships, effective behavior and coping, positive self-concept, and emotional stability.
Mental health has many components, and a wide variety of factors influence it. These factors interact; thus,
a person’s mental health is a dynamic, or ever-changing, state. Factors influencing a person’s mental health
can be categorized as individual, interpersonal, and social/cultural. Individual, or personal, factors include a
person’s biologic makeup, autonomy and independence, self-esteem, capacity for growth, vitality, ability to
find meaning in life, emotional resilience or hardiness, sense of belonging, reality orientation, and coping or
stress management abilities. Interpersonal, or relationship, factors include effective communication, ability to
22
help others, intimacy, and a balance of separateness and connectedness. Social/cultural, or environmental,
factors include a sense of community, access to adequate resources, intolerance of violence, support of diversity
among people, mastery of the environment, and a positive, yet realistic, view of one’s world. Individual,
interpersonal, and social/cultural factors are discussed further in Chapter 7.
Mental Illness
Mental illness includes disorders that affect mood, behavior, and thinking, such as depression, schizophrenia,
anxiety disorders, and addictive disorders. Mental disorders often cause significant distress, impaired
functioning, or both. Individuals experience dissatisfaction with self, relationships, and ineffective coping.
Daily life can seem overwhelming or unbearable. Individuals may believe that their situation is hopeless.
Factors contributing to mental illness can also be viewed within individual, interpersonal, and social/cultural
categories. Individual factors include biologic makeup, intolerable or unrealistic worries or fears, inability to
distinguish reality from fantasy, intolerance of life’s uncertainties, a sense of disharmony in life, and a loss of
meaning in one’s life. Interpersonal factors include ineffective communication, excessive dependency on or
withdrawal from relationships, no sense of belonging, inadequate social support, and loss of emotional control.
Social/cultural factors include lack of resources, violence, homelessness, poverty, an unwarranted negative view
of the world, and discrimination such as stigma, racism, classism, ageism, and sexism.
• To provide a standardized nomenclature and language for all mental health professionals
• To present defining characteristics or symptoms that differentiate specific diagnoses
• To assist in identifying the underlying causes of disorders
The classification system allows the practitioner to identify all the factors that relate to a person’s condition:
• All major psychiatric disorders such as depression, schizophrenia, anxiety, and substance-related disorders
• Medical conditions that are potentially relevant to understanding or managing the person’s mental disorder
as well as medical conditions that might contribute to understanding the person
• Psychosocial and environmental problems that may affect the diagnosis, treatment, and prognosis of mental
disorders. Included are problems with the primary support group, the social environment, education,
occupation, housing, economics, access to health care, and the legal system.
Although student nurses do not use the DSM-5 to diagnose clients, they will find it a helpful resource to
23
understand the reason for the admission and to begin building knowledge about the nature of psychiatric
illnesses.
attempted to relate mental disorders to physical disorders and developed his theory that the amounts of blood,
water, and yellow and black bile in the body controlled the emotions. These four substances, or humors,
corresponded with happiness, calmness, anger, and sadness. Imbalances of the four humors were believed to
cause mental disorders; so treatment was aimed at restoring balance through bloodletting, starving, and
purging. Such “treatments” persisted well into the 19th century (Baly, 1982).
Possessed by demons
In early Christian times (1–1000 AD), primitive beliefs and superstitions were strong. All diseases were
again blamed on demons, and the mentally ill were viewed as possessed. Priests performed exorcisms to rid
evil spirits. When that failed, they used more severe and brutal measures, such as incarceration in dungeons,
flogging, and starving.
In England during the Renaissance (1300–1600), people with mental illness were distinguished from
criminals. Those considered harmless were allowed to wander the countryside or live in rural communities,
but the more “dangerous lunatics” were thrown in prison, chained, and starved (Rosenblatt, 1984). In 1547,
the Hospital of St. Mary of Bethlehem was officially declared a hospital for the insane, the first of its kind. By
24
1775, visitors at the institution were charged a fee for the privilege of viewing and ridiculing the inmates, who
were seen as animals, less than human (McMillan, 1997). During this same period in the colonies (later the
United States), the mentally ill were considered evil or possessed and were punished.Witch hunts were
conducted, and offenders were burned at the stake.
