Professional Documents
Culture Documents
R.M.Vijayakumar, Ph.D,
Professor and Head
J.Auxcilia, Ph.D,
Asst. Professor (Hort)
M.Kavino, Ph.D,
Asst. Professor (Hort)
D.Durga Devi,
Professor (Crop Physiology)
2014
All rights reserved, no part of this publication may be reproduced, stored in a retrieval
system or transmitted in any form or by any means, electronic, mechanical, photocopying,
recording or otherwise without the prior written permission of the publisher.
ISBN: 972-81-73298-00-3
Published by
Department of Fruit Crops
Horticultural College and Research Institute
Tamil Nadu Agricultural University
Coimbatore – 641 003
E-mail: fruits@ tnau.ac.in
Correct citation: Vijayakumar, R.M., Auxcilia, J., Kavino, M., and Durga Devi, D. 2014. A
Knowledge Guide on Fruit Science, Published by Department of Fruit Crops, HC&RI, TNAU,
Coimbatore- 641 003
TAMIL NADU AGRICULTURAL UNIVERSITY
Dr.K.Ramasamy
Vice-Chancellor
FOREWORD
In India, fruit culture has been practiced from ancient times. The wealth and health of any
country can be judged by the percapita consumption of fruits by its people. India is bestowed with
a wide range of agro climatic and soil conditions. Therefore, almost all types of fruits can be grown
in one or the other part of the country. India is the second largest producer of fruits next to China.
According to the latest data, in India, fruits are produced in an area of 6.98 million hectares with a
production of 81.28 million tonnes. But the productivity is only 11.6 t/ha. Since a great demand for
fruits exists in the domestic as well as foreign markets, we have to go a long way in meeting the
demand. Moreover, fruit production has been recognized as a remunerative and sustainable
enterprise in the country. It has a great potential for generating employment both in on and off
farm sectors and therefore, scientific knowledge on fruit cultivation is essential to achieve higher
productivity. Hence, the research programmes on fruit crops has to be still strengthened in order
to achieve higher productivity in many fruit crops.
One of the ways to achieve higher productivity to meet the growing demand in future is that
the students have to be enriched with knowledge on fruit science. This will help the students to take
up innovative projects on improvement and management of fruit crops, which will reflect in the
market in due course.
This comprehensive knowledge on fruit science along with allied sciences will help them to
carry out research on applied science by way of interpretation of results in an appropriate manner.
Hence, the book entitled, “Knowledge Guide on Fruit Science” will definitely help the students as
well as scientists to acquire and refresh their knowledge on Fruit Crops.
I appreciate the efforts taken by the scientists for preparation and release of this book for
the betterment of knowledge of the scientific community.
K. Ramasamy
PREFACE
Fruits had been a staple food for mankind from the time immemorial. The ancient man’s food
habit was mainly based on fruits and tubers which were found naturally in the forest. Before the era
of civilization, the ancient people including saints who lived in forest maintained their livelihood by
consuming fruits only. As per bible, the history of human life started when Adam and Eve
consumed an Apple fruit. Hence, Horticulture, particularly the “Fruit culture” can be considered as
the primitive agriculture. In the past history also, establishment of many mango orchards and
vineyards by the Moghul kings and Nawabs during 16th to 18th century all over the country was
recorded.
India is blessed with a wealthy diversity of fruits owing to the diverse agro-climatic regions.
Right from tropical fruits like mango, banana, papaya, citrus and jack to temperate fruits such as
apple, plum, peach etc. are being grown in various agro-climatic zones of India. Since fruit industry
is one of the remunerative ventures, studying fruit science and research on fruit crops has gained
momentum in the recent days, due to the growing demand from the people mainly on health
grounds
Hence, the students who are studying fruit science have to be imparted with knowledge on
fruit crops besides the basic aspects of plant botany, plant breeding and plant physiology which is
lacking among the students. Hence, the main purpose of this book is to compile information
pertaining to fruit crops including the basic aspects of allied disciplines like botanical, plant
breeding and physiological terminologies which will be much useful in understanding the applied
aspects of crop improvement and crop management programmes.
Taking this in mind, this compilation was made and the book entitled as “Knowledge Guide
on Fruit Science”. We hope this book will be useful for UG, PG and Ph.D students studying
horticulture particularly fruit science to enrich their knowledge in all aspects of fruit science.
Authors
ACKNOWLEDGEMENT
The authors wish to place their sincere and heartfelt thanks for the overwhelming help rendered by the
following staff and students of Department of Fruit Crops in preparing this book in a comprehensive manner
Staff:
Dr.K.Soorianthasundaram, Professor (Hort)
Dr.C.Kavitha, Asst. Professor (Hort)
Dr.Vetrivelkalai, Asst. Professor (Nematology)
Dr.G. Thiribhuvanamala, Asst. Professor (Pathology)
R.Vaishnavi, SRF
South
wales
Abortive : Imperfectly formed or rudimentary
Achene : A small dry one seeded indehiscent fruit
Actinomorphic : Regular and the flower which can be divided into two symmetrical halves by radial
planes
Adventitious : The root which are developed from vegetative parts other than radical
Aestivation : Arrangement of sepals and petals in a flower bud
Aggregate fruit : Cluster of fruits developed from the ovary of a single flower e.g: Polyalthia,
Strawberry, Annona
Albuminous : If the endosperm is not completely consumed the embryo remains comparatively
small and a large quantity of stored food remains within the endosperm which the
embryo could not consume, but would consume in the germinating stage. Such a
seed is termed as albuminous
Alternate : The leaves one after another singly at the nodes
Androecium : Stamens collectively/male part of the flower
Annular : Forming a ring or circle
Anthesis : Opening of flowers
Axile placentation : This is a type in which the placentae are on the axis of the ovary. Seeds are
attached to the axis
Axillary : Arising in the axil of a leaf or shoot
Basifixed : Filament attached to the base of the stem
Berry/ Bacca : Simple, fleshy fruit. The pericarp is fleshy which includes one or more carpels and
seeds e.g guava and tomato
Biseriately : Arranged in two rows
Bracts : The leafy structure beneath the flower/group of flowers
Calyx : Sepals considered as a whorl
Carpel : One of the units composing the gynoecium/pistil and containing one or more
ovules
Cauliflorous : Stem flowering / flowers are borne on trunks or heavy branches.
Ciliate : Fringed with marginal hairs
Colona : Appendage of corolla / by a transverse splitting of the corolla, an additional whorl
may be formed at its throat.
Companulate : Bell shaped flower
Corm : A round swollen, fleshy, underground stem in which the food is stored. eg) Yam,
banana
Corolla : The petals considered as a whorl.
Corymb : The inflorescence axis in this type is not elongated as in raceme. The pedicels of
the flowers are of unequal length. The older flowers have long pedicels and the
younger flowers show short pedicels. Hence all flowers appear at the same level
Cotyledon : A food digesting and food storing part of an embryo.
Cyme Inflorescence in which the main axis terminates in a flower with growth continued
by the side branches. The flower at the centre first opens and matures. eg.
Jasmine, Papaya
Deciduous : Trees which shed / loose or shed their leaves during autumn season or in the fall.
Decurrent (leaf) : Leaves have the base prolonged down the stem often in the form of projections.
Distichous (leaf) : Arranged in two opposite vertical ranks.
Dorsifixed(anther) : Filament is attached to the back of the anther.
Drupe : One seeded fleshy fruit in which the endocarp is stony, mesocarp is fleshy and
exocarp is skin – like.
Egg Female gamete
Embryo : A rudimentary plant with in the seed that usually develops from a fertilized egg
Endocarp : Inner wall of the pericarp.
Endosperm : A reserved food in seeds, which develops from the fertilization of the polar nuclei
of the ovule by the second male nucleus and nourishes embryo
Epicarp or : Outermost layer of the pericarp.
Exocarp
Epicotyl : The growing point of the embryo which gives rise to the shoot or above ground
part of the plant
Epipetalous : Stamens adhere to the corolla wholly or partially by their filaments and the anthers
remain free.
Exalbuminous : If the embryo completely fills up the ovule consuming both endosperm and
nucellus, the seed is exalbuminous, there being no endosperm. In such a seed
food for the embryo is kept stored in the cotyledons.
Extrose anthers : Opening towards outside of the flower
Filament : Stalk of the stamen bearing the anther at its tip
Fruit : Ripened ovary; one or more mature ovaries containing seeds and sometimes
adjacent attached parts
Gamopetalous : Petals united partially or wholly
Gamosepalous : Sepals united partially or wholly
Gynoecium : Female part of the flower including ovary / ovaries, style(s), stigma
Gynophore : Stalk of the ovary
Habit : General form of growth of a plant.
Herb : A plant which does not develop much woody tissue and thus remains soft and
succulent.
Hermaphrodite : Perfect flower; both stamens and pistils are present.
Hesperidium : A berry with a tough, leathery rind.
Imbricate : Petals or sepals are overlapping like the tiles of a roof
Inferior ovary : Epigynous; flower parts above the ovary or apparently growing from its tip
Inflorescence : A group of flowers on a common axis
Introse anthers : Opening inward and shed their pollen towards inside of the flower.
Lateral : Flowers/fruit borne at the side
Mesocarp : The middle layer of the fruit which is mostly fleshy.
Mixed spadix : The banana spadix is a mixed inflorescence as cymose groups of flowers are
arranged acropetally on the flesy axis. Each cymose group is subtended by a
spathe- an older spathe subtending the next younger. This spadix bends down by
its own weight
Monocarpic : A shoot flowers only once and dries after it has borne fruits.
Monocotyledon : A plant whose embryo has one cotyledon
Multiple fruit : A compound fruit formed by all the flowers of a whole inflorescence.
Nerve : Veins or ribs in bracts, leaves, sepals, petals
Node : Slightly enlarged portion of the stem where the leaves and buds arise at the
junction of two internodes.
Nucellus : The tissue in the central part of the ovule in which the embryo sac is embedded
Obtuse : Blunt or rounded at the tip.
