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Unit I
Introduction
Intended Learning Outcomes
Lesson 1
Objectives
Content and Discussion
Assessment
Lesson 2
Objectives
Content and Discussion
Assessment
Lesson 3
Objectives
Content and Discussion
Assessment
Lesson 4
Objectives
Content and Discussion
Assessment
How to earn extra credit?
Unit Test
References
Introduction
Animals are united with all other forms of life by the biological processes that they share
with other organisms. Understanding these processes helps us to know how animals
are united with other forms of life from the evolutionary and ecological processes. This
unit comprises of four lessons. Lesson 1 examines some of these unifying themes and
sets the stage for evolutionary and ecological perspectives. Lesson 2 present cell
structure and inheritance as an important unifying framework within which biologists
approach the diversity of organisms. Lesson 3 explores the work of pioneers of
evolutionary theory, Charles Darwin and Alfred Russel Wallace and how their work
forms the nucleus for modern evolutionary theory. In addition, this lesson also examines
the influence of modern genetics on evolutionary which will provide core knowledge for
understanding the diversity of animal life and how evolution has influenced the animal
structure and function. Lesson 4 presents basic ecological principles that everyone must
understand if we are to preserve the animal kingdom.
A. Define Zoology.
The etymology of the term zoology came from the Greek work, “zoon” and “logos”,
literally translated to the study of animals. It is one of the broadest fields due to
immense variety of animals and complexity of the processes occurring within animals.
There are more than 20,000 described species of bony fishes and more than 300,000
beetle species. The subdisciplines of zoology includes functional, structural and
ecological. Specializations are often pertaining to the type of animal or what animal one
is studying.
For example: Ichthyology is the study of fishes, and ichthyologists work to understand
the structure, function, ecology, and evolution of fishes. These studies have uncovered
an amazing diversity of fishes. One large group, the cichlids, is found in Africa (1,000
species), Central and South America (300 species), India (3 species) and North
America (1 species). Cichlids of Africa (Figures 1a and 1b) exist in an amazing variety
of color patterns, habitats and body forms.
Miller-Harley 2001
Miller-Harley 2001
Figure 1.4. Lakes Victoria,
Tanganyika and Malawi. These
lakes have cichlid populations that
have been traced by zoologist to
an ancestry that is approximately
200,000 years old. Cichlid
populations originated in Lake
Tanganyika and then spread to the
other two lakes.
B. History
1. The prominent ancient Greek philosopher Aristotle took detailed notes on animal
observations. Many other scientists were inspired.
2. Many universities were founded in Europe in the 16th century.
3. Mid 17th century, divisions were founded in universities that focused entirely on
animal research
4. 19th century, the microscope became commonplace in scientific research; cells
of animals could be studied at the microscopic level.
5. Naturalist, Charles Darwin developed the theory of evolution by natural selection
6. More recently, the discovery of DNA as life’s genetic material led to even more
new research about natural world and the evolutionary relationships between
animals
D. An Evolutionary Perspective
Animals share a common evolutionary past and evolutionary forces that influences their
history. Evolutionary processes are remarkable for their relative simplicity, yet they have
had awesome effect on life-forms. These processes have resulted in an estimated 4 to
30 million species of organisms living today (1.4 are described) many more existed in
the past have become extinct. Zoologists must understand evolutionary processes if
they are to understand what an animal is and how it originated.
Main points:
1. Common evolutionary past and forces unravel family relationships.
2. Evolutionary processes affect life-forms thus explains why animals appear and
function.
Evolution not only explains how the animal functions and forms but also unravel the
historical relationships within the animal kingdom.
The main concept for this point is that groups of individuals which share more genetic
material are more related to each other compared to those which do not share common
genetic material;.
Now let’s look closely at the case of African cichlids to better illustrate this evolutionary
perspective.
Like all organisms, animals are names and classified into a hierarchy of
relatedness. Karl von Linne (1707 – 1778) or Carolus Linnæus, proposed the
Binomial Nomenclature: which includes the genus and the species name of an
organism e.g. e.g. Felis domesticus, Canis familiaris. Organisms in the same
species are more closely related than organism in the same genus and so on.
Remember the taxonomic group: species, genus, family, order, class and so on.