Development of Psychopharmacology
A great leap in the treatment of mental illness began in about 1950 with the development of psychotropic
drugs, or drugs used to treat mental illness. Chlorpromazine (Thorazine), an antipsychotic drug, and lithium,
an antimanic agent, were the first drugs to be developed. Over the following 10 years, monoamine oxidase
inhibitor antidepressants; haloperidol (Haldol), an antipsychotic; tricyclic antidepressants; and antianxiety
agents, called benzodiazepines, were introduced. For the first time, drugs actually reduced agitation, psychotic
thinking, and depression. Hospital stays were shortened, and many people became well enough to go home.
The level of noise, chaos, and violence greatly diminished in the hospital setting.
25
1963 with the enactment of the Community Mental Health Centers Construction Act.
Deinstitutionalization, a deliberate shift from institutional care in state hospitals to community facilities,
began. Community mental health centers served smaller geographic catchment, or service, areas that provided
less restrictive treatment located closer to individuals’ homes, families, and friends. These centers provided
emergency care, inpatient care, outpatient services, partial hospitalization, screening services, and education.
Thus, deinstitutionalization accomplished the release of individuals from long-term stays in state institutions,
the decrease in admissions to hospitals, and the development of community-based services as an alternative to
hospital care.
In addition to deinstitutionalization, federal legislation was passed to provide an income for disabled
persons: Supplemental Security Income (SSI) and Social Security Disability Income (SSDI). This allowed
people with severe and persistent mental illness to be more independent financially and to not rely on family
for money. States were able to spend less money on care of the mentally ill than they had spent when these
individuals were in state hospitals because this program was federally funded. Also, commitment laws changed
in the early 1970s, making it more difficult to commit people for mental health treatment against their will.
This further decreased the state hospital populations and, consequently, the money that states spent on them.
26
Revolving door
Shorter, unplanned hospital stays further complicate frequent, repeated hospital admissions. People with
severe and persistent mental illness may show signs of improvement in a few days but are not stabilized. Thus,
they are discharged into the community without being able to cope with community living. However,
planned/scheduled short hospital stays do not contribute to the revolving door phenomenon, and may show
promise in dealing with this issue (see Chapter 4). The result frequently is decompensation and
rehospitalization. In addition, many people have a dual problem of both severe mental illness and substance
abuse. Use of alcohol and drugs exacerbates symptoms of mental illness, again making rehospitalization more
likely. Substance abuse issues cannot be dealt with in the 3 to 5 days typical for admissions in the current
managed care environment.
Homelessness is a major problem in the United States today with 610,000 people, including 140,000
children, being homeless on any given night. Approximately 257,300 of the homeless population (42%) have a
severe mental illness of a chronic substance use disorder. The segment of the homeless population considered
to be chronically homeless numbers 110,000 and 30% of this group has a psychiatric illness and two thirds
have a primary substance abuse disorder or other chronic health condition (Substance Abuse and Mental
Health Services Administration, 2015). Those who are homeless and mentally ill are found in parks, airport
and bus terminals, alleys and stairwells, jails, and other public places. Some use shelters, halfway houses, or
board-and-care rooms; others rent cheap hotel rooms when they can afford it. Homelessness worsens
psychiatric problems for many people with mental illness who end up on the streets, contributing to a vicious
cycle.
Many of the problems of the homeless mentally ill, as well as of those who pass through the revolving door
of psychiatric care, stem from the lack of adequate community resources. Money saved by states when state
hospitals were closed has not been transferred to community programs and support. Inpatient psychiatric
treatment still accounts for most of the spending for mental health in the United States, so community mental
health has never been given the financial base it needs to be effective. In addition, mental health services
provided in the community must be individualized, available, and culturally relevant to be effective.
27
BOX 1.1 HEALTHY PEOPLE 2020 MENTAL HEALTH OBJECTIVES
U.S. Department of Health and Human Services. (2010). Healthy People 2020. Washington, DC: DHHS.