Opposite : two leaves or buds at a node
Orbicular : Approximately circular in outline
Ovary : Enlarged basal portion of a pistil which forms the fruit
Ovate : Egg shaped in outline, broader at the base
Ovoid : Egg shaped
Ovule : Structure containing an embryo sac which becomes the seed after fertilization
Palmate : leaf segments or leaflets radiate from a point like the fingers from the palm of the
hand
Panicle : Compound raceme; type of inflorescence, the main axis of inflorescence is
branched and the flowers are borne not directly on the axis but on its branches in
a racemose pattern
Parietal : Types in which the ovules are attached to the wall of the ovary eg) cucumber,
placentation papaya
Pedicel : Stalk of the individual flowers of an inflorescence
Peduncle : Stalk/stem of main axis of inflorescence
Pentamerous : Having the parts in five or multiples of five
flower
Perennial : Persisting for more than two years
Perianth : Petals and sepals taken together
Pericarp : The ovary wall consisting of three layers exocarp, mesocarp and endocarp
Petalloid : Petal like stamens
Petaloid bract : Bracts sometimes become brightly coloured imitate petals and perform the same
function. These may easily be mistaken for petals
Pinnately veined : Leaf blade with single midrib from which smaller veins branch off like the divisions
of a feather
Pistillode : Abortive or sterile or rudimentary gynoecium
Plumule :A rudimentary structure in the embryo that develops into shoot after germination
Polyadelphous :
Stamens separate or in more than two groups
Polygamous :
A plant develops three kinds of flowers, staminate, pistillate and bisexual
Protandrous :
Stamens mature before the stigma is receptive
Protogynous :
Stigma is receptive before the stamens dehisce
Pubescent :
Covered with fine, soft hairs
Racemose :
The main axis of the inflorescence (rachis) grows indefinitely giving rise to
younger and younger flower in an acropetal order. The rachis never terminates in
a flower the apex being left free for further growth e.g) Mango
Rachis : Main axis of a spike
Receptacle : Enlarged end of pedicel, peduncle to which other flower parts are attached
Reticulate : Forming a network
Serrate : With sharp teeth that point upward/forward
Sessile : Not stalked
Sheath : A tubular envelope above the stem e.g banana
Shrub : A short, much branched woody plant
Spadix : This is a variation in spike in which the rachis is thick and fleshy and the flowers
are covered by one or more spathy bracts.
Spathe : Large bract or pair of bracts subtending a spadix or flower cluster
Spathy bract : These are large, thick, boat-shaped bracts called spathes covering the whole or a
part of a spadix inflorescence e.g) banana
Spike : An indeterminate type of inflorescence like raceme in which numerous flowers
which are sessile and borne on a rachis e.g) coconut
Spine : Sharply pointed structure below or above a leaf axil
Staminate : Flowers bearing stamens only
Staminode : Abortive or sterile stamens/rudimentary stamens
Stipule : An outgrowth at the base of the leaf
Superior ovary : Hypogynous; ovary appearing above other parts of the flower
Sympodial : Formed by several axis;
Taproot : A well developed vertical main root bearing/producing lateral roots
Tendril : Tender, coiled organ used in climbing
Tepal : Individual segments of perianth
Tomentose : Covered with dense, wool like hair
Truncate : Appearing to be cut off squarely at the tip
Tubercles : Small projections in any part of the plant
Unisexual : One sex only
Versatile anthers : Anthers attached near the middle of the filament in such a manner that they swing
freely
Zygomorphic : Irregular, one or more of the parts of a series are dissimilar
TERMINOLOGIES RELATED TO POLLINATION MECHANISMS/FLOWERS/FRUITS
Andromonoecious : Male and bisexual flowers borne on the same tree e.g) Papaya
Dichogamy : The maturation of the male and female reproductive organs at different
periods to ensure cross-pollination.
Dioecious : Male and female flowers borne on different plants e.g) Papaya
Fascicle : A special type of cymose corymb flower. In a polychasial inflorescence the
successive axes are of equal lengths bringing the flowers to the same level
in a corymbose manner. The flowers are very much clustered together. This
is seen in some members of Garcinia of Guttiferae
Gynodioecious : Female plants and andromonoecious plants will be segregated in 1:2 ratio
e.g) Papaya
Heterostyly : The phenomenon in higher plants in which the style in the flower is of a
different length in different individuals. Generally the stamens also are
attached at different levels in the flower, or are of different lengths. This
mechanisms favours cross-fertilization between individuals differing in this
kind of flower structure.
Hypanthodium : This is the typical inflorescence of figs. Here the cup-shaped receptacle is
formed by the condensation of the rachis of three closely placed cymes. The
receptacle is concave and cup shaped. The upper end has an opening called
ostiole, which is protected by scales. Inside the receptacle three types of
flowers are present. Male flowers are present in the upper part, female
flowers towards the base and the neutral flowers are found in the middle
between the male and female flowers
Malformation/Stamen : In gynodioecious papayas when temperature goes below 20 o C or above
carpellody/Cat-faced 35oC, the stamens adhere to the ovarian wall and the resultant fruit is
malformed or mis-shapened.
Monoecious : Male and female flowers borne on the same plants.g) Mango
Nomenclature : Naming of plants. Normally the scientific name/botanical name is in Latin
language, because this language is concise or precise in expression
Parthenogenesis : Development of fruit without pollination and fertilization but still produces
viable pollen
Parthernocarpy : The production of fruit without fertilization and formation of normal seeds
e.g., bananas. This may be caused by 1. Lack of pollination with or without
stimulation, 2. Lack of fertilization (by incompatibility or gametic sterility) or 3.
Lack of embryo development.
Phyllotaxy : Mode of arrangement of leaves on the stem
Sex-reversel/Sexual : Ability of a plant to change its flower type. Change of sex from one phase to
ambivalence the other. This occurs in papaya due to environmental changes or any stress
situations. Male tree produces bisexual or female flowers and produces fruit
Stenospermocarpy : Pollination and fertilization takes place but the embryo gets aborted and
results in seedless fruit e.g Thompson seedless variety of Grapes
Stimulative : Development of fruits is due to the stimulus of the pollen, only pollination and
parthenocarpy no fertilization e.g Black Corinth & Papaya
Taxonomy : It is the branch of botany which deals with study and description of the
variations in plants. The term systematic is commonly synonym.
Thyrsus : In this type a number of cymose clusters are borne acropetally on an
unlimited axis in the manner of a raceme. This is seen in grapevine.
Vegetative : Fruits develop without pollination/fertilization or even without stimulus
parthenocarpy
DESCRIPTION OF TYPES OF FRUITS
Amphisarica : This is a superior many-chambered and many seeded fruit with a woody
pericarp. The inner layers of the pericarp become pulpy and edible. The
testa is mucilaginous. Eg). Aegle marmelos and Feronia limonia of
Rutaceae.
Balusta : This is the special inferior, many-celled and many seeded fruit. The pericarp
is tough and two rows of carpels are placed one above the other. Seeds are
attached irregularly and the calyx persists on the top. The succulent testa is
edible. Eg). Pomegranate
Berry/Bacca : This term is usually applied to any fleshy fruit in which there is no hard part
excepting the seeds. One or the other of epicarp, mesocarp and endocarp
may, sometimes be differentiated or they may form a pulp in which the seeds
are embedded. Eg) Papaya, banana and guava
Drupe : This is normally a one-seeded fruit with a skinny epicarp, a fibrous and
usually juicy mesocarp and a stony hard endocarp. Drupes are called
―Stone fruits‖ because of the stony endocarp.
Hesperidium : This is superior and many-chambered, developing from syncarpous ovary
with axile placentation. The endocarp projects inwards forming distinct and
sometimes separable chambers while the epicarp and mesocarp are fused
together form the skin. The juicy, unicellular hairs are outgrowths from the
endocarp. Eg). Oranges and lemons
Pepo : This is an inferior one-celled or spuriously three celled, many-seeded fruit as
seen in the Cucurbitaceae. The epicarp forms a tough rind and the seeds are
firmly attached to the placenta. This is an inferior one-celled or spuriously
three celled, many-seeded fruit and epicarp forms a tough rind and the
seeds are firmly attached to the placenta. Eg) Cucurbitaceae
Pome : This is an inferior fruit developed out of a syncarpous ovary. The fruit is false
as it is surrounded by a fleshy thalamus which is edible. Eg).Apple (Pyrus
malus) and pear (Pyrus communis).
SPECIES NAME AND ITS SIGNIFICANCE
Albinism: Absence of chlorophyll. This is due to genetic causes and is in contrast to chlorosis or etiolation
caused by environmental factors
Allele: Alleles are alternative forms of the gene and are located at the same point (locus) in homologous
chromosomes
Allogamy: Syn: Cross-fertilization: Pollen grains from one plant pollinate the flowers of other plants
Amphidiploids: An allopolyploid. The result of hybridization between two plant species or genera where
the chromosome set in each parent undergoes doubling and produces a hybrid which has the sum of the
diploid number of the two parent forms as chromosome number. It has two copies of each of the two or
more different genomes present. Thus an amphidiploids have the diploid set of chromosome derived from
each parent
Aneuploid: An aneuploid organism has a chromosome number that is not an exact multiple of the basic
chromosome number/ haploid number. This is an unbalanced polyploidy. Hyperploid=Higher;
Hypoploid=lower.
Anther culture: Culture of anthers or pollen grains on a suitable medium for production of callus and or
haploid plants.
Antibodies: Specific reaction products produced in a living organism due to the presence of antigens. The
antibodies are specific and only act against those antigens that have evoked formation of the antibodies.
Antibodies are of considerable importance for immunity reactions.
Antigens: Substances that stimulate the production of antibodies when introduced into an organism.
Apogamy: Development of embryo from synergids or antipodal cells without fertilization; this is a form of
apomixes
Apomixis: The development of an individual from an unfertilized egg without sexual fusion, whether the
egg be normally haploid or abnormally diploid through failure of reduction division.
Apospory: A form of apomixes in which the embryo sac develops from a vegetative cell of the ovule
Auto tetraploid: A tetraploid whose nuclei contain four sets of chromosomes of the same origin
(duplicational ployploid).
Auto triploid: An individual having three homology sets of chromosomes in the body cell.
