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All protists, fungi, plants, and animals are eukaryotes. Some of these organisms
are independent, free-living cells (Figure 3.1); others consist of many cells working together as a body.
By defi nition, a eukaryotic cell starts out life with a nucleus (eu– means
true; karyon means nut, or kernel). Like many other organelles, a nucleus has a
substances that cross it. Such control maintains a special internal environment
that allows the organelle to carry out its particular function. That function may
be isolating toxic or sensitive substances from the rest of the cell, transporting
substances through cytoplasm, maintaining fl uid balance, or providing a favorable environment for a
special process.
cells also have other special structures (Figures 3.9 and 3.10). Much as interactions among organs keep
an animal body alive and well, interactions among
The Nucleus A nucleus serves two important functions. First, it keeps the
cell’s genetic material—its one and only copy of DNA—safe and sound. Isolated
in its own compartment, DNA stays separated from the bustling activity of the
The second function of a nucleus is to control the passage of certain molecules between the nucleus and
the cytoplasm . The nuclear membrane, which is
called the nuclear envelope, carries out this function. A nuclear envelope consists of two lipid bilayers
folded together as a single membrane. Receptors and
transporters stud both sides of the bilayer; other proteins cluster to form tiny
pores that span it. These molecules and structures work as a system to selectively
differ in structure and metabolic details. Some characteristics of archaeans indicate they are more
closely related to eukaryotic cells than they are to bacteria.
Most prokaryotic cells are not much bigger than a few micrometers. None
has a complex internal framework, but protein fi laments under the plasma
membrane reinforce the cell’s shape. Such fi laments also act as scaffolding for
internal structures.
Figure 3.8 shows a general body plan of a prokaryotic cell. The cytoplasm of
these cells contains many ribosomes (organelles upon which polypeptides are
assembled), and in some species, additional organelles. The cell’s single chromosome, a circular DNA
molecule, is located in the cytoplasm, in an irregularly
shaped region called the nucleoid. Most nucleoids are not enclosed by a membrane. The cytoplasm of
many prokaryotes also contains plasmids. These small
circles of DNA carry a few genes (units of inheritance) that can provide advantages, such as resistance to
antibiotics.
Many pr