Professional Documents
Culture Documents
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Engineering
Fundamentals
An Introduction to Engineering
6th Edition SI Edition
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Engineering
Fundamentals
An Introduction to Engineering
6th Edition SI Edition
Saeed Moaveni
Minnesota State University, Mankato
Professor Emeritus
of Mechanical Engineering
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Important Notice: Media content referenced within the product description or the product
text may not be available in the eBook version.
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Engineering Fundamentals: An Introduction © 2020, 2016, 2011 Cengage Learning, Inc.
to Engineering, Sixth Edition, SI Edition Unless otherwise noted, all content is © Cengage
Saeed Moaveni WCN: 02-300
ALL RIGHTS RESERVED. No part of this work covered by the copyright
Product Director, Global Engineering: herein may be reproduced or distributed in any form or by any means,
Timothy L. Anderson except as permitted by U.S. copyright law, without the prior written
Senior Product Assistant: Alexander Sham permission of the copyright owner.
Cengage
20 Channel Center Street
Boston, MA 02210
USA
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Contents
Preface xiii
Preface to the SI Edition xxi
Digital Resources xxii
PART 1
Engineering 2
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vi Contents
3.3 Teamwork 85
3.4 Project Scheduling and the Task Chart 87
Before You Go On 89
3.5 Engineering Standards and Codes 89
3.6 Water and Air Standards in the United States 95
Before You Go On 100
Professional Profile 101
Summary 102
Key Terms 103
Apply What You Have Learned 103
Problems 103
Impromptu Design II 106
Civil Engineering Design Process 106
Mechanical/Electrical Engineering Design Process 110
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Contents vii
PART 2
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viii Contents
Summary 278
Key Terms 279
Apply What You Have Learned 279
Problems 280
11 Temperature and T
emperature-Related Variables in Engineering 368
11.1 Temperature as a Fundamental Dimension 371
11.2 Temperature Difference and Heat Transfer 381
Before You Go On 396
11.3 Thermal Comfort 396
11.4 Heating Values of Fuels 399
Before You Go On 401
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Contents ix
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x Contents
PART 3
PART 4
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Contents xi
Summary 667
Key Terms 668
Apply What You Have Learned 668
Problems 669
An Engineering Marvel 678
PART 5
PART 6
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xii Contents
Appendix 849
Index 855
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Preface
2. Discussion Starter
Pertinent articles serve as chapter openers to engage students and promote
active learning. The discussion starters provide a current context for why
the content that the students are about to learn is important. An instructor
can start class by asking students to read the Discussion Starter and then
ask the students for their thoughts and reactions.
3. Before You Go On
This feature encourages students to test their comprehension and under-
standing of the material discussed in section(s) by answering questions
before they continue to the next section(s).
Vocabulary—It is important for students to understand that they need to
develop a comprehensive vocabulary to communicate effectively as well-
educated engineers and intelligent citizens. This feature promotes growing
vocabulary by asking students to state the meaning of new words that are
covered in section(s).
xiii
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xiv P r e fa c e
5. Summary
Each chapter concludes by summarizing what the student should have
gained from studying the chapter. Moreover, the learning objectives and
the summary are tied together as a refresher for the students.
6. Key Terms
The key terms are indexed at the end of each chapter so that students may
return to them for review.
Organization
This book is organized into six parts and twenty chapters. Each chapter begins
by stating its objectives and concludes by summarizing what the student
should have gained from studying the chapter. I have included enough mate-
rial for two semester-long courses. The reason for this approach is to give the
instructor sufficient materials and the flexibility to choose specific topics to
meet his or her needs. Relevant, everyday examples with which students can
easily associate are provided in every chapter. Each chapter includes many
hands-on problems, requiring the student to gather and analyze information.