Autogamy: Pollen grains of a flower pollinate the same flower; Spontaneous self-fertilization
Autopolyploid: A polyploidy that has more than two copies of the same genome
Auxotroph: A mutant that is unable to grow on a minimal nutrient medium, nut that will develop normally if
a certain substance is added.
Backcross breeding: Breeding method based on repeated back-crossing of the F1 and the subsequent
generations to the recurrent parent
Biotype: A population of individuals with identical genetic constitution. A biotype may be homozygous or
heterozygous.
Bud selection: A form of clonal selection in which mutant buds are selected
Bud-mutation/Bud sport: A mutation in the bud. Such a mutated bud may grow into a branch or flower
differing in one or more characters from the parent plant. Somatic mutation.
Centres of Diversity: Areas where cultivated plant species and or their wild relatives show much greater
variation than anywhere in the rest of the world
Centres of origin: Areas where cultivated plant species are supposed to have originated.
Certified seeds: Seeds produced from foundation or registered seeds. Its purity is certified by a seed
certification agency, and is usually used for commercial crop production
Chimera: A plant composed of tissues of two or more genetically distinct types, as a result of mutation,
irregular mitosis, plastid segregation or artificial fusion by grafting.
Chromosome: Thread like structures present in nucleus which show distinct change in their morphology
during cell division. They are deeply stained with some dyes, particularly with the Feulgen‘s reagent. Genes
are located in chromosomes in a linear order like beads on a string
Cleistogamy: Pollination and fertilization without the opening of flowers; the flowers do not open at all i.e)
there is no anthesis
Combining ability: The relative ability of a biotype to transmit desirable performance to its crosses.
General combining ability: The average or over-all performance of a genetic strain in a series of crosses:
Specific combining ability: The performance of specific combinations of genetic strains in crosses in
relation to the average performance of all combinations.
Correlation: The relationship of one attribute to another attribute of an individual or population. The co-
efficient of correlation is a measure of the amount of the relationship either positive or negative ranging ±
1.0. If it is zero, then there is no correlation.
Cross pollination: The transfer of pollen from the anther of one flower to the stigma of another.
Distant hybridization: Hybridization between individuals belonging to two different species of the same
genera or between two genera
Double cross: A hybrid between two single crosses involving four different inbred lines.
Emasculation: Removal of stamens from the bisexual flower to avoid genetic contamination
Embryo culture: A method of inducing growth of embryos artificially. The technique is of value in
obtaining progeny in inter-specific crosses in which partially developed embryos normally abort. It consist
of excising embryos under aseptic conditions and placing them on suitable nutrient media for further
growth.
Euploid: An organism with chromosome number in exact multiple of the haploid number of the species
from which it arose. It may be diploid, triploid, tetraploid etc.
Evolution: A process that has occurred during previous geological periods and by which the living
organisms, broadly speaking, have evolved form more primitive ancestors to higher and more complicated
types of plants and animals.
Fertilization: The fusions of the nuclei form male and female gametes
Geitonogamy: Self-pollination in which the pollen is derived from another flower on the same plant
Genetic drift: A change in the frequency of a gene/allele in a population due to random sampling.
Genetic incompatibility: Incompatibility reaction depends directly on the relation existing between the
genetic constitution of the female gamete and the genetic constitution of the male gamete.
Heterosis: It indicates the hybrid vigour. The performance of offspring are better than parents.
Heterozygote: A zygote formed from the union of two gametes which differ in their chromosome
constitution; with reference to gene differences, having different alleles
Homology: Similarity in the structure of plant organs or chromosome which is due to their descent from a
common ancestor.
Inbreeding: Mating of closely related plants. This is done by ‗selfing‘ herma-phrodite flowers. Inbreeding
increases homozygosity in further generation of inbred-line.
Introgression: The process in which genes are transferred from one species to another through repeated
hybridization and back-crossing.
Male sterility: A condition in which the pollen is not able to produce functional pollen grains
Mass selection: Several plants are selected on the basis of their phenotype and their seed is composited
to raise the next generation. The process is further repeated till desired results are achieved.
Maternal inheritance: Inheritance where the offspring takes after the mother
Meristem culture: Cultivation of apical meristems, particularly shoot apical meristem, for production of
shoots and plantlets
Metaxenia: Immediate effect of pollen grains on maternal tissues of fruits like pulp colour, size, shape of
fruit and duration of harvest.
Modifying genes: Genes with no phenotypic effect of their own but change the expression of some
oligogene.
Monosomic: An individual with one chromosome less than the somatic chromosome number (2n-1)
Mutation: A sudden heritable change in characteristics of an organism. The term is widely used to cover
gene mutations and other changes due to chromosomal variations such as deletions, duplications,
inversions, inter-changes, ploidy, etc. In a strict sense, a mutation implies a chemical alteration of a gene or
a small structural chromosome change
Nullisomic: An individual having a pair of homologous chromosomes less than the somatic chromosome
number of the species (2n-2).
Ontogeny: The development history of an individual from the zygote to the adult stage
Pedigree method: A method of breeding in which individual plants are selected based on their previous
parental records
c. Phenotypic – the total variance, the sum of the environmental and genetic variance
Phenotypic ratio: The proportion in which the phenotypes occur in a segregating population
Physical purity: Freedom of seed from inert matter, including broken seeds, mechanical admixture
Plant breeding: The branch of biology concerned with changing the genotype of plants so that they
become more useful. It is an art and science to change the traits of a plant character
Pleiotropism: Multiple effect of a gene. Phenomenon of a single gene affecting two or more different
characters
Point mutation: A mutation that only affects a minimal part of the chromosome, i.e, a gene or intra-genic
region
Pollen culture: Cultivation of pollen grains in vitro for producing haploid plants. Generally referred as
Anther culture
Population: A group of individuals belonging to different biotypes. A gene pool from which new types may
arise through mutation and gene recombination under the influence of natural selection
Prepotency: Ability of an individual to produce progeny which are similar to each other and to itself.
Pseudogamy: Parthenogenetic development of the egg which requires stimulation from the male gamete,
but there is no true fertilization
Qualitative character: Character showing distinct classes and little or no effect on environment
Quantitative character: Character showing continuous variation and considerable effect of environment
Race: A group of biotypes with certain properties in common, or with a certain average genetic constitution.
Recombination: The phenomenon by which the characters found distributed in the parents is combined
into an offspring by hybridization. The rearrangement of linked genes due to crossing over
Regression coefficient: The rate of change of the dependent variate on the independent variate
Restitution: The reunion of the two broken surfaces of a fragmented chromosome, the original structure
thereby being restored
Sib or Sibling: Individuals having both the parents common. Crossing between brother and sister plants.
Strain: A group of individuals similar in phenotype and often in genotype. A strain is known as variety when
released for commercial cultivation by a variety release committee.
Syngamy: Fertilization; union of the male and female gametes to form the zygote
Synthetic variety: A term used particularly with cross-pollinated plants to refer to a variety produced by the
combination of selected lines or plants.
Tapetal cell: Arises from the archesporial cells in the anther. Sorrounding the pollen mother cells, they
provide nourishment to them. These cells often have deviating number of chromosomes (polyploidy)
Variance: The average of the squared deviations of the observations from the mean of a sample drawn
from a population
Variation: Differences among individuals belonging to a single species or different species. Variation may
be due to environment or due to both genotype and environment
Variety: An agricultural variety is a group of similar plants which by structural features and performance
can be identified from other varieties within the same species
Xenia: The immediate effect of the pollen on the endosperm characters of the crossed seed
Zygote:
TERMINOLOGIES RELATED TO PLANT PHYSIOLOGY
Abaxial: Surface facing away from the stem of a plant (in particular denoting the lower surface of a leaf).
Situated out of or directed away from the axis.
Adaxial: The upper surface/side of a leaf, the surface facing the stem.
Adsorption: A chemical process that takes place when a liquid or, most commonly, a gas (adsorbate)
accumulates on the surface of a solid (adsorbent), forming a molecular or atomic film.
Aerobic respiration: The process in which glucose is converted into CO 2 and H2O in the presence of
oxygen, releasing large amounts of ATP. This process includes the krebs cycle, electron transport chain
and oxidative phosphorylation.
Agrose gel: Agarose gel electrophoresis is a method of gel electrophoresis used in biochemistry,
molecular biology, and clinical chemistry to separate a mixed population of DNA or proteins in a matrix of
agarose.
Aleurone layer : The outermost layer of the endosperm, followed by the inner starchy endosperm. This
layer of cells is sometimes referred to as the peripheral endosperm. It lies between the pericarp and the
hyaline layer of the endosperm.
Alkaloids: A class of nitrogenous organic compounds of plant origin which have pronounced physiological
actions on humans. They include many drugs (morphine, quinine) and poisons (atropine, strychnine).
Allelopathy: The chemical inhibition of one plant (or other organism) by another, due to the release into the
environment of substances acting as germination or growth inhibitors.
Amino-acids: A simple organic compound containing both a carboxyl (—COOH) and an amino (—NH2)
group, Examples: glycine, alanine, valine, leucine, iosleucine, serine, thereonine, phenylalanine, tyrosine,
tryptophane, cystine, cysteine, methionine, proline, hydroxyproline, aspartic acid, glutamic acid, hystidine,
ariginine and lysine.
Amylose: A type of starch which has a straight chain polysaccharide consisting of glucose units with ∞ 1-4
links. It is less soluble in water and gives red to purple colour with iodine.
Amylopectin: It is a branched molecule of ∞ glucose units with ∞ 1-6 links in addition to ∞1-4 links. It is
less soluble in water and gives red to purple colour with iodine.
Anabolism: The synthetic processes by which the plant makes new and more complex materials from
simple substances. These processes are generally energy consuming, Example: photosynthesis.
Anaerobic respiration: An anaerobic (without oxygen) cellular process in which organic foods are
converted into simpler compounds and chemical energy (ATP) is produced.
Anion: A negatively charged ion, i.e. one that would be attracted to the anode in electrolysis. Example:
Cl, S¯.
Anoxia: Absence of oxygen.