In naming, how much DNA are shared has been aside from variations in traits
are important to include in the classification of an organism. When zoologists
classify animals into taxonomic groupings, they are making hypotheses about the
extent to which groups of animals share DNA, even when they study variations in
traits like color patterns, behaviors, because these kinds of traits ultimately are
based on the genetic material.
Evolutionary concepts hold the key to understanding why animals look and act in
their unique ways, live in their particular geopgraphical regions and habitats, and
share characteristics with other related animals.
Example of Hierarchy of Relatedness:
Figure 1.9. The classification of a housefly, horsefly, cichlid fish and human illustrates
how the classficiation system depicts degree of relatedness.
E. An Ecological Perspective
Ecology (Greek oikos, house and logos, to study) is the study of relationships between
organisms and their environment. Central to this relationship is dependence (or
interdependence) of one animal to another animal or vice-versa. Humans (animal higher
taxa) depend on animals (lower taxa) for food, medicines and clothing. However, human
activities upset the ecological balances.
Let’s take a look at this ecological situation as an example. In the 1950’s, the giant Nile
perch (Lates niloticus) was introduced into Lake Victoria (in Africa) in an attempt to
increase the lake’s fishery, for human consumption. This voracious predator reduced
the cichlid population from 99% to less thatn 1% of the total fish population and has
resulted in the extinction of many cichlid species. Because many of the cichlids fed on
algae, the algae in the lake grew uncontrolled. When algae died and decayed, much of
the lake became depleted of its oxygen. To make matters worse, when Nile perce are
caught, their excessively oily flesh must be dried. Fishermen cut local forests for the
wood needed to smoke the fish. This practice has resulted in severe deforestation
around Lake Victoria. The resulting runoff of soin into the lake has caused further
degradation.
World Population and its Resources
Over all, there are two main problems that are the focus of our ecological concerns: 1)
global overpopulation and 2) the exploitation of world resources.
1) Overpopulation
Global overpopulation is at the root of virtually all other environmental problems.
Human population growth is expected to continue in the the 21 st century. It is estimated
that the world population will reach 10.4 billion by the year 2100. As the human
population grows, the disparity between the wealthiest and the poorest natons is likely
to increase. This will have negative implications in the space we live in, in the resources
we have and the pollution we will cause to the environment.
b)
a)
Summary of Lesson 1.
1. Zoology is the study of animals. A broad field of specializations and
subdisciplines.
2. Animals share a common evolutionary past and evolutionary forces that influence
their history.
3. Evolution explains how the diversity of animals arose.
4. Evolutionary relationships are the basis for the classification of animals into a
hierarchical system. The classification is binomial nomenclature. Higher level of
classification denote more distinct evolutionary relationships
5. All animals share a common environment, and ecological principles help us to
understand how animals interact within that environment
6. Human overpopulation is at the root of virtually all other environmental problems,
stresses world resources and results in pollution, global warming, deforestation,
and the extinction of many plant and animal species.
Assessment:
Short Quiz
Direction. For items 1-4, select the best answer that pertains to the unifying principles in
zoology. For items 5-8, give the correct answer.
1. All life forms descended from a common ancestor through a branching lineage.
2. Large differences in anatomical traits that characterize different species originate
through the accumulation of many small incremental changes over very long
periods of time.
3. The origin and diversification of any taxon or the evolutionary history of its origin
and diversification.
4. Changed in the population of organisms over time.
Options.
Thinking questions:
1. How is zoology related to biology? What major biological concepts, in addition to
evolution and ecology, are unifying principles within the two disciplines?
2. Why should people in all parts of the world be concerned with the extinction of
cichlids in Lake Victoria?
Lesson 2: Levels of Organizations in Animal
Complexity
Objectives:
1. Recall cell as the most basic unit of life
2. Review the functions of the different cell organelles
3. Differentiate the types of animal tissue
4. Define organ and organ systems
Because all animals are made of cells, the cell is as fundamental to an understanding of
zoology.
The cells is the fundamental unit of life, in which all of the chemical reactions necessary
for the maintenance and reproduction of life take place. It is the smallest independent
unit of life. Structurally, it is either prokaryotic or eukaryotic. Discovered fist by Robert
Hooke and was described to be a boxlike cavities in slices of cork and leaves which he
referred calling as “little boxes of cells”.