Community-Based Care
After deinstitutionalization, the 2000 community mental health centers that were supposed to be built by
1980 had not materialized. By 1990, only 1300 programs provided various types of psychosocial rehabilitation
services. Persons with severe and persistent mental illness were either ignored or underserved by community
mental health centers. This meant that many people needing services were, and still are, in the general
population with their needs unmet. The Treatment Advocacy Center (2015) reports that about one half of all
persons with severe mental illness have received no treatment of any kind in the previous 12 months. Persons
with minor or mild cases are more likely to receive treatment, whereas those with severe and persistent mental
illness are least likely to be treated.
Community support service programs were developed to meet the needs of persons with mental illness
outside the walls of an institution. These programs focus on rehabilitation, vocational needs, education, and
socialization as well as on management of symptoms and medication. These services are funded by states (or
counties) and some private agencies. Therefore, the availability and quality of services vary among different
areas of the country. For example, rural areas may have limited funds to provide mental health services and
28
smaller numbers of people needing them. Large metropolitan areas, although having larger budgets, also have
thousands of people in need of service; rarely is there enough money to provide all the services needed by the
population. Chapter 4 provides a detailed discussion of community-based programs.
The community-based system did not accurately anticipate the extent of the needs of people with severe
and persistent mental illness. Many clients do not have the skills needed to live independently in the
community, and teaching these skills is often time-consuming and labor intensive, requiring a 1:1 staff-to-
client ratio. In addition, the nature of some mental illnesses makes learning these skills more difficult. For
example, a client who is hallucinating or “hearing voices” can have difficulty listening to or comprehending
instructions. Other clients experience drastic shifts in mood, being unable to get out of bed one day and then
unable to concentrate or pay attention a few days later.
Despite the flaws in the system, community-based programs have positive aspects that make them
preferable for treating many people with mental illnesses. Clients can remain in their communities, maintain
contact with family and friends, and enjoy personal freedom that is not possible in an institution. People in
institutions often lose motivation and hope as well as functional daily living skills, such as shopping and
cooking. Therefore, treatment in the community is a trend that will continue.
29
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Otoconcha is peculiar. The fresh-water Mollusca, besides the Isidora
characteristic of the sub-region, are partly related to New Caledonia
through the occurrence of Melanopsis, partly to Tasmania through
Potamopyrgus, while the peculiar Latia is possibly akin to
Gundlachia (Tasmania). The land operculates number only 5 genera
and 14 species in all, excluding a doubtful Diplommatina.[374]
Land and Fresh-water Mollusca of the Neozealanian Province
Schizoglossa 1
Paryphanta 5
Rhytida 6
Rhenea 2
Helicarion 1
Otoconcha 1
Microcystis 1
Trochonanina 1
Phacussa 3
Thalassohelix 5
Gerontia 2
Allodiscus 10
Pyrrha 1
Therasia 7
Phenacohelix 3
Suteria 1
Flammulina 13
Laoma 23
Endodonta 10
Charopa 28
Placostylus 1
Carthaea 1
Tornatellina 1
Janella 3
Latia 2
Ancylus 2
Limnaea 5
Amphipeplea 2
Planorbis 1
Isidora 7
Melanopsis 2
Potamopyrgus 4
Paxillus 1
Lagochilus 7
Omphalotropis 1
Realia 4
Hydrocena 1
Unio 9
Sphaerium 1
Pisidium 2
Lord Howe’s I. is remarkable as containing a Placostylus, which
thus links the island with this province. The remainder of the fauna is
Polynesian, with the exception of a species (common to the Fijis) of
Parmella, a slug akin to Helicarion, Parmacochlea, and Cystopelta.
(3) The Polynesian Sub-region includes all the island groups of
the central and southern Pacific (except those classified in the
Papuan and Australian sub-regions), from the Pelews and Carolines
in the west to the Marquesas and Paumotus in the east, and from
the Tonga group in the south to the Sandwich Is. in the north. It may
be subdivided into (a) the Polynesian province proper, and (b) the
Hawaiian province, which includes the Sandwich Is. only.