Antagonism: Refers to reversing the usual effect of one compound (or nutrient) by another.
Anthocyanin: A group of colored glycosylated flavanoid pigments usually imparting blue, purple and red
colours to flowers and fruits in plants.
Anti-auxins: A chemical compound that inhibit completely the action and transport of auxins, Example:
TIBA, coumarin, maleic hydrazide, etc..
Anti-gibberellins: Synthetic chemical compounds that antagonize the action of gibberellins, e.g. CCC,
AMO-1618, Phosphon-D, etc.
Antitranspirants: The substances when applied to plant are able to retard their transpiration.
Apomixis: Asexual reproduction in plants, in particular agamospermy. The development of an embryo
without the occurrence of fertilization. Parthenogenesis is one form of apomixis. In plants, another form of
apomixis also occurs, in which the embryo develops from the somatic cells of the ovule surrounding the
embryo sac, not from the egg cell within the embryo sac itself. Such embryos are clones of the parent plant,
and valuable cultivars of plants such as the fig are propagated using seeds produced through this kind of
apomixis.
Assimilation: The conversion of simple molecules into the complex constituents of the living body.
Atom: The smallest unit of an element that can take part in a chemical reaction. It consists of a minute
nucleus carrying positive charged protons and neutral neutrons, surrounded by negatively charged
electrons that revolve around the nucleus in orbits. Atom as a whole is electrically neutral.
Auxins: Plant growth hormones with biological activities similar to indole 3 acetic acid (IAA) in
physiological action. They characteristically induce cell elongation in shoot cells. These are active in
processes like root initiation, fruit set etc. The naturally occurring auxin is indole 3 acetic acid. Some
synthetic auxins are indole butyric acid (IBHA) and alpha- napthalene acetic acid (NAA) etc.
Basipetal: Movement of substances from the growing tip of a shoot towards the base (junction of the root
and shoot).
Basipetal transport: Transport of substances away from the apical meristem in both root and shoot.
Beta carotene: A carotenoid hydrocarbon pigment (C 40H56) found widely in plants. Always associated with
chlorophylls.
Bioassay: Quantitative estimation of a known or suspected biologically active substance such as a
hormone by measuring its effect on a living organism in standard condition.
Biological yield: It refers to total dry matter produced by a plant or a crop in unit land area.Total dry
matter includes both straw and grain yields.
Biosynthesis: The synthesis of organic molecules by the living organisms using energy (ATP) and
reducing powers (NADPH2) generated from respiration or photosynthesis.
Biotic: Refers to something related to living. The term is used for a stress which is caused by living
organisms like insects and pests.
Biotin: A vitamin that acts as a coenzyme in carboxylation reaction.
Bolting or shooting: Elongation of stem that bears flower primordia in a rosette plants.
Bud: An undeveloped rudimentary shoot or flower.
C3 plants: Plants with C3 cycle (Calvin cycle) in which first stable product of CO 2 fixation in photosynthesis
is 3 PGA (a 3 carbon compound). Examples: rice, wheat, barley.
C4 plants: Plants with Krantz type of anatomy of leaves and Hatch-Slack pathway in which first stable
product of photosynthesis CO2 fixation is oxaloacetate (a 4 carbon compound). Examples: maize,
sugarcane, sorghum , etc.
CAM plants: Plants (some non-halophytic succulent and semi-succulent) in which crassulacean acid
metabolism (CAM) occurs wherein atmospheric CO 2 is fixed at the night into malate that is stored in
vacuole. During the following day, malate moves out from vacuole into the cytoplasm and is decarboxylated
to release CO2. The later is then is assimilated in chloroplast stroma through Calvin cycle.
Catabolism: A degradation processes by which complex molecules are converted to simple ones.
Generally energy is liberated in such reactions, Example: respiration.
Chelator: A carbon compound that can bind with certain metal cations thereby facilitating their biological
activity.
Cytochrome: Haem (iron-porphyrin) containing protein, functions as electron carrier in oxidative
phosphorylation and other oxidation-reduction reactions.
Cytokinins: A group of naturally occurring plant growth hormones that are amino-purine derivatives having
specific effect on cytokinesis (cell division), also have other physiological effects in plants. Zeatin is the
most abundant and widely distributed natural cytokinin in higher plants.
Degeneration: Loss of vitality or vigour of a biological system due to physiological disorder, disease or
other unfavourable conditions.
Desiccation tolerance: In some xerophytic plants, the protoplasm is able to tolerate or endure extremely
negative water potential or extreme water stress without being killed.
Devernalization: The positive effect of low temperature treatment on the vernalization of the plants can be
counteracted by subsequent high temperature treatment.
Dialysis: Removal of small molecules from a macromolecule preparation by allowing them to diffuse
across a semi-permeable membrane into water.
Dicotyledonous plants: The plants which have two cotyledenary leaves in the seed. They generally
possess reticulate venation and tap root system.
Diffusion: The net movement of molecules of a substance from a place of its higher concentration to a
place of its lower concentration.
Distal or proximal end: The part of fruit, leaf, tuber or root farthest from its connections with the plant
bearing is distal end.The part of the organ that is closer to the connection with the plant bearing is proximal
end.
Dormancy or rest: A state of arrested growth of a living plant or its part such as seed, bud, tuber, bulb
etc. It is due to unfavourable environmental factors or biochemical and physical factors. Examples: bud
dormancy in winter, seed dormancy due to thick seed coat or due to high inhibitory activity.
Drought resistance: The capacity of a plant to limit and control consequences of water deficit.
Economic yield: Economically useful part produced by a plant or crop in unit land area, e.g. in case of
wheat, it is the grain yield.
Ecotype: A variety or stain adapted to a specific environment.
Elasticity: Refers to reversible extension.
Enzyme: A substance produced by a living organism which acts as a catalyst to bring about a specific
biochemical reaction.
Epinasty: Downward curvature of leaf resulting from asymmetric growth on two sides of petiole, more
growth on upper (adaxial) side than on lower (abaxial) side.
Ethylene: A gaseous plant hormone with diverse effects on growth and development of plants and
commonly known as fruit ripening hormone
Ethephon:(2-Chloroethyl phophonic acid). An ethylene-releasing compound that is practical in use. Its
trade name is ethrel.
Flooding injury: Water logged condition may injure or kill plants as it creates oxygen deficiency in the root
zone. This results in the ‗physiological drought‘ in plant.
Florigen: An universal floral hormone in plants, synthesized in leaves and tranlocated to the apical tips of
stem resulting in flowering.
Foliar nutrition: A method of supplying nutrients by spraying on aerial portion of plant, particularly on
leaves.
Free radical: An atom or group of atoms that has at least one unpaired electron and is therefore unstable
and highly reactive. Free radicals can damage the cells and cellular constituents.
Gas chromatography: A method for separating the complex mixture of molecules in which a mobile
phase is caused to move in contact with selectively absorbant stationary phase.
Gel: A highly hydrated, dispersed network of long polymers, with elastic properties intermediate between
that of a liquid and a solid.
Glycolysis: A series of biochemical reactions in respiratory process, in which hexose sugar is converted
to two molecules of pyruvic acid with the net gain of two ATP molecules. This process takes place in
cytoplasm.
Glycoprotein: In addition to simple proteins, glycoproteins contain some carbohydrates.
Harvest index: The ratio of harvest yield such as grains to the total biological yield (total dry matter of
shoot with reproductive parts and root), expressed in per cent.
Heat shock proteins: A specific group of proteins produced in cells in response to rapid rise in
temperature; act as chaperons that facilitate denatured or unfolded polypeptides to snap back to their
native conformation
Heliotropism: The growth of plants or plant parts (esp. flowers) in response to the stimulus of sunlight, so
that they turn to face the sun
Hydrolysis: The breaking down of a chemical compound into two or more simpler compounds by reacting
with water.
Hydrophilic: Meaning, "water loving". Substances that have an affinity for water often because of the
formation of hydrogen bonds.
Hydrophobic: It is the property of a substance to repel water. Hydrophobic molecules tend to be nonpolar
molecules and group together. Examples: Oils and fats.
Hydroponics: The practice of growing plants in nutrient enriched water without soil.
Hypoxia: A condition in which oxygen concentrations fall below the level necessary to sustain plant life.
Hypoxic condition occurs in water logged soil.
Ideotype: A plant with its morphology ideally suited for getting maximum produce under a particular
environmental condition or set of conditions.
Incipient wilting: Initial sage of wilting of leaves which can easily recover with the availability of water.
Induced thermotolerance: Tolerance of the plants to lethal high temperature stress as the result of pre
treatment with sub lethal temperature.
Isotopes: Atoms of the same element with the same atomic number but with different atomic weights.
Example: C14 is an isotopic carbon.
Isozymes: Multiple forms of an enzyme catalyzing the same biochemical reaction but have different
molecular structure and kinetic properties.
Jasmonic acid: An important signaling agent in plant defense response against fungal, pathogens, insects
and other herbivorous. Derived from unsaturated fatty acid (linolenic acid) and found in membrane lipids.
Also act as a plant growth regulator.
Leaf area index: Leaf Area Index (LAI) is a dimensionless quantity that characterizes plant canopies. It is
defined as the one-sided green leaf area per unit ground surface area (LAI = leaf area / ground area, m2 /
m2) in broad leaf canopies.
Leaf primordia: Region of the shoot apical meristem that forms a leaf during the normal course of
development.
Lenticels: Openings in the bark through which gas exchange can take place in the stems of woody plants.
Meiosis: Cell division in which chromosome pair separates and passes to daughter cells resulting in
reduction of chromosome number into half. It occurs during production of gametes in the sexual
reproductive growth.
Membrane permeability: The extent to which membranes permits or restricts the movement of the
substances.
Meristem: The region whose tissue is composed of highly active cells undergoing cell division during
growth, e.g. growing shoot and root apex and cambium.
Metabolism: The sum total of the enzymatic reactions (anabolism and catabolism) that occur in living cells.
Mineralization: Process of breaking down of organic compounds by soil microorganisms that release
mineral nutrients in forms that can be absorbed by the plants.