Prokaryotes:
• “Before nucleus”
• Independent, single-celled organisms (e.g. bacteria)
• Describes cells in which DNA is localized in a region but is not bound by a
membrane
Eukaryotes:
• “true nucleus”
• Have cells with a membrane-bound nucleus containing DNA
• Contain other structures called organelles (“little organs) that perform specific
functions
• Have a network of specialized structures called filaments and tubules organized
into the cytoskeleton (shape + intracellular movements)
Cell Membranes:
1. Plasma membrane surrounds the cell
2. Other membranes inside the cells enclose some organelles and have properties
similar to the plasma membrane
Additional Concepts:
• The ability of the plasma membrane to let some substances in and let others out
is called selective permeability, essential for cellular homeostasis (latin
permeare or per, through + meare, pass)
• Homeostasis is the maintenance of a relatively constant internal environment
despite fluctuations in the external environment (Greek homeo, always the same
+ stasis, standing)
a. Pinocytosis - (“cell drinking” from “pinein” – to drink) when a small portion of the plasma membrane indents
(invaginates) forming a small vesicle and moves into cytoplasm
b. Phagocytosis (“cell eating” from “phagein” to eat) similar to pinocytosis except that the cell takes in solid
material rather than liquid. A lysosome combines with the vesicle to form a phagolysosome
c. Receptor- mediated endocytosis – involves a specific receptor protein on the plasma membrane that recognizes
an extracellular molecule and binds with it. This reaction somehow stimulates the membrane to indent and create
a vesicle containing the selected molecule. A variety of important molecules (such as cholesterol) are brought into
cells in this manner.
7. Exocytosis: Golgi apparatus packages proteins and other molecules that the cell produces into vesicles fro
secretion. In the process of exocytosis, these secretory vesicles fuse with the plasma membrane and release their
contents into the extracellular environment.
Cytoplasm, Organelles and Cellular Ccomponents
1. Cytoplasm
2 parts: a) cytomembrane system – well-defined structures (ER, GA, vacuoles,
vesicles; b) cytosol – fluid suspends the structures of the cytomembrane system and
contains various dissolved molecules
2. Ribosomes
-non-membrane bound structures
-sites for protein synthesis
-contain ~equal amounts of protein and ribosomal RNA (rRNA)
-some attach to the ER; some float freely: cluster in groups connected by a strand of
messenger RNA (mRNA) called polyribosomes or polysomes
3. Endoplasmic Reticulum (ER): production and transport
-complex, membrane-bound labyrinth of flattened sheets, sacs and tubules that
branches and spreads throughout the cytoplasm
-continuous from the nuclear envelope to the plasma membrane
-series of channels that helps various materials to circulate throughout the cytoplasm
-storage unit for enzymes and other proteins, point of attachment for ribosomes
-Two types: a) rough ER: with attached ribosome, and b) smooth ER: site for lipid
production, detoxification of organic mol, storage of Ca ions in muscle cells
4. Golgi Apparatus (GA) or complex: packaging, sorting, export
-named after Camillo Golgi, 1898
-collection of membranes associated physically and functionally with the ER in the
cytoplasm
-flattened stacks of membrane-bound cisternae (closed spaces serving as fluid
reservoirs)
-sorts, packs, secretes proteins and lipids
-e.g. pancreatic cells secreting digestive enzymes, nerve cells: neutotransmitters
5. Lysosomes: Digestion and degradation
-(Greek lyso, dissolving + soma, body)
-membrane-bound spherical organelles that contain enzymes called acid hydrolases,
which digest organic molecules (lipids, proteins, nucleic acids and polysaccharides)
under acidic conditions
6. Mitochondria: power generators, powerhouse of the cell
-double-membrane-bound, spherical to elongate in share
-small space separates the outer membrane from the inner
-inner membrane folds and doubles in on itself to form incomplete partitiosn called
cristae
-cristae increases the SA available for the chemical reactions that trap usable energy for
the cell space between cristae: matrix, containing ribosomes, circular DNA, and others
7. Cytoskeleton: Microtubules, Intermediate Filaments, Microfilaments
-latticed framework extends throughout the cytoplasm, connecting various organelles
and cellular components
a) Microtubules: hollow, slender, cylindrical structures in animals cells
-subunit of globular proteins: tubulin
-Function: 1) movement of organelles, e.g. SV and in chromosome movt during cell
nucleus division 2) part of transport system within the cells e.g. in nerve cells 3) overall
shape changes that cells undergo during periods of specialization
b) Intermediate filaments – chemically heterogeneous group of protein fibers, the
specific proteins of which can vary with cell type
-help maintain cell shape and spatial organization of organelles
-promote mechanical activities within the cytoplasm
c) Microfilaments – solid strings of protein (actin) molecules
-actin: most highly developed in muscle cells as myofibrils helping muscle cells shorten
of contract but in nonmuscle cells, providing mechanical support for cellular structures
and help contractile systems responsible for some cellular movements (e.g. amoeboid