Moreover, information collection and proper use of information are encour-
aged in this book by asking students to complete a number of assignments
that require information gathering by using the Internet as well as by employ-
ing traditional methods. Many of the problems require students to make brief
reports so that they learn that successful engineers need to have good written
and oral communication skills. To emphasize the importance of teamwork
in engineering and to encourage group participation, many of the assign-
ment problems require group work; some require the participation of the
entire class.
The main parts of the book are:
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P r e fa c e xv
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xvi P r e fa c e
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P r e fa c e xvii
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xviii P r e fa c e
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P r e fa c e xix
Impromptu Designs
I have included seven inexpensive impromptu designs that could be developed
during class times. The basic ideas behind some of the impromptu designs have
come from the ASME.
References
In writing this book, several engineering books, websites, and other materi-
als were consulted. Rather than giving you a list that contains hundreds of
resources, I cite some of the sources that I believe may be useful to you. All
freshmen engineering students should own a reference handbook in their
chosen field. Currently, there are many engineering handbooks available in
print or electronic format, including chemical engineering handbooks, civil
engineering handbooks, electrical and electronic engineering handbooks, and
mechanical engineering handbooks. I also believe all engineering students
should own chemistry, physics, and mathematics handbooks. These texts
can serve as supplementary resources for all your classes. Many engineers
may find useful the ASHRAE handbook, the Fundamental Volume, by the
American Society of Heating, Refrigerating, and Air Conditioning Engineers.
In this book, some data and diagrams were adapted with permission from
the following sources:
• Baumeister, T., et al., Mark’s Handbook, 8th ed., McGraw Hill, 1978.
• Electrical Wiring, 2nd ed., AA VIM, 1981.
• Electric Motors, 5th ed., AA VIM, 1982.
• Gere, J. M., Mechanics of Materials, 6th ed., Thomson, 2004.
• Hibbler, R. C., Mechanics of Materials, 6th ed., Pearson Prentice Hall.
• U.S. Standard Atmosphere, Washington D.C., U.S. Government Printing
Office, 1962.
• Weston, K. C., Energy Conversion, West Publishing, 1992.
Instructor Resources
Lecture Note PowerPoints and the full Instructor’s Solutions Manual are
available online via a secure, password-protected Instructor’s Resource Center
at http://login.cengage.com.
Acknowledgments
I would like to express my sincere gratitude to the editorial and produc-
tion team at Cengage, especially Timothy Anderson and Alexander Sham.
I am also grateful to Rose Kernan of RPK Editorial Services, Inc. I would
also like to thank Dr. Karen Chou of Northwestern University, Mr. James
Panko, and Paulsen Architects, who provided the section on civil e ngineering
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xx P r e fa c e
design process and the related design case study, and the late Mr. Pete Kjeer
and Johnson Outdoors, who provided the mechanical/electrical engineer-
ing case study. I am also thankful to all the reviewers who offered general
and specific comments including Whitney Blackburn-Lynch, University of
Kentucky, Lance Crimm, Kennesaw State University, Ali Eydgahi, Eastern
Michigan University, David Feinauer, Norwich University, Sungwon Kim,
Minnesota State University—Mankato, Jennifer O’Neil, Rochester Institute
of Technology, Farhad Reza, Minnesota State University—Mankato, Abiye
Seifu, Columbus State University, and Chengyang Wang, Montgomery County
Community College.
I would also like to thank the following individuals for graciously providing
their insights for our Student and Professional Profiles sections: Nahid Afsari,
Jerry Antonio, Celeste Baine, Suzelle Barrington, Steve Chapman, Karen
Chou, Ming Dong, Duncan Glover, Dominique Green, Lauren Heine, John
Mann, Katie McCullough, Susan Thomas, and Nika Zolfaghari.
Thank you for considering this book, and I hope you enjoy the Sixth
Edition.
Saeed Moaveni
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Preface to the SI Edition
xxi
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Digital Resources
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Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it.
D i g i ta l R e s o u r c e s xxiii
MindTap Reader
Available via WebAssign, MindTap Reader is Cengage’s next-generation
eBook for engineering students.