Mitosis: Cell division in which each chromosome splits longitudinally and the halves pass on to the two
daughter cells, each of which is identical to the original keeping the same number of chromosome.
Molar solution: A solution in which one gram molecular weight of a substance is dissolved in a solvent and
made up to one liter in volume.
Molality: The expression of the number of moles of solute per litre of solution.
Molecular biology: The branch of biology that deals with the structure and function of the macromolecules
(e.g. proteins and nucleic acids) essential to life.
Nucellus: The tissue in the central part of the ovule in which the embryo sac is embedded.
Osmotic potential: The change in free energy of water due to the addition of solute is called the osmotic
potential of the solution.
Paclobutrazol: An inhibitor of gibberellin biosynthesis that blocks synthesis of GA 12-aldehyde onthe
endoplasmic reticulum in stage II of GA biosynthesis,
Paraquat: A herbicide that blocks photosynthetic electron flow by accepting electrons form photosystem I.
Permanent wilting percentage: The percentage of soil water left after the plant growing in that soil has
permanently wilted.
pH: The negative logarithm of the hydrogen ion concentration in aqueous solution.
Photoperiod: The relative length of certain day and night required by the plants in order to flower.
Photoperiodism: The response of plants to the photoperiod expressed in the form of flowering.
Phototropism: Tropical movements in plants in response to unilateral light stimulus enabling stem tips
bends towards the latter.
Physiological drought: Non-availability of water to the plant due to unfavourable physiological conditions
such as water logging, soil salinity, low temperature etc.
Phytoalexins: Antimicrobial substances produced in plants in response to fungal or bacterial infection.
Phytochrome: A proteinaceous pigment responsible for inhibition of flowering in short day plants and the
stimulation of flowering in long day plants. It is present in two interconvertible forms Pr and Pfr.
Phytohormones: Organic substances produced naturally in higher plants, controlling the growth or other
physiological functions at a site remote from its place of production and active in minute amounts.
Phytotron: A plant growth chamber with environment control facility where it is possible to make available
various combinations of light, temperature and humidity.
Potometer: An apparatus used for measuring transpiration (or more appropriately water absorption).
Primary growth: Growth, which is initiated by apical stem and root meristems.
Primary metabolites: Substances that are found in all plants and play reasonably well-understood roles in
the physiology of the plant. Example: carbohydrates and proteins
Reducing and non reducing sugars: Sugars possessing a free aldehyde or ketone group are reducing
sugars. Examples: Glucose, fructose.
When the above groups are locked up in glycosidic linkages in the formation of complex sugars, the
resulting sugars are said to be non-reducing sugars. Example: sucrose
Relative humidity: The amount of water vapor in the air at any given time is usually less than that required
to saturate the air. The relative humidity is the percent of saturation humidity.
Root pressure: A positive hydrostatic pressure in xylem of roots
Salinity: High concentration of total salts in the soil.
Salinization: The accumulation of salts particularly sodium chloride and sodium sulphate in soil often due
to irrigation of underground water.
Stomatal conductance: A measurement of the flux of water and carbon dioxide through the stomata in
and out of the leaf. Inverse of stomatal resistance.
Stomatal resistance: A measurement of the limitation to the free diffusion of gases from and into the leaf
through stomatal pores. Inverse of stomatal conductance.
Temporary wilting: Wilting condition of the plant due to loss of water. The plants are able to recover from
wilted condition during the night without addition of any water.
Thermoperiodism: The effect on the growth and reproduction of plants of timed exposures to varied
temperatures.
Thigmotropism: Induced movement of growth that occur in response to an external touch stimulus and are
very common in plants which climbed by tendrils.
Tissue culture: Method by which isolated plant cells are grown in laboratory on a liquid culture medium.
Totipotency: The capability of any living cell of a plant in generating an entire new plant
Toxicity: Harmful effects of a substance on plants due to its application above the tolerance limit.
Transcription: Transfer of genetic information from DNA to RNA by synthesis of complementary m-RNA
molecule on DNA template.
Translation:The process in which a particular base sequence on mRNA specifies the sequence of amino
acids during protein synthesis.
Transmination: Enzyme catalyzed transfer of amino group from an α-amino acid to keto acid. The
enzymes catalyzing the reactions are known as transaminases. Example: aminotransferases.
Transamination reactions involving glutamic acid are most prevalent in plants.
Transpiration: The loss of water from the aerial portions of living plants in the form of vapour. This occurs
principally from leaves through stomata. It may be stomatal, cuticular and lenticular transpiration.
Vernalization: The acquisition or hastening of the ability of the plants to flower by chilling treatment.
Vernalization treatment may be given to seed or whole plant. Generally 0 to 7°C temperatures are
effective for vernalization. For low temperature to be effective, oxygen, water and adequate supply of
carbohydrates for respiration are essential.
Wilting: Loss of turgidity in plants leading to a flaccid state, due to turgor pressure falling to zero.
Xylem: Conductive tissues in higher plants through which water and solutes move from roots to shoot. It
consists of trecheary elements, vessel elements, xylem fibers and xylem parenchyma.
Xylem loading: The process whereby ions exit the symplast and enter the conducting cells of the xylem.
MULTIPLE CHOICE QUESTIONS
2. Fruit crop suitable for high soil pH (7.2 - 8.0 & above) is
a. Aonla b. Custard apple
c. Guava d. Ber
c. Pome d. Berry
5. Origin of Aonla is
a. China b. India
c. Chakkaiya d. Francis
7. which of the following anolavariety which highly susceptible to fruit necrosis disorder?
a. Francis b.Chakaiya
c. BSR-1 d.Banarasi
9. The fruit crop which can be grown in wide range of sodic soil is
a. Papaya b. Aonla
c. Guava d. Sapota
10. Three tier fruit crop model system which suits for sodic soil is
a. 70% b. 20%
c. 50% d.10%
13. Aonla is best suited to grow under which one of the following soil condition?
14. Which one of the following banana cultivar has got ―Geographical Indication‖ status?
a. Poovan b. Rasthali
c. Monthan d. Robusta
18. ----------------------- is the fruit crop that occupies the highest area in India
a. Mango b. Banana
c. Citrus d. Grapes
a. Banana b. Apple
c.Papaya d. Citrus
23. The explant used for commercial tissue culture of banana is
a. Leaf bits b. Stem bits
c. Shoot tips d.Petiole
24. The genomic constitution of Cavendish group of banana is
a. AAA b. AAB
c. ABB d. AA
25. The operation in banana where 60 cm of the mother plant from the ground level after harvest is retained
for nutrient supply to daughter suckers is
a. Desuckering b. Denavelling
c. Mattocking d. Propping
26. --------------state is the leader in banana production in India
a. Uttar Pradesh b. Maharastra
c. Andhra Pradesh d. Tamil Nadu
27. The spacing recommended for HDP(3 suckers/clump) in banana by TNAU is
a.1.8m x 1.8m b. 2.0m x 2.0m
c. 3.6m x 1.8m d.1.8m x 3.6m
28. Which of the following is banana hybrid
a. Co-1b b. PKM-1
c. Poovan d. IIHR -1
29. Banana is otherwise called as
a. King of the fruit b. Queen of the fruit
c. Apple of Paradise d. Prince of fruit
30. Which of the following is triploidaccuminata commercial variety of banana?
a. Grand Naine b.Nendran
c.Poovan d.Virupakshi
31. Banana is commercially propagated through
a.Water suckers b. Sword suckers
c.Peepers d. Tissue culture of floral buds
32. Removal of male bud after completion of female phase in banana is referred as
a.Tipping b.Denavelling
c.Thinning d.Pinching
33. The banana fruits are packed in 100 gauge polybags with following % holes to increase the shelf life
a. 0.2% b. 2.0%
c.5.0% d.10.0%
34. Ney poovan variety of banana belongs to ------------------------- genome group of banana.
a.AA b. AAA
c. AB d.AAB
35. Paring and Pralinage is followed in banana to control
a. Aphids b. Stem borer
c. Nematodes d. Fusarium wilt
36. Tissue culture propagation is gaining importance in -------------------- crop.
a. Citrus b. Grape
c. Banana d.Papaya
37. --------------- is the one of the popular parthenocarpic fruits.
a. Banana b. Mango
c. Citrus d.Grapefruit
38. Storage temperature for banana is --------------.
a. 5-10°C b. 10-15°
c. 13°C d.20-21°C
39. The major problem in banana breeding is----------------------
a. Long duration b. Seedlessness
c. Complex flower structure d. Plant height
40. For ripening in banana, ----------------------- is kept along with Ethrel
a. Sodium chloride b. Calcium Chloride
c. SodiumHydroxide d. None of these
41. Which of the following is a multiple hybrid of Banana
a. Bombay Green b. CO.1
c. Plantain d. None of these
42. Banana genomic classification was given by
a. Capsule b. Nut
c. Pome d. Berry
44. AmritSagar is a variety of
a. Banana b. Guava
c. Fig d. Avocado
45. A chemical used for artificial ripening in banana
a. Ethephon b. Auxin
c. KNO3 d. Calcium carbide
46. Bunchy top viruswas entered into India from
a. USA b. Japan
c. Australia d. Sri Lanka
47. Choke throat in banana is due to
a. High Humidity b. Low Humidity
c. High temperature d. Low Temperature
48. Kokkan disease in banana is caused by
a. Fungus b. Bacteria
c. Virus d. MLOs
49. Major bottleneck in banana breeding is
a. Pollen sterility b. Non- receptivity of stigma
c. Vegetative parthenocarpy d. All the Above
50. Banana Bract Mosaic Virus (BBMV) is serious disease in
a. Poovan b. Rasthali
c. Monthan d. Hill Banana
51. Seediness in banana can be controlled by the spraying of
a. NAA @ 25 ppm b. 2,4-D @ 25 ppm
c. Kinetine @ 25 ppm d. ABA @ 25 ppm
52. Improper bunch filling in banana is due to
a. K deficiency b. N deficiency
c. B deficiency d. Zn deficiency
53. Tetrazolium test is used for the diagnosis of presence of
a. Fungus b. Bacteria
c. Virus d. MLOs
54. Water logging condition in banana cultivation leads to the more incidence of
a. Panama wilt b. Sigataka leaf spot
c. Anthracnose d. Mokowilt
a. Seeds b. T - budding
c. Inarch grafting d. Air layering
75. Salinity and alkalinity tolerant variety of ber is ……………
a. Gola b. Umaran
c. Mahatawali d. Mundia
76. Ber varieties Katha, Tikdi, Sanaur No.1, Mehrun and Dodia are
a. Ber b. Litchi
c. Guava d. Mango
78. Pollen Sterility is major problem in …………fruit
a. Grape b. Guava
c. Ber d. Pomegranate
79. Which of the following fruit has non-endospermic seed?
a. Ber b. Beal
c. Aonla d. Mango
80. Ganesh Kirti is a variety of
a. Grape b. Guava
c. Ber d. Pomegranate
81.India is the largest producer of .................in the world.