movement)
8. Centrioles and Microtubule-organizing Center
-specialized nonmembranous regions of cytoplasm near the nucleus
-centers of dense material give rise to a large no. of microtubules with different functions
in the cytoskeleton e.g. gives rise to centriole
9. Vacuoles: Cell maintenance
-membranous sacs, part of the cytomembrane system
-various functions, e.g. protozoa and sponges have contractile vacuoles that collect
water and pump it to the outside to maintain the organisms’ internal envt; others for food
storage
10. Nucleus – Information Center
• Control and the information center for the eukaryotic cell
• 2 major functions:
a) directs chemical reactions in cells by transcribing genetic information from DNA into
RNA, which then translates this specific info into proteins (e.g. enzymes) that determine
the cell’s specific activities and
b) stores genetic info and transfers it during cell division from one cell to the next and
from one generation of organisms to the next
Parts/Contents
• Nuclear envelope – membrane separating the nucleus from the cytoplasm that is
continuous with the ER at a no. of points
-nuclear pore: over 3000 for direct contact with the ER; each is composed of an ordered
array of globular and filamentous granules (proteins) which forms the nuclear pore
complex which governs the transport of molecules;
• Chromosomes:
-nucleoplasm: inner mass of the nucleus; contain genetic material, chromatin
-uncoiled chromatin containing hereditary information in segments of DNA called genes
• Nucleolus: preassembly point for ribosomes
-nonmembrane bound structure in the nucleoplasm that is present in nondividing cells
-preassembly points for ribosomes and usually contain proteins and RNA in many
stages of synthesis and assembly
Below are images of the organelles and their parts (Miller-Harley 2001).
Endoplasmic Reticulum
Ribosomes in Rough ER
Smooth ER
Golgi Apparatus
How proteins are transported
from ribosomes to ER to GA
via vesicles
Cytoskeleton – Microtubule,
Intermediate Filament, and
Microfilament
Nucleolus
Tissues
group of similar cells specialized for the performance of a common function
classified into four types:
1. epithelial
2. connective
3. muscle
4. nervous
1. Epithelial Tissue
-many structural forms
-generally, covers or lines something and typically consists of renewable sheets
of cells that have surface specializations adapted for their specific roles
-usually, a basement membrane separates it from underlying adjacent tissues
-Functions: 1) absorb (small intestine lining), 2) transport (kidney tubules) 3)
excrete (sweat glands) 4) protect (skin) and 5) contain nerve cells fro sensory
reception (taste buds) and so the size, shape and arrangement are directly
related
-Classified on the basis of shape and no. of layers:
1) simple: one layer of cells
2) stratified: two or more stacked layers
-individuals cells can be flat, cube or columnlike
2. Connective Tissues
-support and bind
-distributed throughout an extracellular matrix (unlike epithelial) which contains
fibers embedded in a ground substance with a consistency from liquid to solid
-Two General Types:
a) Loose: strong flexible fibers of the protein collagen are interwoven with fine,
elastic, and reticular fibers giving its elastic consistency making it a binding tissue
(e.g. binding the skin to muscle tissue)
b) Fibrous: the collagen fibers are densely packed and lie parallel to one another,
creating a very strong cords such as tendons (connect muscles to bones or to
other muscles) and ligaments (bones to bones)
Specific Examples:
• Adipose Tissue; loose connective tissue that consists of large cells that store
lipids; cells accumulate in large nos. to form fats
• Cartilage: hard yet flexible tissue that supports such structures as the outer ear
and forms the entire skeleton of such as animals as sharks and rays
Chondrocytes – cells lie within spaces call lacunae that are surrounded by a
rubbery matrix that chondroblasts secrete
• Bone cells: (osteocytes) lie within lacunae but the matrix around them is heavily
impregnated with calcium phosphate making it hard suitable for support and
protection
• Blood: a connective tissue in which a fluid called plasma suspends specialized
RBCs and WBCs and platelets; transports various subsstances
3. Muscle Tissue
-for movement
-a contractile process
-three kinds: 1) smooth 2) skeletal 3) cardiac
4. Nervous Tissue
-for communication
-composed of several different types of cells:
1) neurons: impulse-conducting cells: 2) involved with protection, support and
nourishment are called neuroglia; 3) cells that form sheaths and help protect,
nourish and maintain cells of the peripheral nervous system are called
peripheral glial cells
Organs
Functional units of an animal’s body that are made up of more than one type of
tissue
Heart, lungs, liver, spleen, kidneys
Organ Systems
Association of organs that together perform an overall function
The highest organization is the: organismic level
In the higher vertebrate:
a) integumentary
b) skeletal
c) muscular
d) nervous
e) endocrine
f) circulatory
g) lymphatic
h) respiratory
i) digestive
j) urinary
k) reproductive
Cell division and Inheritance
Concepts:
1. The genetic material is organized into chromosomes. Chromosome may be
represented differently in males and females. However, the number of
chromosomes is constant for a given species.