Fig. 216.—Characteristic
Polynesian Mollusca: A,
Achatinella vulpina Fér.,
Sandwich Is.; B, Partula
planilabrum Pease, Society
Is.
(a) The general features of the Polynesian province are very
similar throughout, although the Mollusca of each island group are in
the main peculiar. The species are mostly small and obscure. Helix
scarcely occurs, its place being taken by small Zonitidae
(Microcystis, Charopa, Trochomorpha, etc.), and by groups of so-
called Patula (Endodonta, Pitys, etc.), the exact position of which is
not yet settled. Libera, remarkable for its method of ovipositing (p.
128), is peculiar to the Society and Hervey Is.; Partula is almost
universal, attaining its maximum (40 sp.) in the Society Is.;
Tornatellina, Pupa, and Vertigo occur throughout.
The land operculates consist chiefly of Omphalotropis, Pupina,
Realia, and Helicina. Diplommatina and Palaina are abundant on the
Pelews, and a Moussonia occurs in the Samoa Is. Ostodes, a small
form of Cyclophorus, is found in some of the southern groups. The
fresh-water operculates are Melania, Neritina (including Clithon, a
sub-genus furnished with spines), and Navicella; there are no
Unionidae, while fresh-water Pulmonata are very scarce.
(b) The land Mollusca of the Hawaiian province are distinguished
by the possession of four entirely peculiar genera—Achatinella,
Leptachatina, Carelia, and Auriculella. More than 300 of the two
former genera have been described, every mountain valley of some
of the islands having its own peculiar species. The destruction of the
indigenous herbage by goats is rapidly extinguishing many forms.
Partula, and the small land operculates, so characteristic of the other
groups, are, with the exception of Helicina, entirely wanting. The
occurrence of one of the Merope group of Helix (Solomon Is.) is
remarkable, and there is a rich development of Succinea. “Patula,”
Microcystis, Tornatellina, and the other small Polynesian land
Pulmonata are well represented. The presence of Isidora, absent
from the central Pacific groups, is remarkable, and Erinna is a
peculiar genus belonging to the Limnaeidae.
CHAPTER XI
GEOGRAPHICAL DISTRIBUTION OF LAND MOLLUSCA (continued)—THE
ETHIOPIAN, NEARCTIC, AND NEOTROPICAL REGIONS
Fig. 220.—Mollusca
characteristic of L.
Tanganyika: A, Nassopsis
nassa Woodw.; B, Spekia
zonata Woodw.; C,
Syrnolopsis lacustris E. A.
Smith.
Fig. 221.—Achatina zebra
Lam., S. Africa. × ½.
(2) The South African Sub-region.—The principal characteristic
of the Mollusca of S. Africa is the occurrence of numerous small
species of Helicidae, belonging chiefly to the groups Pella, Phasis,
Dorcasia, and Sculptaria, all of which are practically peculiar.
Carnivorous genera are also prominent, Ennea here attaining its
maximum. Rhytida (to which several species still regarded as Pella
belong) is common only to the S. Pacific and Australasia, and forms,
with Isidora among the fresh-water pulmonates, a remarkable link of
connexion. Aerope, the largest of all helicoid carnivorous genera,
and Chlamydephorus, a carnivorous slug with an internal shell, are
peculiar. Achatina is still abundant, but Limicolaria is wanting.
Livinhacea, a form with a continuous peristome, perhaps akin to
Bulimus; Apera, a form of slug; and Coeliaxis, a genus perhaps akin
to the Papuan and Queensland Perrieria, are all peculiar. The land
operculates, which are not numerous, are of the East African type.