a. Lime b. Sweet orange
c. Grapefruit d. Mandarin
82. Mosambi, a popular sweet orange cultivar is predominantly grown in
a. Punjab b. Maharashtra
c. Rajasthan d. Andhra Pradesh
96. Colour pigment in the flavedo of mature ripe orange fruit is………….
a. Hesperidine b. Naringin
c. Carotene d. Chlorophyll
97. Common rootstock for budding of mandarin is
a. C. aurantium b. C. sinesis
c. C. limonia d. C. jambhiri
98. Degreening in citrus is done with application of
a. Ethephon b. GA3
c. Cytokinin d. Auxin
99. Flying dragon is the most dwarfing rootstock for…………
a. Citrus b. Grape
c. Mango d. Apple
100. Granulation can be reduced by the spraying of…………..
a. Leaf arsenate b. Urea
c. Copper oxide b. KNO
d. Urea3
101. Growth regulator commercially used for control for fruit drop in citrus is
a. NAA b. 2, 4 - D
c. IBA d. ABA
102. Hesperidium is the botanical form of ------------fruit
a. Citrus b. Grape
c. Mango d. Apple
103. Indicator plant for Triteza virus in citrus is ----------------------
a. C. aurantifolium b. C. sinesis
c. C. reticulata d. C. paradisi
104. Kinnow is a cross between
a. C. nobilis and C. deliciosa b. C. reticulata and C. sinesis
c. C. reticulate and C. sinensis d. C. nobilisand C. sinensis
105. Monoembryonic species of citrus is
a. Pummello b. Grapefruit
c. Sweet orange d. Mandarin
106. On tree storage in citrus is possible with application of
a. GA3 b. 2, 4 - D
c. Thio urea d. ABA
107. Only citrus fruit which contain Malic acid is……………
a. Kagzi lime b. Sweet lime
c. Sweet orange d. Kinnow
108. Lemon squash is preserved for a longer period without the discolouration by use of
a. Malic acid b. Citric acid
c. Benzoic acid d. Potassium meta bisulphite
a. Kinnow b.Atemoya
c. Cultivated strawberry d. All of these
130. Origin of Citrus reticulata is
a. Japan b. China
c. India d.West Indies
131. Fruits of which of the following group is mostly seedless?
a. Hexaploid b. Tetraploid
c. Diploid d. Triploid
208. Among the following Artocarpus species,------------ is edible species.
a. A.marianensis b. A. hirsutus
c. A. chaplasha d. A. lakoocha
209. In Jack, female flower(spike) is having………surface and……..in size than male flower
a. Pineapple b. Mulberry
c. Jackfruit d. All of the above
211. Precocious bearing cultivar of Jackfruit is
a. Stimulative b. Stenospermocarpy
c. Vegetative d. None of these
248. Alphonso a variety of mango is mostly grown in
a. Punjab b. UP
c. Maharashtra d. Bihar
249. Early maturing mango variety of south India is
a. Neelum b. Dashehari
c. Alphonso d. Banganapalli
250. Which of the following is an off season variety of mango in north India ?
a. Mallika c. Fazli
c. Niranjan d. Olour
251. Ideal variety for long distance transporting in mango is
a. Neelum b. Langra
c. Bombay green d. Rumani
252. ------------------ is a mango hybrid released by IARI, New Delhi.
a. Mallika b. Rathna
c. Sindhu d. Alphonso
253. Alphonso is the one of the parents for which of the following mango hybrid?
a. ArkaAruna b. Pusa Lalima
c. PusaPeetamer d. Pusa Giant
254. Sunlight interception is the primary aim in mango for
a. Training b. Pruning
c. Top working d. Smudging
255. Jelly seed is a physiological disorder in -------------------.
a. Apple b. Mango
c. Guava d. Grapes
256. In which group will you classify mango according to its ethylene production rate?
a. Very low b. Low
c. Moderate d. High
257. Sap burn is a postharvest disorder in ----------------
a. Mango B. Apple
c. Orange d. Grape
258. Among fruits, the highest vitamin A content is in mango. It is about
a. 200 IU/100g b. 1500 IU/100g
c. 4000 IU/100g d. 3000 IU/100g
259. Mango hybrid Mallika is a cross between
a. Dashehari X Langra b. Neelum X Dashehari
c. Neelum X Langra d. Banganapalli X Alphonso
260. Spongy tissue, a serious problem in mango is seen in
a.Totapuri b. Malda
c. Alphonso d. Langra
261. Tapka is associated with maturity of
a. Litchi b. Aonla
c. Pineapple d. Mango
262. A common pest during flowering time of mango (Feb-March) is
a. Red ants b. Shoot borer
c. Hoppers d. Mealy bug
263. A disease which can transform an ―On year‖ of fruiting to ―Off year‖ in mango is
a. Powdery mildew b. Anthracnose
c. Leaf blight d. Stem canker
264. A serious physiological disorder of mango orchards near brick kiln is
a. Black tip b. Black rot
c. Spongy tissue d. Malformation
265. Amimgola is a variety of
a. Litchi b. Guava
c. Fig d. Mango
266. Black tip of mango is also known as
a. Chimney disease b. Soft nose of mango
c. Witch‘s broom d. Seed jelly of mango
267. Creeper is a variety of
a. Litchi b. Kiwifruit
c. Grape d. Mango
268. Dwarf mango suitable for kitchen garden is
a. Amrapali b. Alphonso
c. Mallika d. Dashehari
269. Flowering in mango can be suppressed by the application of
a. Auxin b. Gibberellins
c. Kinetin d. Paclobutrazol
270. Hot water treatment for control of anthracnose in mango is at
a. 52oC b. 32oC
c. 70oC d. 100oC
271. In situ method of mango grafting is
a. Soft wood grafting b. Inarching
c. Epicotyl grafting d. Stooling
a. Bangalora b. Neelum
c. Amrapali d. Dashehari
296. Which of the following fruits even though propagated by seed do not have variability
a. Litchi b. Mangosteen
c. Loquat d.Papaya
297. Which of the following pair is not correctly matched?
aDatepalm - Monoecious. b.Papaya - Polygamous
c. Walnut - Monoecious d.Pistachionut – Dioecious
298. A transgenic papaya variety is
a. Maradol b. Red Lady
c. Sunset Solo d.Sun Up
299. Scientist associated with studies on sex forms in papaya is
a.Drew b. N.W.Simmonds
c. W.B.Storey d.R.W.Allard
300. Dioecious mutant variety of papaya is
a. CO.8 b. Washington
c. Pusa Giant d.PusaNanha
301. Papain is ------- in water
a. Insoluble b. Highly soluble
c. Moderately d. Less soluble
302. ---------------- is a transgenic papaya variety developed against Papaya Ring spot Virus
a. Surya b. Rainbow
c. PusaNanha d.Red lady
303. --------------------- is added as preservative to wet latex from papaya before drying
a. Potassium chloride B. Sodium chloride
c. Potassium meta bisulphite D.Potassium dihydrogen phosphate
304. CO. 1 papaya is derived from,
a. Ranchi b. Washington
c. Sun Rise Solo d.Coorg Honey Dew
305. The seed rate for papaya for one hectare is
a. 250 g b. 500 g
c. 625 g d. 1000 g
306. The most devastating disease in papaya is
a. Foot rot b. Collar rot
c. Papaya ring spot virus d. Leaf spot
307. Cold tolerant papaya species is known as ----------------------------.
a. Caricacandamarcensis b.Carica papaya
c. Caricacauliflora d.Caricagoudotiana
308. CO 3 and CO 7 are examples of --------------------- varieties of papaya
a. Dioecious b. Monoecious
c. Gynodioecious d.Andromonoecious
309. Femaleness in papaya is controlled by genotype
a. M1M b. M2M
c. M1M2 d. mm
310. CO.8 papaya variety is a cross between
a. CO.2 X Sunrise Solo b. Sunrise Solo X Pink Flesh Sweet
c. Co.2 X Red Anthered Male parent (Co-2) d. CO.1 X Washington
a. 7-9˚C b. 14-15˚C
c. 10-13˚C d. 6-7˚C
329. Among, which species of papaya is resistant to frost
a. C. pubescence b. C. monoica
c. C. pentagona d. C. cauliflora
330. Among the following varieties, which variety suitable for dual purposes(Dessert purpose and papain
extraction)………………?
a. CO.3 b. CO.2
c. CO.4 d. CO.5
331. The only red flesh cultivar in dioecious type of papaya is
a. PusaNanha b. CO.3
c. Solo d. CO-8
332. Popular IIHR variety with good peeling quality
a. Arkasurya b. Solo
c. Washington d. Ranchi
333.Proteolytic enzyme present in papaya is……………
a. Bromeline b. Caricaxanthin
c. Papain d. none of these
a. Taiwan b. Solo
c. Red lady d. Eksotica
336. The seed rate for gynodioecious cultivar of papaya (g/ha) is
a. 200-250 b. 300
c. 400 d. 500
337. The ratio of male and female in dioecious line is maintained by
a. 1:20 b. 1:10
c. 1:15 d. 1:8
338. Which of the following fruit crop is called as ‗tree melon‘?
a. Papaya b. Jackfruit
c. Mango d. Durian
339. Example for continuous flowering and fruiting crop is
a. Papaya b. Sapota
c. Mango d. Banana
340. Sib mating is practiced in
a. Papaya b. Banana
c. Mango d. Guava
341.Example for monoecious papaya species is………….