2. Mitosis is the form of cell division that results in growth and repair processes. It
ensures an orderly and accurate distribution of chromosomes during cell cycle.
Cytokinesis results in the division of the cytoplasm.
3. Meiosis is the form of cell division that results in the formation of gametes. It
reduces the chromosome number by half and allows for the random distribution
of one member of each pair of parental chromosomes to the offspring.
4. Deoxyribonucleic acid (DNA is the genetic material of the cell. Its double helix
structure suggests how it can replicate itself and how it can code for the
sequences of amino acids that make proteins.
5. Proteins synthesis involves two processes. Transcription is the production of a
messenger RNA (mRNA) molecule that is complementary to a gene in DNA.
Translation is the assembly of proteins at the ribosomes based on the genetic
information in the transcribed mRNA.
6. Changes in DNA and chromosomes increase the variation within a species and
account for evolutionary process.
7. Principles of classical genetics explain the inheritance patterns of many animal
traits, including dominance, segregation, and independent assortment.
8. Many alternative forms of a gene may exist in a population, and these alternative
forms may interact in different ways.
9. Patterns of inheritance observed at an organismal level are explained at a
molecular level by the presence or absence of functional enzymes.
Assessment:
Short Quiz: (link for the google form will be given in case)
Direction: Fill in the blanks for the movement of molecules through the cytomembrane
system. (2 points each item).
1. _________________________ that ribosomes synthesize are sealed off in
little packets called 2. _____________________ . 3. __________________ pass from
the 4. __________________ , and then to the 5. ___________________ and fuse with
it. In GA, proteins are concentrated and 6. _____________________ . Proteins then are
packaged into 7. __________________________ and released into the 8. __________
close to the plasma membrane. Secretory vesicles fuse with 9. _______________ and
release contents to the outside via 10. __________________________ .
Homework:
Topic: Organ Systems
Direction: In the google classroom, you will be assigned an individual work. You will be
given a specific set of organ system and discuss it with the following guide questions.
Format is TNR, 12, A4-size, 1-inch margin. Write your name, subject and section, and
date submitted.
1. Describe concisely the overall function of the system. File name should be as
follows: HW1_LastName_OrganSystems
2. Enumerate the major parts of the system.
3. Give the main function of each part in relation to the overall function of the
system. If there are other functions being performed, identify them.
4. Explain the interconnectedness of the system to other systems, if any.
Lesson 3: Evolution: A Historical Perspective
and the Modern Synthesis
Objectives:
1. Define evolution
2. Discuss pre-Darwininan, Darwinian and modern synthesis
General Description/Definition:
Organic Evolution
introduced by Charles Darwin
refer to as “descent with modification” which means species change over time
evolution by itself does not imply any particular lineage or any particular
mechanism, evidence for change in organisms over long time periods is
overwhelming
natural selection is the mechanism for evolution
Figure 3.1. Geographical distribution of lions. a. African lion (Panthera leo) has a similar ecological role
as b. mountain lion (Felis concolor) of north and south America. Their similar form suggests a distant
common ancestry. Obvious differences, however result from millions of years of independent evolution.
c. distribution of lions found in Africa and North, South America.
