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1894.
SCORPIONS.
355
2A 2
356 NATURAL SCIENCE. MAY,
side ; but in some of the Buthidae there are as m a n y as five, and the
fact that these five m a y be o b s e r v e d in genera that are not especially
related points to the conclusion that the ancestors of this family
possessed this number ; and this conclusion seems to be borne out by
the c i r c u m s t a n c e that the nearest living a l l y of the S c o r p i o n s , namely,
Thelyphonus, which we believe to be the most p r i m i t i v e of all the
1
E p e c t i n a t e A r a c h n i d a , m a y also present five lateral ocelli on each
side.
T h e conclusion that the Buthidae are, on the w h o l e , the most
a r c h a i c of living Scorpions is borne out by a s t u d y of the g e o g r a p h i c a l
distribution of these a n i m a l s ; for, as will be seen in the sequel, this
family is represented by peculiar genera in all the quarters of the
globe w h e r e S c o r p i o n s exist.
T h e a b s e n c e of proofs of their great antiquity, which w a s referred
to in connection w i t h the inter-relationships of the e x i s t i n g species of
S c o r p i o n s , is also r e v e a l e d no less r e m a r k a b l y by a study of their
g e o g r a p h i c a l distribution ; for in this latter respect t h e y show con-
siderable r e s e m b l a n c e to such a r e l a t i v e l y recent g r o u p as the
Mammalia. No doubt our k n o w l e d g e of the recent species of this
latter class is v e r y much more complete than is our k n o w l e d g e of the
A r t h r o p o d s in question ; but so far as we h a v e been able to ascertain,
W a l l a c e ' s m a p of the Z o o l o g i c a l R e g i o n s of the w o r l d , if we eliminate
the northern parts of the so-called Palaearctic and N e a r c t i c R e g i o n s
w h e r e S c o r p i o n s do not occur, is closely a p p l i c a b l e to this group of
animals. W e m a y , therefore, without further words o f preface,
proceed to discuss the c h a r a c t e r i s t i c features of the v a r i o u s regions.
In the Palaearctic R e g i o n the northern limit of S c o r p i o n s in
E u r o p e and W e s t e r n A s i a a p p e a r s in a general w a y to be defined by
the mountain-chain of w h i c h the P y r e n e e s is the western and the
C a u c a s u s the eastern e x t r e m i t y . E a s t of the C a s p i a n they h a v e not
been recorded north of the 40th parallel, but they are k n o w n to occur
at t w o places upon this line, n a m e l y , at K a s h g a r in T u r k e s t a n , and
at P e k i n in C h i n a . T h e t w o species from these localities are
nearly allied to each other, and they u n m i s t a k a b l y belong to the
M e d i t e r r a n e a n section of the g e n u s Buthus. F u r t h e r m o r e , a species of
Buthus, with distinct Palaearctic affinities, h a s recently been recorded
from S h a n g h a i . C o n s e q u e n t l y , we must conclude that the Medi-
terranean fauna extends from S p a i n and A l g e r i a to the coast of
C h i n a , with the probable omission of the table-land of T i b e t . The
southern limit of this area in C h i n a is, for w a n t of d a t a , uncertain ;
but since the S c o r p i o n s of B u r m a , A s s a m , and N o r t h I n d i a are totally
different from those of T u r k e s t a n , A f g h a n i s t a n , and P e r s i a , we m a y
1
Since, as Mr. Laurie has pointed out, the term Ctenophora, which I have
recently assigned to the Scorpions as opposed to all other Arachnida which were
called Lipoctena, is generally recognised as signifying a group of Ccelenterata,
I wish to propose the terms Pectinifera and Epectinata as substitutes for the
above.
1S94. SCORPIONS. 357