a. Vasconcelleamonoica b. V. candamorcensis
c. V. cauliflora d. V. gudotiana
342. Red pulpcolour is present only in .......................papaya
a. Monoecious b. Dioecious
c. Gynodioecious d. None of these
343. How many species in the genus Carica ?
a. 2 b. 4
c. 1 d. 3
344. Jalore seedless, a pomegranate variety is recommended for cultivation in the state of
a. Rajasthan b. U.P
c. Gujarat d. Maharashtra
345. Jyothi pomegranate is a clonal selection from
a. Kandhari b. Alandi
c. Ganesh d. Bassein seedless
a. M. hexandra b. M. kauki
c. M. bileratad. none
a.Spinddle b. Oval
c. Round d. Oblong
363. Edible part of Sapota is
a. Placenta b. Mesocarp
c. Endocarp d. Endosperm
364. Chickle is extracted from………..parts of sapota
a. Root, stem b. Bark, immature fruit
c. Ripen fruit, seed d. none
365. Flattening of branches in Sapota is due to
a. Fungus b. Bacteria
c. Virus d. MLO
366. Round shape variety of Sapota is
a. CO1 b. PKM2
c. CO3 d. PKM1
367. Central Institute for Arid Horticulture (CIAH) is located at __________.
a. Jodhpur b. Abohar
c. Bikaner d. Hissar
368. The headquarters of Bioversity International is located at ___________.
a. Nigeria b. Germany
c. France d. Italy
369. International year of biodiversity as per declaration of UN…………….
a. 1997 b. 2009
c. 2010 d. 2011
370. Which is the cheapest preservation method of fruits?
a. Canning b. Freezing
c. Drying d. Fermentation
371. In which form, pectin is found in fruit?
a. Pectin b. Calcium pectate
c. Proto-pectin d. None of these
372. During controlled atmospheric storage, composition of which of the following set of gases is
controlled?
a. O2+N2 b. CO2+N2
c. C2H4+N2 d. CO2+O2
373. In precooling field heat is mostly removed by -----------------.
a. Conduction b. Convection
c. Radiation d. None of these
374. Degreening of fruits is caused by -----------------.
a. Loss of chlorophyll b. Loss of lycopene
c. Loss of anthocyanin d. Loss of carotene
375. The enzyme which is responsible for softening of fruits
a. Poly sulphonase b. Catalase
c. Polyhydrase d. Polygalactouranase
376. ------------------- hormone helps in improving fruit set in fruit trees
a. Ethylene b. Abscissic acid
c. Salicylic acid d. NAA
377. Which of the following chemical is used as Cryoprotectant
a. DMSO b. 2, 4-D
c. Ethylene d. EMS
378. Enzyme responsible for formation of ethylene is --------------------.
a. ACC synthase b. ACC transferase
c. ACC Oxidase d. Adenosyl methionine
379. The fruit product order (FPO) license is given by -----------------
a. GOI b. NHB
c. ICAR d. State Government
380. As fruits mature, the specific gravity will ----------------.
a. Increase b. Decrease
c. Remains constant d. None of these
381. In India, processing industries utilize ……..% of total fruit production?
a. >1% b. <1%
c. 2-3% d. 5%
382. Which of the following is a rapid precooling method?
a. Forced air cooling b. Hydro cooling
c. Vacuum cooling d. Evaporate cooling
383. Calcium carbide releases ---------------.
a. Carbondioxide b. Ethylene
c. Acetylene d. Methane
384. SINGER is related to __________.
a. Field gene bank b. In vitro conservation
c. Data base management d. Cryo preservation
385. The aroma in fruit is caused due to the formation of ---------------------.
a. Fats b. Essential oil
c. Esters d. Aromatic compounds
398. ------------- is the important certificate needed for export of horticulture produce
a. HACCP b. Sanitary &Phytosanitary
c. Organic certification d. GAP
399. ----------------------- is the most important activity expected in plant system after layering.
a. Rooting b. Heeling
c. Shooting d. Flowering
400. CISH, is located at _____________.
a. Bangalore b.Trichy
c. Lucknow d. Srinagar
401. The nucellar embryos develop through the process of__________.
a. Poly embryony b. Bud mutation
c. Parthenocarphy d. Apomixis
402. -------------------- hormone helps in improving fruit set in fruit trees.
a. Ethylene b. Abscissic acid
c. Salicylic acid d. NAA
403. Cryo preservation is associated with -------------------.
a. Liquid Nitrogen b. Liquid oxygen
c. Liquid Carbon dioxide d. None of the above
404. Indian Institute of Horticultural Research (IIHR. is located at ----------.
a. Lucknow b. New Delhi
c. Bangaluru d. Ranchi
405. ---------------- is one of the popular hormones used for root induction.
a. GA3 b. Cytokinin
c. IBA d. ABA
406. ------------------- is the method of grafting by which old unproductive orchards are regenerated
a. Top working b. Bridge grafting
c. Epicotyl grafting d. In situ grafting
407. Chilling requirement means prevalence of low temperature conditions for a duration in case of ----------
----------- crops.
a. Humid zone b. Subtropical
c. Temperate d. Aridzone
408. ---------------- is the hormone which promotes vegetative growth thereby delaying flowering
a. Auxin b. Gibberellin
c. Cytokinin d. Abscissic acid
409. Alternate bearing is a problem caused by ---------------
a. Nutritional deficiency b. Unbalanced growth of vegetative anreproductive
c. Fungal infestation parts
d. Water deficit
410. --------------- is the basic concept in organic farming.
a. LEISA b. IFOAM
c. EUREPGAP d. HACCP
411. ------------------nutrient is required at higher quality in fruit crops for quality enhancement
a. N b. P
c. K d. Fe
412. Under high density planting, -------------- is an important requirement.
a. Gap filling b. Weeding
c. Nutrition at higher doses d. Application of growth regulator
413. Pollinizers are essential in --------------- fruit crops for good fruitfulness
a. Tropical b. Subtropical
c. Temperate d. Arid
414. --------------- is the recent and improved method of storage for fruits and vegetable.
a. Cold storage b. MA storage
c. Ambient conditions d. Vacuum packing
415. By waxing -------------- process is showed down in fruits and vegetable during storage
a. Ripening b. Respiration
c. Transpiration d. Photosynthesis
416. Which refrigerant is commonly used in cold storage in India?
a. Ethylene b. Calcium carbide
c. Ammonia d. Sodium benzoate
417. Chemical used for removing the spray residues of Arsenic and Lead from the fruit is ----------
a. NaOH b. HCl
c. NaCl d. H2SO4
418. Alcholol content of wine ranges from -------------.
a. 5-10% b. 10-12%
c. 7-20% d. 12-17%
419. Which of the following is a rapid precooling method?
a. Forced air cooling b. Hydro cooling
c. Vacuum cooling d. Evaporate cooling
420. Amount of pectin in jelly is tested by -----------.
a. Penetrometer b.Refractometer
c. Thermometer d. Jelmeter
421. ---------------- is the recent method followed in citrus for production of disease free planting materials.
a. Callus culture b. Micro grafting
c. In situ grafting d. Shoot top culture
422. The most common method for grape preservation is ----------------.
a. Jam making b. Jelly making
c. Dehydration d. Pickling
423. Stenospermocarpy is expressed in
a. Italia b. Tas-A-Ganesh
c. Red Globe d. Anab-E-Shahi
424. Plant growth regulator used to increase berry size in grape is ------------------
a. IAA b. GA3
c. Cytokinin d. ABA
425. Among the following which is false
a. Most of the viniferagrapes are self fertile b. Crosses between different Euvitisspecies can
be easily made
c. Parents with high seed index are preferable d. Euvitisand Muscadina hybrids are fertile
for breeding seedless varieties and vigourous
426. In grapes, fruit buds appear in ------------------ which is a past season shoot.
a. Shoot b. Spur
c. Arm d. Cane
427. --------------- is the rootstock which has good tolerance against nematodes and drought in grapes.
a. Dog Ridge b. 16-1
c. Khirnee d. Vellaikolumban
428. Saline tolerant root stock in grape is __________
a. Salt Creek b. St. George
c. 1613 d. Rupestnis Du Hot
429. NRC grape is located at ___________
a. Nagpur b. Dehradun
c. Hyderabad d. Pune.