Theories of Evolution
1. Pre-darwinian
2. Lamarck
3. Charles Darwin
4. Alfred Russel Wallace
5. Modern Synthesis
1. Pre-darwinian –
Ancient Greeks and philosphers
Empedocles (495 – 435 B.C.) and Aristotle (384 – 322 B.C.): concepts of
change in living organisms over time
Georges-Louis Buffon (1707 – 1788) studied comparative anatomy
Erasmus Darwin (1731 – 1802), physician who questions origin and
change; believed in the common ancestry of all organisms
2. Lamark –
Jean Baptise Lamarck, French zoologist
Animal Classification
Theory of how change occurs: “species are not constant and that existing
species were derived from preexisting species”
1800’s, Theory of Inheritance of Acquired Characteristics- use of an organ
resulted in that organ becoming highly developed and that disuse resulted
to degeneration
Now, found to be an incorrect theory since neck elongation did not
originate as changes in the genetic material
Figure 3.8. Voyage of the HMS Beagle. Darwin went to the Galapagos Island where he
observed animals and soon proposed the theory of evolution.
Galapagos Island as basis for his theory of evolution by natural selection
The inset shows two of the islands 1) Ferdinandina and Isabela (Wolf,
Darwin, Alcedo)
He spent 5 weeks on the Galapagos Islands, a group od volcanic islands
900 km off the coast of Ecuador
“I shall always maintain [the theory of evolution by natural selection] to be actually yours
and yours only. You had worked it out in details I had never thought of years before I
had a ray of light on the subject.”
-Wallace to Darwin
5. Modern Synthesis –
A. Hardy-Weinberg Theorem
B. Evolutionary Mechanism
C. Species and Speciation
D. Rates of Evolution
E. Molecular Evolution
F. Mosaic Evolution
*Notice that the definition of organic evolution has changed from the Darwinian theory.
In modern synthesis,
Understanding genetics: why variations exist and how to pass on
Genetic variation: may confer advantage to individuals
Crossing-over, multiple alleles and mutations add to variations
No two individuals are genetically identical except for twins
Chance combinations of genes
A. Hardy-Weinberg Theorem
1908, Godfrey H. Hardy and Wilhelm Weinberg
The mixing of alleles at meiosis and their subsequent recombination do not alter
the frequencies of the alleles in future generations, if certain assumptions are
met
Assumptions:
1. The population size must be large.
2. Mating within the population must be random.
3. Individuals cannot migrate into, or out of, the population.
4. Mutations must occur.
B. Evolutionary Mechanisms:
Four Types: genetic drift, gene flow, mutation, and selection
Evolution is a result of some individuals in a population surviving and being more
effective at reproducing than others in a population, leading to changes in gene
frequencies.
Smaller the population, more significant chance there may be,
2. Gene Flow/Migration
• Immigration or emigration results in changes in gene frequencies
• Can decrease genetic variation in a pop by working towards same direction as
other evolutionary forces
• Can increase variation if it puts back in the pop the genes because of other
processes
Caveats:
1. Some organisms do not reproduce sexually (e.g. microbes)
2. When applied to fossil material (dead organisms)
3. Populations of similar organisms may be so isolated from each other the gene
exchange is geographically impossible
Allopatric Speciation
occurs when subpopulations become geographically isolated from one another
Parapatric Speciation
occurs in small, local populations called demes
E.g. all of the frogs in a particular pond or all of the sea urchins in a particular
tidepool make up a deme
Individuals of a deme a re more likely to breed with one another than with others
Same envt, same pressure
Demes – not completely isolated from each other
Relative isolation means its members experience different selection pressures
than other members of a population
Sympatric Speciation
occurs within a single population in the same area
-though organisms are sympatric, they still may be reproductively isolated from
one another.
-plant species producing viable forms with multiple sets of chromosomes
-uncommon to animals but some species of bats and insect and fish display this
D. Rates of Evolution
1) Phyletic gradualism- evolutionary change as occurring gradually over millions
of years which traditionally interprets the tempo or rate of evolution
Stasis (or equilibrium)- studies of fossil record show that many species do not
change significantly over millions of years
2) Punctuated equilibrium- long periods of stasis interrupted by brief periods of
change
E. Molecular Evolution
Evolutionary relationships among organisms by studying DNA and protein e.g.
cytochrome c
Gene Duplication- accidental duplication of a gene on a chromosome; one way
that extra gene material can arise e.g. vertebrate hemoglobin (carries O2) and
myoglobin (O2 storage molecule in muscle cells)
F. Mosaic Evolution
A species is a mosaic of different molecules and structures that have evolved at
different rates.