The MindTap Reader provides more than just text learning for the student.
It offers a variety of tools to help our future engineers learn chapter concepts
in a way that resonates with their workflow and learning styles.
• Personalize their experience
Within the MindTap Reader, students can highlight key concepts, add notes,
and bookmark pages. These are collected in My Notes, ensuring they will have
their own study guide when it comes time to study for exams.
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xxiv D i g i ta l R e s o u r c e s
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Engineering
Fundamentals
An Introduction to Engineering
6th Edition SI Edition
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Engineering
An Exciting Profession
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PART
1
I
n Part One of this book, we will introduce you to the engineering profession.
Engineers are problem solvers. They have a good grasp of fundamental physi-
cal and chemical laws and mathematics and apply these laws and principles to
design, develop, test, and supervise the manufacture of millions of products and ser-
vices. Engineers, regardless of their background, follow certain steps when designing
the products and services we use in our everyday lives. Successful engineers possess
good communication skills and are team players. Ethics plays a very important role in
engineering. As eloquently stated by the National Society of Professional Engineers
(NSPE) code of ethics, “Engineering is an important and learned profession. As mem-
bers of this profession, engineers are expected to exhibit the high-
Good engineers are problem est standards of honesty and integrity. Engineering has a direct
solvers and have a firm grasp and vital impact on the quality of life for all people. Accordingly,
of mathematical, physical, and the services provided by engineers require honesty, impartiality,
chemical laws and principles. They fairness and equity, and must be dedicated to the protection of
apply these laws and principles to the public health, safety, and welfare. Engineers must perform
design products and services that under a standard of professional behavior which requires adher-
we use in our everyday lives. They ence to the highest principles of ethical conduct.” In the next five
also have good written and oral chapters, we will introduce you to the engineering profession,
communication skills. how to prepare for an exciting engineering career, the design
process, engineering communication, and ethics.
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C HAPTER
1
Introduction to the Engineering
Profession
Learning Objectives
LO1 Engineering Work is All Around You: give examples
of products and services that engineers design that
make our lives better
LO2 Engineering as a Profession: describe what engi-
neers do and give examples of common careers for
engineers
LO3 Common Traits of Good Engineers: describe the
important traits of successful engineers
LO4 Engineering Disciplines: give examples of common
engineering disciplines and how they contribute to
the comfort and betterment of our everyday lives
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Discussion Starter 5
Engineers are problem solvers. Successful engineers develop, test, and supervise the manufacture of millions
possess good communication skills and are team play- of products and services. They consider important fac-
ers. They have a good grasp of fundamental physical tors such as efficiency, cost, reliability, and safety when
and chemical laws and mathematics. Engineers apply designing products. Good engineers are dedicated to
physical and chemical laws and mathematics to design, lifelong learning and service to others.
Discussion Starter
Who are Engineers?
W
e all want to make the world a better Engineers are problem solvers. They have a
place, but how do we do it, and where good grasp of fundamental physical and chemi-
do we start? Leo Tolstoy, a Russian nov- cal laws and mathematics and apply these laws
elist and philosopher, once said: and principles to design, develop, test, and
supervise the manufacture of millions of prod-
Everyone thinks of changing the world, but no
ucts and services. Engineers, regardless of their
one thinks of changing oneself.
background, follow certain steps when designing
Increasingly, because of worldwide socioeco- the products and services we use in our everyday
nomic trends, environmental concerns, and the lives. Successful engineers possess good commu-
Earth’s finite resources, more is expected of all of nication skills and are team players.
us. As future engineers, you are expected to con-
sider the link among the Earth’s finite resources
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and environmental, social, ethical, technical,
and economical factors as you make decisions
regarding the services that you provide and the
products you design. This book is designed to
introduce you—someone who is interested in
studying engineering regardless of your area
of specialization, personal interests, and future
career path—to important concepts that every
engineer should know.