430. Development of seedless varieties is possible in grape due to ------------------------.
a. Triploid breeding b. Stenospermocarpy
c. Parthenocarpy d. Hormones
431. ------------------- is a mutant of grape derived from Thompson seedless
a. Sharad seedless b. Sonaka
c. PusaUrvasi d. Tas-e-Ganesh
432. The richest source of iron is __________
a. Dry karonda b. Date palm
c. Bael d. Woodapple
433. Patch budding is generally practiced in ----------------.
a. Guava b. Sapota
c. Aonla d. Ber
434. Which of the following belongs to family Rutaceae?
a. Wood apple b. Mulberry
c. Jamun d. Ber
435. Heat units required for flowering, fruit development and ripening in date palm is ------------.
a. 3300 b. 4000
c. 2250 d.1100
a.E.amylovora b.E.carotovora
c. P.infestance d.P.syringae
490. Colouring pigment of plum is ____________
a. Carotenoides b. Anthocyanins
c. Betalains d. None of the above
491. Internal browning of apple is due to __________
a. Apple b. Pear
c. Pineapple d. Avocado
493. Richest source of Vitamin C is _________
a. January-February b. May-July
c. March-May d. February-March
495. Storage temperature of pomegranate is ____.
a. 2°C b. 7°C
c. 5°C d. 12°C
496. Highest productivity of pineapple is in ____________
a. Quince b. Apple
c. Plum d. Peach
498. African pride is a Annona variety is developed by cross between ________ and ___________
a. Sugar apple and Annonareticulata b. Sugar apple and A.glabra
c. Sugar apple and A.cherimaya d. Sugar apple and A.squamosa
499. Pollinizer for Red delicious Apple is __________
a. Jonathan b. Ambri
c. Golden Delicious d. Early Shanburry
500. Type of parthenocarpy in Litchi is _________
a. Stimulative b. Vegetative
c. Stenospermocarpy d. None of these
501. Harvesting period of cherry is
a. April b. July
c. October d. December
502. Low chilling plum variety is
a. Peach b. Plum
c. Pear d. Cherry
504. Thames Pride introduced from Japan is a variety of
a. Loquat b. Kiwifruit
c. Avocado d. Mangosteen
505. Which of the following organic acid is predominantly present in the Carambola?
a. Ca deficiency b. Mg deficiency
c. S deficiency d. None of these
507. Macadamia nut belongs to the family
a. Ebenaceae b. Actinidiaceae
c. Oxilidaceae d. Proteaceae
508. Protandry type of dichogamy obtained in
a. A.squamosa b. Fig
c. Pomegranate d. Passion Fruit
509. Gynodioecious cross pollinated fruit crop is
a. Grape b. Fig
c. Banana d. Kokum
510. In temperate fruit crops, the ―root cutting‖ are prepared during the month of
a. January b. February
c. October d. December
511. Abakka is a variety of ___________
a. Guava b. Grape
c. Pineapple d. Carambola
512. Which variety of apple is resistant to all fungal diseases?
a. Liberty b. Freedom
c. Ambri d. Ambred
513. Gamboge is a physiological disorder of
a. Durian b. Mangosteen
c. Pear d. None of these
514. Interveinalchlorosis of apple is due to the deficiency of
a. Bo b. Mo
c. Mg d. Cu
515. Placentia is a variety of
a. Pecanut b. Walnut
c. Apple d. Almond
516. Heterodichogamy is found in
a. Walnut b. Pecanut
c. Almond d. Chestnut
517. Which temperate fruit crop require highest chilling requirement?
a. Apple b. Pear
c. Almond d. Cherry
518. Flying dragon is the most dwarfing rootstock for
a. Citrus b. Guava
c. Apple d. Plum
519. Which one of the following temperate fruit is earlier in flowering?
a. Apple b. Cherry
c. Peach d. pear
520. Self sterile variety of pear is __________
a. China pear b. Patharnakh
c. Leconte d. Magness
521. Type of self incompatability in loquat
a. Sporophytic b. Gametophytic
c. Both d. None
522. Origin of custard apple is
a. Apple b. Pear
c. Peach d. Plum
526. Mulching is the most important cultural practice in the cultivation of
a. Apple b. Pear
c. Peach d. Strawberry
527. Training method followed in peach
a. September b. October
c. November d. December
529. Which chemical serves as an alternative to caprification in fig
a. SADH b. CPA
c. BOA d. CCC
530. Which variety of pear has highest TSS
a. Strawberry b. Plum
c. Peach d. Apple
533. Fruit cracking in pear is due to the deficiency of
a. Ca b. Mg
c. B d. N
534. Largest producer of Almond in the world is
a. USA b. Japan
c. China d. Afganistan
535. Origin of Apple is
a. N.America b. S.America
c. S.E.Asia d. S.W.Asia
536. Ultra dwarf root stock of apple is
a. M-9 b. M-27
c. MM-106 d. MM-111
537. Fig is
a. Dioecious b. Monoecious
c. Gynodioecious d. Andromonoecious
538. Kiwi fruit is a
a. Seeds b. layering
c. Cuttings d. stooling
540. Australian nut is
a. Walnut b. Pecanut
c. Macadamia nut d. Hazel nut
ANSWER KEYS
1. b 2. a 3. d 4. a 5. b 6. a 7. a
8. c 9. b 10. a 11. a 12. a 13. b 14. b
15. a 16. c 17. c 18. a 19. c 20. c 21. b
22. a 23. c 24. a 25. c 26. d 27. c 28. a
29. c 30. a 31. b 32. b 33. a 34. c 35. c
36. c 37. a 38. c 39. b 40. c 41. a 42. a
43. d 44. a 45. d 46. d 47. d 48. d 49. c
50. c 51. b 52. a 53. c 54. a 55. d 56. c
57. b 58. a 59. c 60. a 61. d 62. a 63. a
64. d 65. d 66. d 67. d 68. a 69. a 70. a
71. d 72. c 73. c 74. d 75. a 76. b 77. a
78. c 79. a 80. c 81. a 82. b 83. a 84. a
85. a 86. d 87. b 88. b 89. c 90. a 91. a
92. d 93. a 94. c 95. c 96. c 97. a 98. a
99. c 100. b 101. a 102. a 103. a 104. a 105. a
106. b 107. d 108. a 109. c 110. a 111. b 112. a
113. d 114. d 115. a 116. b 117. b 118. b 119. d
120. a 121. b 122. c 123. d 124. b 125. c 126. d
127. c 128. b 129. d 130. b 131. c 132. a 133. b
134. a 135. a 136. b 137. c 138. a 139. b 140. a
141. a 142. b 143. c 144. d 145. a 146. a 147. b
148. c 149. a 150. c 151. c 152. a 153. a 154. b
155. b 156. a 157. a 158. b 159. a 160. c 161. b
162. c 163. c 164. a 165. a 166. c 167. a 168. d
169. b 170. c 171. d 172. a 173. a 174. d 175. b
176. c 177. c 178. b 179. a 180. a 181. a 182. a
183. a 184. a 185. a 186. a 187. a 188. c 189. b
190. a 191. a 192. a 193. c 194. a 195. a 196. a
197. b 198. b 199. b 200. b 201. b 202. b 203. b
204. b 205. a 206. a 207. b 208. d 209. c 210. d
211. a 212. b 213. d 214. b 215. a 216. a 217. d
218. c 219. d 220. b 221. a 222. d 223. c 224. a
225. c 226. d 227. b 228. d 229. b 230. a 231. c
232. d 233. c 234. c 235. a 236. b 237. d 238. c
239. b 240. a 241. a 242. a 243. c 244. b 245. c
246. d 247. b 248. c 249. d 250. c 251. a 252. a
253. a 254. b 255. b 256. c 257. a 258. c 259. b
260. c 261. d 262. c 263. a 264. a 265. d 266. a
267. d 268. a 269. b 270. a 271. a 272. a 273. c
274. b 275. a 276. d 277. a 278. a 279. b 280. c
281. d 282. a 283. a 284. a 285. b 286. a 287. a
288. a 289. b 290. a 291. a 292. b 293. c 294. a
295. d 296. d 297. b 298. d 299. c 300. d 301. a
302. d 303. c 304. a 305. b 306. b 307. a 308. c
309. d 310. c 311. c 312. a 313. b 314. a 315. a
316. a 317. c 318. d 319. a 320. c 321. c 322. c
323. d 324. b 325. b 326. d 327. b 328. c 329. a
330. b 331. d 332. a 333. c 334. a 335. c 336. b
337. a 338. a 339. a 340. a 341. a 342. c 343. c
344. a 345. d 346. a 347. d 348. c 349. b 350. a
351. c 352. a 353. d 354. a 355. a 356. a 357. a
358. c 359. d 360. b 361. b 362. b 363. b 364. b
365. a 366. a 367. a 368. d 369. c 370. c 371. c
372. d 373. b 374. a 375. d 376. d 377. a 378. a
379. a 380. a 381. b 382. b 383. c 384. c 385. c
386. a 387. c 388. b 389. c 390. c 391. b 392. a
393. b 394. a 395. c 396. c 397. d 398. b 399. a
400. c 401. d 402. c 403. a 404. c 405. c 406. a
407. c 408. b 409. b 410. d 411. b 412. c 413. c
414. b 415. b 416. d 417. b 418. c 419. c 420. d
421. b 422. c 423. b 424. b 425. d 426. d 427. a
428. a 429. d 430. b 431. b 432. a 433. c 434. a
435. a 436. b 437. d 438. a 439. a 440. b 441. a
442. a 443. d 444. a 445. b 446. b 447. a 448. b
449. a 450. a 451. c 452. b 453. b 454. c 455. a
456. d 457. b 458. d 459. b 460. d 461. c 462. d
463. b 464. c 465. a 466. a 467. c 468. a 469. c
470. a 471. c 472. b 473. c 474. a 475. c 476. a
477. c 478. c 479. d 480. c 481. d 482. a 483. b
484. c 485. b 486. c 487. b 488. a 489. a 490. a
491. d 492. c 493. b 494. c 495. c 496. a 497. c
498. c 499. c 500. a 501. a 502. b 503. b 504. a
505. a 506. a 507. d 508. d 509. b 510. d 511. c
512. a 513. c 514. b 515. b 516. d 517. a 518. c
519. d 520. b 521. b 522. c 523. c 524. a 525. d
526. b 527. b 528. b 529. b 530. a 531. a 532. a
533. c 534. a 535. d 536. b 537. c 538. d 539. c
540. c
REFERENCES
1. Bose, T.K., Mitra, S.K and Sanyal.2002. Fruits : Tropical and Subtropical, Vol & II (Third Revised
Edition), Naya Udyog, Calcutta – 700 006
2. Chandrasekharan, S.N, Parthasarathy, S.V., Krishnaswamy and N.Hrishi. 1975. Cytogenetics and
Plant Breeding. Published by Varadachary & Co, 8, Linghi chetty street, Madras
3. Gangulee, Das and Datta, 1997. College Botany Vol.I. New Central Book Agency Pvt. Ltd., 14,
Chatterjee Street, Calcutta – 700 012
4. Gupta Narendra, K and Sunita Gupta. 2005. Plant Physiology. Oxford & IBH Publishing
Co.Pvt.Ltd., New Delhi
5. Purseglove, J.W.1969. Tropical Crops Dicotyledons Vol 1 & 2 combined. Longman, Group Ltd.,
London
6. Singh, B.D.1983. Plant breeding . Kalyani Publishers, 1/1, Rajendra Nagar, Ludiana – 141 008
7. Veeraragavathatham, D., Jawaharlal, M., Jeeva, S., Rabindran, R., and Umapathy. 2004.
Published by Suri Associates, Thadagam Road, Coimbaore - 2