Some are conserved; Some changed rapidl e.g. design of a bird (feathers, bills,
body form);
Particular parts are less conservative and have a higher rate of change e.g.
wings have been modified for hovering, soaring and swimming while legs for
wading, swimming and perching
Assessment:
Short Quiz:
1. What is the general definition of evolution in Darwinian era?
2. In modern synthesis, define evolution.
3. Enumerate disciplines as evidences for evolution.
4-11 Identify which Evolutionary/genetic mechanism each item refers to.
4. range of mechanisms
5. could occur at the gametic or zygotic levels
6. can decrease genetic variation in a population as well as increase that variation if
genes where put back
7. Dunkers
8. cytosine changes into guanine in a gene segment
9. the case of cheetah population
10. affects the reproductive success of a genotype
11. What is a species?
Lesson 4: Ecology: Preserving the Animal
Kingdom
Objectives:
1. Cite some interactions prevailing between animals and its abiotic
environment
2. Define population
3. Enumerate interspecific interactions among animals
4. Give the unique attributes of communities
5. Demonstrate actions to resolve ecological problems
Content:
Habitat
Refers to the environment including all living and nonliving characteristics of the
area in which the animal lives
Abiotic- oxygen, inorganic ions, light, T, current, wind velocity
Tolerance range - certain range of values for any envt’l factor that animals can
thrive
range of optimum- a certain range of values within the tolerance range where the
animal is most successful
Figure 4.1. Tolerance
range of an animal
Energy
Refers to the ability to do work (foraging, cellular activities)
Heterotrophic- animals ingesting other animals
Autotrophic- photosynthesizing or other carbon-fixing activities that supply their
own food source
Temperature
An animal expends part of its existence energy in regulating body temperature
Temperature influences the rates of chemical reactions in animal cells (metabolic
rate) and affects animal’s overall activity
Not constant due to heat loss and gain
Torpor- time of decreased metabolism and lowered body T that occurs in bats,
hummingbirds who must feed constantly when they are active
Hibernation- time of decreased metabolism and lowered body T that may last for
weeks or months. e.g. rodents, shreds, bats
Winter sleep- in larger animals; large energy are reserved through periods of
winter inactivity
Aestivation- period of inactivity that must withstand an extended periods of drying
(e.g. invertebrates, reptiles amphibians)
B. Populations
-defined as the group of individuals of the same species that occupy a given area at the
same time and have unique attributes
Population Growth
Change over time as a result of birth, death and dispersal
Survivorship curves- characterize a population based in death of individuals
Population growth- the potential for a population to increase in numbers of
individuals in an exponential growth
Four factors influencing reproductive potential
1. Number of offspring produced
2. Likelihood of survival to reproductive age
3. Duration of the reproductive period
4. Length of time to maturity
Environmental resistance – refers to the constraints that climate, food, space and other
environmental factors place on a population
Population Regulation
- to regulate the population, its density is considered
Factors:
1. Density-independent factors- influence the no. of animals in a population without
regard to the no. of individuals per unit space (density)
2. Density-dependent factors- more severe when population density is high than
they are at other densities
Intraspecific Competition
Competition occurs when animals utilize similar resources and in some way
interfere with each other’s procurement of those resources
Intraspecific competition refers to competition among members of the same
species; often intense
C. Interspecific Competition
Example:
D. Communities
- defines as different populations of species living in an area
Community Stability
• Succession- a process in which the dominant members of a community often
change a community in predictable ways
• Pioneer community- first community to become established in an area. Death,
decay and additional nutrients add to the community.
• Seral stage- each successional stage (over thousands of years nutrients
accumulate, and the characteristics of the ecosystem change)
• Sere- entire successional sequence
• Climax community- final community which can tolerate its own reaction and
usually have complex structure and high species diversity
Important concepts:
• Food Chain- sequence of organisms through which energy moves in an
ecosystem
• Food webs- complexly interconnected food chains
• Trophic levels- groupings of organisms according to the form of energy used
• Keystone species- the very important species in the food web that when
depleted, all the other animals will collapse
Food Webs
• Producers (autotrophs)- obtain nutrition (complex organic compounds) from
inorganic materials and an energy source
• Consumers (heterotrophs)-eat other organisms to obtain energy
• Herbivores (10 consumers)-eat producers
• Carnivores (20 consumers) - eat herbivores and eat other carnivores (30
consumers)
• Decomposers-break down dead organisms and feces by digesting organic matter
extracellularly and absorbing products of digestion
Figure 6.5. Flow of energy through ecosystems. 1.5 million kcal of radiant
energy strike a square meter of the earth’s surface each year. Plants convert
less than 1% into chemical energy. Of this 60% is converted into a new
biomass and 40% is lost in respiration.