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6 CHAPTER 1 Introduction to the Engineering Profession
Ethics plays a very important role in engi- all people. Accordingly, the services provided
neering. As eloquently stated by the National by engineers require honesty, impartiality,
Society of Professional Engineers (NSPE) code of fairness and equity, and must be dedicated to
ethics, “Engineering is an important and learned the protection of the public health, safety and
profession. As members of this profession, engi- welfare. Engineers must perform under a stan-
neers are expected to exhibit the highest stan- dard of professional behavior which requires
dards of honesty and integrity. Engineering has adherence to the highest principles of ethical
a direct and vital impact on the quality of life for conduct.”
To the Students: Why do you want to study engineering? Name at least two products
or services that are not available now that you think will be readily available in the
next 20 years. Which engineering disciplines do you think will be involved in design
and development of these products and services?
P
ossibly some of you are not yet certain you want to study engineering
during the next four years in college and may have questions similar to
the following:
Do I really want to study engineering?
What is engineering and what do engineers do?
What are some of the areas of specialization in engineering?
How many different engineering disciplines are there?
Do I want to become a mechanical engineer, or should I pursue civil engi-
neering? Or would I be happier becoming an electrical engineer?
How will I know that I have picked the best field for me?
Will the demand for my area of specialization be high when I graduate,
and beyond that?
The main objectives of this chapter are to provide some answers to these
and other questions you may have and to introduce you to the engineering
profession and its various branches.
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1.1 Engineering Work Is All Around You 7
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Figure 1.1 Examples of products and services designed by engineers.
get your morning news or to see how your favorite team did—was designed by
electrical and electronic engineers. There are, of course, engineers from other
disciplines involved in creating the final product; for example, manufacturing
and industrial engineers. When you are getting ready to take your morning
shower, the clean water you are about to use is coming to your home thanks
to civil engineers. Even if you live out in the country on a farm, the pump you
use to bring water from the well to your home was designed by mechanical
and civil engineers. The water could be heated by natural gas that is brought
to your home thanks to the work and effort of chemical, mechanical, civil, and
petroleum engineers. After your morning shower, when you get ready to dry
yourself with a towel, think about what types of engineers worked behind the
scenes to produce the towels. Yes, the cotton towel was made with the help of
agricultural, industrial, manufacturing, chemical, petroleum, civil, and mechan-
ical engineers. Think about the machines that were used to plant and pick the
cotton, transport the cotton to a factory, clean it, and dye it to a pretty color
that is pleasing to your eyes. Then other machines were used to weave the fab-
ric and send it to sewing machines that were designed by mechanical engineers.
The same is true of the clothing you are about to wear. Your clothing may con-
tain some polyester, which was made possible with the aid of petroleum and
chemical engineers. “Well,” you may say, “I can at least sit down and eat my
breakfast and not wonder whether some engineers made this possible as well.”
But the food you are about to eat was made with the help and collaboration
of various engineering disciplines, from agricultural to mechanical. Let’s say
you are about to have some cereal. The milk was kept fresh in your refrigera-
tor thanks to the efforts and work of mechanical engineers who designed the
refrigerator components and chemical engineers who investigated alternative
refrigerant fluids with appropriate thermal and other environmentally friendly
properties that can be used in your refrigerator. Furthermore, electrical engi-
neers designed the control and the electrical power units.
Now you are ready to get into your car, take the bus, or ride the subway to
go to school. The car you are about to drive was made possible with the help
and collaboration of automotive, mechanical, electrical, electronic, industrial,
material, chemical, and petroleum engineers. So, you see there is not much that
you do in your daily life that has not involved the work of engineers. Be proud
of the decision you have made to become an engineer. Soon you will become
one of those whose behind-the-scenes efforts will be taken for granted by bil-
lions of people around the world. But you will accept that fact gladly, knowing
that what you do will make people’s lives better.