Land Mollusca of the S. African Sub-region
Chlamydephorus 1
Ennea 31
Aerope 5
Rhytida 3
Helicarion 3
Trochonanina 1
Trochozonites 1
Limax 1
Apera 1
Vitrina 7
Nanina 6
Conulus 2
Patula 2
Pella 44
Dorcasia 8
Phasis 1
Sculptaria 2
Helix (inc. sed.) 4
Rachis 1
Pachnodus 3
Buliminus (?) 4
Pupa 20
Vertigo 2
Achatina 18
Livinhacea 1
Stenogyra 4
Coeliaxis 1
Succinea 3
Vaginula 2
Cyclophorus 1
Cyclostoma 7
Cyclotus (?) 1
Blanfordia 1
St. Helena.—The Molluscan fauna of St. Helena is perhaps the
most puzzling, as regards its geographical affinities, of any in the
world. It consists of 29 peculiar species of land Mollusca (fresh-water
species being unknown), 19 of which are recently extinct, partly
owing to the destruction of the forest, but are found in considerable
abundance in a state of good preservation.[375] The genera are—
Hyalinia 1
Patula 4 (3 extinct)
Endodonta 10 (7 extinct)
Bulimulus 7 (5 extinct)
Pachyotus 1 (extinct)
Tomigerus (?) 1 (extinct)
Pupa 2 (extinct)
Succinea 3
The 5 genera which concentrate our attention are Patula,
Endodonta, Pachyotus (Fig. 222), Tomigerus, and Bulimulus, all of
which appear utterly strange to an oceanic island in the middle of the
S. Atlantic. Patula and Endodonta are essentially Polynesian forms,
occurring abundantly on all the island groups in the Central Pacific.
Pachyotus, Tomigerus (assuming its correct identification), and
Bulimulus are all S. American forms, the two former being especially
characteristic of Brazil. How this mixture of genera now confined to
regions so widely distant, not only from St. Helena itself, but from
one another, became associated here, is a problem obviously not
easy of solution. The fauna is probably a remnant of a very ancient
type, possibly at one time much more widely distributed. Endodonta
(an essentially insular form, like Omphalotropis) actually occurs on
Fernando Noronha, off the Brazil coast, and we shall see how an
Indian and even a Polynesian element is present off the eastern
coasts of Africa.
Ascension I.—One indigenous species, a so-called Limax, is all
that has ever been discovered.
(3) The Malagasy Sub-region includes Madagascar with its
attendant satellites Bourbon, Mauritius, and Rodriguez, and the
Seychelles and Comoro groups. No land Mollusca are known from
the Amirantes, the Chagos, or from Aldabra. The special
characteristics of the sub-region are the great development of the
carnivorous land Mollusca (Ennea, Gibbus), the occurrence of a
considerable number of true Helicidae of great size and beauty, and
the prominence of the genus Cyclostoma.
(a) The Madagascan Province.—The land Mollusca of
Madagascar, although as yet imperfectly known, possess a striking
individuality. Two of the chief characteristics of the Ethiopian region
are the paucity of its land operculate and of its Helix fauna;
Madagascar is especially distinguished by the rich development of
both these groups. For size, colouring, and beauty of shape, the
Helicidae of the two sub-genera Ampelita and Helicophanta rival, if
they do not surpass, any in the world. They are quite peculiar to this
sub-region, not a trace of them occurring on the Mascarenes,
Seychelles, or even on the Comoros. Helicophanta is distinguished
by the enormous size of its embryonic shell, which persists in the
adult (Fig. 223), and in this respect the group appears to be related
to Acavus (Ceylon, Fig. 204) and Panda (N.E. Australia). As is usual
when Helix is well developed, Nanina (about 12 sp.) is
proportionately scanty.
The African Bulimini (Pachnodus and Rachis) are represented by
two species, but Achatina, so abundant on the mainland, is scarce.
Two other groups of Buliminus, Leucotaenia and Clavator, are
peculiar. The presence of a single Kaliella, specifically identical with
a common Indian form, is very remarkable.
Cyclostoma proper, of which Madagascar is the metropolis, is
richly developed (54 sp.). Many of the species are of great size and
of striking beauty of ornamentation. Unlike its Helicidae, this genus is
not restricted to Madagascar; several species occur on the mainland,
6 on the Comoros, one on the Seychelles, and 16 in Mauritius. The
sub-genera Acroptychia and Hainesia are peculiar.