Figure 6.6. The Antarctic Food Web. An Antarctic food web: small crustaceans called krill support nearly
all life in Antarctic. 6 species of baleen whales, 20 species of squid, 100 species of fish, 35 species of
birds and 7 species of seals eat krill. Krill feed on algae, protozoa, other small crustaceans and various
larvae.
Cycling Within Ecosystems
• Unlike energy, all matter is cycled from nonliving reservoirs to living systems and
then back to nonliving reservoirs
• Matter is constantly recycled within ecosystems
Biogeochemical Cycles
• Nutrient- any element essential for life
• 97% of living matter is made of C,H,O,N (often in gaseous cycles); but carbon is
most abundant
• S, P, Ca – less abundant, equally important
• Nonliving reservoir: in the earth stored as coal, gas and oil
1. Sedimentary cycles- cycles involving these elements in the earth only
2. Hydrological cycle- water cycles through ecosystems
3. Carbon Cycle – movement and conversion and sequestration of Carbon into
the earth, water and atmosphere
But let’s only discuss the Carbon Cycle in detail:
• Carbon- plentiful in the earth and rarely a limiting factor
• Carbon dioxide- reservoir for carbon in the atmosphere or water
• Carbonic acid in the waters and calcium carbonate (from the shells of molluscs
and skeletons of echinoderms) in the waters
• Calcium carbonate deposits results to limestone formation. Geologic uplifts,
volcanic eruptions and weathering return this carbon back to the earth’s surface
and atmosphere
• Coal, oil and gas- in the earth
• Other carbon are tied up in fossil fuels. Burning fossil fuels returns large
quantities of this carbon to the atmosphere as carbon dioxide
E. Ecological Problems
1. Human overpopulation is the root of all environmental problems. The earth
has a carrying capacity and a limited supply of resources. When human
populations achieve that carrying capacity, populations should stabilize.
Otherwise, war, famine, and/disease will occur.
Assessment:
No quiz.
Unit test: Not Applicable since two major exams will be given. The first major exam will
be given as a midterm exam.
References:
• Hickman, Cleveland P., et al. 1988. Integrated Principles of Zoolgy. Quezon City:
Copyright by Times Mirror/ Mosby College Publishing and Printed by JMC Press,
Inc.
• Miller, Kenneth R. and Levine, joseph S. 2006. Prentice Hall Biology. Boston,
Massachusetts: Pearson Education, Inc., publishing as Pearson Prentice Hall.
• Miller, Stephen A. and John P. Harley. 2001. Zoology, Fifth Edition. The
McGraw-Hall Companies.
Extra Credit:
Students can earn extra points in two ways:
1. Define and present in class a not-yet-discussed zoological term pertaining to a
body part significant to any animal.
Example:
Proboscis –
A proboscis refers to an elongated or projecting appendage in the head region of
certain animals. In invertebrates, such as certain insects, annelids, mollusks, and
ribbon worms, the proboscis is a hollow tubular cephalic appendage. In
butterflies, in particular, their proboscis is connected to the oral region. Their
proboscis, therefore, serves as a feeding or a sucking organ. In certain
vertebrates, the proboscis is not attached to the mouth but may be formed as a
fusion of the nose and the upper lip (as in snouts). Common examples of
vertebrate proboscises are the elephant trunk and the elongated nose of the
proboscis monkey.
References:
Molecular Expressions: Science, Optics & You – Olympus MIC-D: Darkfield
Gallery – Butterfly Proboscis. (2019). Fsu.Edu.
https://micro.magnet.fsu.edu/optics/olympusmicd/galleries/darkfield/butterflyprob
oscis.html
Zenker, M. M., Penz, C., Paris, M. de, & Specht, A. (2011). Proboscis
Morphology and Its Relationship to Feeding Habits in Noctuid Moths. Journal of
Insect Science, 11(42), 1–10. https://doi.org/10.1673/031.011.0142