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8 CHAPTER 1 Introduction to the Engineering Profession
We Need Clean Air Every day, human activities through stationary and mobile
sources contribute to the pollution of outdoor air. Power plants, factories, and dry
cleaners are examples of stationary sources that create outdoor air pollution. The
mobile sources of air pollution, such as cars, buses, trucks, planes, and trains, also
add to the level of outdoor air pollution. In addition to these man-made sources,
natural air pollution also occurs due to forest fires, windblown dust, and volcanic
eruptions. Moreover, because most of us spend approximately 90 percent of our
time indoors, the indoor air quality is also very important to our short-term and
long-term health. In recent years, we have been using more synthetic materials in
newly built homes that can give off harmful vapors. We also use more chemical
pollutants, such as pesticides and household cleaners.
We Need Clean Water Our next essential need is water. Droughts are good
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1.1 Engineering Work Is All Around You 9
TAB L E 1 . 1 Estimated Calorie Needs per Day by Age, Gender, and Physical Activity Level.
Estimated amounts of caloriesa needed to maintain calorie balance for various gender and age groups at three
different levels of physical activity. The estimates are rounded to the nearest 200 calories for assignment to a
USDA food pattern. An individual’s calorie needs may be higher or lower than these average estimates.
Male Female
Activity Moderately Moderately
Levelb (Age) Sedentary Active Active Sedentary Active Active
TAB L E 1 . 1 Estimated Calorie Needs per Day by Age, Gender, and Physical Activity
Level. (continued)
Male Female
Activity Moderately Moderately
Levelb (Age) Sedentary Active Active Sedentary Active Active
Not all Americans are able to follow this dietary guideline. Here are some
facts about American calorie imbalance that are worth noting:
• According to the USDA Economic Research Service, in recent years,
nearly 15 percent of American households have been unable to get
enough food to meet their daily Calorie needs.
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1.1 Engineering Work Is All Around You 11
• At the other end of the spectrum, many Americans (among all subgroups
of the population) are overweight or obese because their daily Calorie
intake exceeds their activity level needs.
• In an article entitled “U.S. Lets 141 Trillion Calories of Food Go to
Waste Each Year,” Eliza Braclay writes that “The sheer volume of
food wasted in the U.S. each year should cause us some shame, given
how many people are hungry both in our own backyard and abroad.”
This is happening in America, while 1 in 9 people in the world (data
from World Food Programme Organization) do not have enough
food to lead a normal life. The 141 trillion Calories that represent
approximately 1,250 Calories per person per day in the United States
is the result of nearly 59 billion kg of food that was lost. Moreover,
it is important to understand that the wasted food is worth over
$100 billion. According to the USDA, the top three food groups lost
(in a recent year) were dairy products (11.3 billion kg, or 19 percent
of all the lost food); vegetables (11.3 billion kg, or 19 percent); and
grain products (8.4 billion kg, or 14 percent). The USDA’s Economic
Research Service also points out that, if we were to reduce this waste,
the price of food worldwide might go down. In addition, most of us
do not realize that a vast amount of energy is spent in the food supply
chain, and when food is wasted, valuable resources such as water and
fossil fuels (that go into growing, processing, and transporting the food)
are also wasted.
Have you ever thought about how much energy it takes to feed you every
day? Let’s go back to the simple example we used earlier—having some cereal
for breakfast. Now think about what it takes to grow a cereal crop such as corn
and wheat (incidentally, corn, wheat, and soybeans make up the majority of
field crop inputs to the U.S. food supply). Think about the energy that needs
to be spent to plant the seeds, make and apply fertilizers, irrigate the field, har-
vest the crop, and finally transport it to a processing plant. Next consider how
much energy it takes to process the corn into the cereal, make plastic bags and
attractive boxes to contain it, and deliver the cereal boxes to the supermarkets.
Moreover, we like to have some milk with our cereal. Well, that is going to
require additional energy to make the cattle feed, run the milking machines,
produce milk containers, and build and operate refrigerated trucks. Then after
the milk gets to the supermarket, it needs to be placed in cold storage, requir-
ing energy to maintain its low temperature. After you bring the milk home,
you need to store it in a refrigerator, which also consumes energy. You get the
picture! Now, think about all of the other food and drinks that you consume
in a single day.