Fig. 224.—Cyclostoma
campanulatum Pfr., Madagascar.
The fresh-water Mollusca of Madagascar contain further traces of
Indian relationship. Thus we find two species of Paludomus, a genus
whose metropolis is Ceylon, India, and Further India, and which is
barely represented on the Seychelles and in the Somali district.
Melanatria, which is peculiar to Madagascar, has its nearest affinities
in the Cingalese and East Indian faunas. Several of the Melania and
the two Bithynia are of a type entirely wanting in Africa, but common
in the Indo-Malay sub-region. Not a single one of the characteristic
African fresh-water bivalves (Mutela, Spatha, Aetheria, Galatea, etc.)
has been found in Madagascar. On the other hand, certain African
Gasteropoda, such as Cleopatra and Isidora, occur, indicating, in
common with the land Mollusca, that an ultimate land connexion with
Africa must have taken place, but at an immeasurably remote period.
Land and Fresh-water Mollusca of Madagascar
Ennea 9
Urocyclus 2
Helicarion (?) 1
Macrocyclis (?) 1
Kaliella 1
Nanina (inc. sed.) 9
Ampelita 35
Helicophanta 17
Pachnodus 2
Rachis 2
Leucotaenia 2
Clavator 2
Achatina 3
Opeas 2
Subulina 3
Vaginula 4
Limnea 2
Planorbis 3
Isidora 3
Melania 7
Melanatria 4
Paludomus 2
Vivipara 1
Bithynia 2
Cleopatra 2
Ampullaria 6
Cyclophorus 2
Cyclotus (?) 1
Cyclostoma 54
Otopoma 5
Lithidion 1
Acroptychia 3
Hainesia 3
Unio 1
Corbicula 2
Sphaerium 1
Pisidium 1
The Comoro Islands.—This isolated group possesses about 100
species, almost all of which are peculiar. The principal feature is the
rich development of Ennea (30 sp.). On the whole the group shows
more relationship to Madagascar than to the mainland. Thus we
have six species of true Cyclostoma, and only one Achatina, while
among the fresh-water genera is Septaria, which is characteristic of
the whole Malagasy Sub-region, but is absent from the mainland.
The Helicidae are all of insignificant size. Peculiar to the group is the
remarkable genus Cyclosurus (Fig. 152, p. 247).
(b) The Mascarene Province (Mauritius, Bourbon, Rodriguez, and
the Seychelles).—The percentage of peculiar species, which is very
high, can only be paralleled in the case of some of the West Indian
islands, and sufficiently attests the extreme isolation of the group
from Madagascar. We have—
Total sp. Land sp. Fresh-water Peculiar Peculiar to
sp. group
Mauritius 113 104 9 78 102 (90 p.c.)
Bourbon 45 40 5 19 38 (84 p.c.)
Rodriguez 23 19 4 15 21 (95 p.c.)
Seychelles 34 27 7 24 30 (90 p.c.)
The Mollusca of the group exhibit three distinct elements, the
Indigenous, the Madagascan, and the Indian and Australasian.
The genus Pachystyla (Naninidae) is quite peculiar, forming the
main portion of the land snails proper. It attains its maximum in
Mauritius (17 sp.), with 5 sp. in Bourbon and one sub-fossil sp. in
Rodriguez, while in the Seychelles it is absent. But the principal
feature of the Mascarene group is the extraordinary development of
the carnivorous genus Gibbus, which has 27 sp. in Mauritius, 8 in
Bourbon, 4 in Rodriguez; in the Seychelles, it is replaced by
Edentulina and Streptostele. The principal link with Madagascar is
found in a part of the operculate land fauna. Cyclostoma is present
(with Otopoma) in several fine living forms, and the number of sub-
fossil species is a clear indication that this group was, not long ago,
much more abundant, for of the 16 Cyclostoma known from
Mauritius 10 are sub-fossil. The operculates form a decided feature
of the land fauna; thus in Mauritius there are 32 species, or more
than 28 per cent of the whole.