Now let’s consider what happens to a dollar spent on food. According
to USDA, for a typical dollar spent (in a recent year) by U.S. consumers on
domestically produced food (including both grocery store and eating-out pur-
chases), 12.1 cents went to agribusiness and farm production, 15.8 cents to
food processing, 2.7 cents to packaging, 3.3 cents to transportation, 9.3 cents to
a wholesale trader, 13 cents to the food retailer, 31.1 cents to services provided
by food service establishments, 5.6 cents to energy costs, 3.3 cents to finance
and insurance costs, and 3.8 cents to pay for activities such as advertising, legal,
and accounting services (see Figure 1.2).
As you can see, in the United States, if you were to buy a loaf of bread,
say for one dollar, approximately 12 cents goes to the actual cost of the flour
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12 CHAPTER 1 Introduction to the Engineering Profession
e
nc
ura
ing
on
de
ion
s
cti
Ins
tra
ss
ce
pro ss
du
e
Wh or tat
ce
e
i
le
ns ng
ad
Ot ce &
erv
Fa usin
pro
sa
Tra gi
l tr
Fin y
s
p
ole
cka
erg
r
an
r ib
rm
tai
od
od
he
Re
En
Ag
Pa
Fo
Fo
¢
¢
3.3¢
¢
¢
1¢
¢
¢
¢
2.4
9.7
.8
2.7
9.3
13
5.6
3.3
3.8
31.
15
and the remaining 88 cents goes to paying for processing, packaging, trans-
portation, advertisement, and so on. But if the price of wheat doubles from
12 cents to 24 cents—assuming no changes in other costs—the bread will cost
only an additional 12 cents, which is an increase of 12 percent in total cost
(incidentally, in the United States, we spend approximately 12.9 percent of
our income on food). The share of U.S. household consumer expenditures by
major categories for year 2013 is shown in Figure 1.3. In contrast, in develop-
ing countries, poor people may spend as much as 80 percent of their income
on food.
Other Education,
Apparel, 3.1% 3.1% reading, 2.4%
Savings, 3.6%
Entertainment, alcoholic
beverages, 5.8%
Housing
Health care, 7.1% 33.6%
Personal
insurance,
pensions Food Transportation
10.8% 12.9% 17.6%
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1.1 Engineering Work Is All Around You 13
35%
<5% 5–14, 9% 15–24, 9% 25–34, 9%
and over
Very low Moderately low Moderately high High Very high Missing
or insufficient data
It costs as little
as US 25 cents a day
to feed a hungry child
and change her life
forever.
Together we are
About 795 million making progress
people, or 1 in 9 of the in reducing the number
world’s population, of hungry people in the
go to bed hungry world. We can achieve
each night. zero hunger in our
lifetimes.
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14 CHAPTER 1 Introduction to the Engineering Profession
RECHARGEABLE BATTERIES:
Lithium metallic oxide
Based on EPA
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1.1 Engineering Work Is All Around You 15
Evan Lorne/Shutterstock.com
In addition, we need various modes of transportation to get to different
places, because we may live and work in different cities or wish to visit friends
and relatives who may live elsewhere. We also like to have some sense of secu-
rity, to be able to relax, and to be entertained. We need to be liked and appreci-
ated by our friends and family, as well.
Increasingly, because of worldwide socioeconomic population trends, envi-
ronmental concerns, and the earth’s finite resources, more is expected of engi-
neers. Future engineers are expected to provide goods and services that increase
the standard of living and advance health care while also addressing serious envi-
ronmental and sustainability concerns. At the turn of the 21st century, there were
approximately six billion of us inhabiting the earth. As a means of comparison,
it is important to note that the world population about 120 years ago, at the start
of the 20th century, was one billion. Think about it. It took us since the beginning
of human existence to reach a population of one billion. It only took 120 years to
increase the population by sixfold. Some of us have a good standard of living, but
some of us living in developing countries do not. You will probably agree that our
world would be a better place if every one of us had enough to eat, a comfort-
able and safe place to live, meaningful work to do, and some time for relaxation.
According to the latest estimates and projections of the U.S. Census Bureau,
the world population will reach 9.3 billion people by the year 2050. Not only will
the number of people inhabiting the earth continue to rise, but the age structure
of the world population will also change. The world’s elderly population—the
people at least 65 years of age—will more than double in the next 25 years (see
Figure 1.5).
How is this information relevant? Well, now that you have decided to study
to become an engineer, you need to realize that what you do as an engineer is
very important to all of us. The world’s current economic development is not
sustainable—the world population already uses approximately 20% more of
the world’s resources than the planet can sustain. (United Nations Millenium
Ecosystem Assessment Synthesis Report, 2005.) You will design products and
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Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it.
16 CHAPTER 1 Introduction to the Engineering Profession
10
9
Population (billions) 8
7
6
5
4
3
2
1
0
1950 1960 1970 1980 1990 2000 2010 2020 2030 2040 2050
Year
(a)
1990 37,307
Year
2050 834,000*
(b)
Figure 1.5 ( a) The latest projection of world population growth. (b) The latest estimate of U.S.
elderly population growth.
Data from the U.S. Census Bureau
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1.2 Engineering as a Profession 17
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18 CHAPTER 1 Introduction to the Engineering Profession
risteski goce/Shutterstock.com
Oleksiy Mark/
Shutterstock.com
Yulia Nikulyasha Nikitina/Shutterstock.com
BassKwong/Shutterstock.com
DM7/Shutterstock.com
Figure 1.6 s an engineer, you will apply physical and chemical laws and principles and
A
mathematics to design various products and services.
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Another random document with
no related content on Scribd:
(above), boundary between pairing animals; (below), first fission; single
vertical line, continuity or enlargement. M, Meganucleus; µ, micronucleus;
Z, zygote-nucleus.
In the Peritrichaceae the mates are unequal; the larger is the normal
cell, and is fixed; the smaller, mobile, is derived from an ordinary
individual by brood-divisions, which only occur under the conditions
that induce conjugation (Fig. 60). Here, though the two pairs of
nuclei are formed, it is only the migratory nuclei that unite, the
stationary ones aborting in both mates. During the final processes of
conjugation the smaller mate is absorbed into the body of the larger,
and so plays the part of male there. But this process, though one of
true binary sex, is clearly derived from the peculiar type of equal
reciprocal conjugation of the other Infusoria.
The Ciliata are almost all free-swimming animals with the exception
of most of the Peritrichaceae, and of the genera we now cite.
Folliculina forms a sessile tube open at either end; and Schizotricha
socialis inhabits the open mouths of a branching gelatinous tubular
stem, obviously secreted by the hinder end of the animal, and forking
at each fission to receive the produce. A similar habit to the latter
characterises Maryna socialis; all three species are marine, and
were described by Gruber.[168] Stentor habitually attaches itself by
processes recalling pseudopodia, and often forms a gelatinous
sheath.
Acineta, Ehrb. (Fig. 61, 2); Amoebophrya, Koppen; Choanophrya, Hartog (Fig.
62); Dendrocometes, St. (Fig. 61, 4); Dendrosoma, Ehrb. (Fig. 61, 9);
Endosphaera, Engelm.; Ephelota, Str. Wright (Fig. 61, 5, 8); Hypocoma,
Gruber; Ophryodendron, Cl. and L. (Fig. 61, 7); Podophrya, Ehrb. (Fig. 61, 1);
Rhyncheta, Zenker (Fig. 61, 3); Sphaerophrya, Cl. and L. (Fig. 61, 6),
Suctorella, Frenzel; Tokophrya, Bütschli.
BY
CHAPTER VII
PORIFERA (SPONGES)[185]
INTRODUCTION—HISTORY—DESCRIPTION OF H A L I C H O N D R I A
P A N I C E A AS AN EXAMPLE OF BRITISH MARINE SPONGES AND OF
E P H Y D A T I A F L U V I A T I L I S FROM FRESH WATER—DEFINITION—
POSITION IN THE ANIMAL KINGDOM.
The familiar bath sponge was naturally the earliest known member of
the phylum. It is dignified by mention in the Iliad and in the Odyssey,
and Homer, in his choice of the adjective "full of holes," πολύτρητος,
shows at least as much observation as many a naturalist of the
sixteenth and seventeenth centuries. Aristotle based his ideas of
sponges entirely upon the characters of the bath sponge and its near
allies, for these were the only kinds he knew. With his usual
perspicuity he reached the conclusion that sponges are animals,
though showing points of likeness to plants.
It was not till 1825 that attention was again turned to the current,
when Robert Grant approached the group in a truly scientific
manner, and was ably supported by Lieberkühn. It would be
impossible to do justice to Grant in the brief summary to which we
must limit ourselves. The most important of his contributions was the
discovery that water enters the sponge by small apertures scattered
over the surface, and leaves it at certain larger holes, always
pursuing a fixed course. He made a few rough experiments to
estimate the approximate strength of the current, and, though he
failed to detect its cause, he supposed that it was probably due to
ciliary action. Grant's suggestion was afterwards substantiated by
Dujardin (1838), Carter (1847), Dobie (1852), and Lieberkühn
(1857). These five succeeded in establishing the claims of sponges
to a place in the animal kingdom, claims which were still further
confirmed when James-Clark[190] detected the presence of the
protoplasmic collar of the flagellated cells (see pp. 171, 176). Data
were now wanted on which to base an opinion as to the position of
sponges within the animal kingdom. In 1878 Schulze[191] furnished
valuable embryological facts, in a description agreeing with an earlier
one of Metschnikoff's, of the amphiblastula larva (p. 226) and its
metamorphosis. Then Bütschli[192] (1884) and Sollas[193] on
combined morphological and embryological evidence (1884)
concluded that sponges were remote from all the Metazoa, showing
bonds only with Choanoflagellate Protozoa (p. 121). This the exact
embryological work of Maas, Minchin, and Delage has done much to
prove, but it has to be admitted that unanimity on the exact position
of the phylum has not yet been attained, some authorities, such as
Haeckel, Schulze, and Maas still wishing to include sponges in the
Metazoa.
Halichondria panicea.
One of the commonest of British sponges, which may be picked up
on almost any of our beaches, and which has also a cosmopolitan
distribution, is known by the clumsy popular name of the "crumb of
bread sponge," alluding to its consistency; or by the above technical
name, with which even more serious fault may be found.[194]
Bidder has proposed to call the different forms of the same species
"metamps" of the species. Figures of the metamps of H. panicea will
be found in Bowerbank's useful Monograph.[195]
Sections show that the ostia lead into spaces below the thin
superficial layer or "dermal membrane"; these are continued down
into the deeper parts of the sponge as the "incurrent canals,"
irregular winding passages of lumen continually diminishing as they
descend. They all sooner or later open by numerous small pores
—"prosopyles"—into certain subspherical sacs termed flagellated
chambers. Each chamber discharges by one wide aperture
—"apopyle"—into an "excurrent canal." This latter is only
distinguishable from an incurrent canal by the difference in its mode
of communication with the chambers.
Fig. 64.—H. panicea: the arrows indicate the direction of the current, which is
made visible by coloured particles. (After Grant.)
The excurrent canals convey to the osculum the water which has
passed through the ostia and chambers. All the peripheral parts of
the sponge from which chambers are absent are termed the
"ectosome," while the chamber-bearing regions are the
"choanosome."