Bol. R. Soco Esp. Rist. Nat. (Sec. Biol.), 88 (1-4), 1992, 79-98.

A preliminary key to the World genera of the subfamilies

Toxotarsinae, Chrysomyinae and Rhiniinae (Diptera,
Calliphoridae)
Claves preliminares para los géneros de las subfamilias Toxotarsinae,
Chrysomyinae y Rhiniinae (Diptera, Calliphoridae) en el mundo
Salvador V. Peris (*)

KEY wüRDS: Calliphoridae: Toxolarsinae, Chrysomyiinae, Rhiniinae. Genera World Fauna.

PALABRAS CLAVE: Calliphoridae: Toxolarsinae, Chrysomyiinae, Rhiniinae. Géneros del mundo.

ABSTRACT

The genera of the Calliphoríd subfamilies Toxotarsinae, Chrysomyinae and Rhiniinae are keyed and
listed. The World Fauna is covered.

RESUMEN

Los géneros de las subfamilias de Califóridos Toxotarsinae, Chrysomyinae y Rhiniinae se incluyen

en una clave y en un listado. Se añaden las citas mundiales de estas subfamilias.

l. INTRODUCTlON

The family Calliphoridae have been the object of two papers on their general
classification (SEGUY, 1935; LEHRER, 1970). Neither of them have had wide ac­
ceptance. Probably the time is not yet ripe for that, the present day set-up is a rather
conservative one, and far more knowledge is required of early stages and biology
before a sound classification be possible. This paper does not pretend to be dealing
with classification but rather with identification.
To make clear my intention, in this and other papers that may appear adopting
similar heading, it is just to make a path towards generic distinction of adult flies.
1 know that perhaps it would have been better to make specific revisions of different
genera and only latter prepare the generic key; this is what 1 did, with Rhiniinae in

(*) Departamento de Biología Animal 1 (Zoología-Entomología). Facultad de Biología. Universidad

Complutense. 28040 Madrid.

Bol. R. Soco Esp. Hist. Nat. (Sec. Biol.), 88 (1-4). 1992.

80 S. V. PERIS

a former paper (PERIS, 1952). But doing so is a time-consuming effort, and take a
necessary time is not available at presento Given the opportunity to study big collec­
tions from all parts of the world for a rather long period of time this task may be
achived. My intention is far more simple and this may explain the word «prelimi­
nary» of the heading.
In the following keys all type-species of genera ineluded have been seen, unless
specifically stated other wise. It is on the characters shown by them that key is manu­
factured, but in large genera other species have been taken into consideration as far
a possible, and availability allows.
The intention has been to inelude as many groups proposed as possible instead
of diminishing them, that is by using small genera instead of large ones. With this
principIe lumping is always possible if desired, but the world fauna must be considered
before doing so, as only then are the limits of genera in their proper perspective.
The results here given are the ones that seemed to me more reasonable, but 1
am sure that future knowledge, or particular personal tastes of researches, will change
the outlines of several groups. Personally, and for biogeographical reasons and com­
parisons, 1 prefer reasonably restricted groups.
The key is followed by an alphabeticallist of genera considered, including their
type-species nominations and reasons for it and also synonymies. Names in synonymy
are included under the valid adopted names.
A few abbreviations are used, but they are so simple that atable for them seems
useless. However, if necessary it may be found in PERIS (1952) and GONZÁLEZ-MoRA
& PERIS (1988).

2. KEY TO SUBFAMILIES OF CALLlPHORIDAE

Keys to recognize the high rank groups were published elsewhere (PERIS, 1952;
GROSSKEY, 1965; MARILUIS & PERIS, 1984) but they are also presented here for the
sake of completeness.
Mesembrinellinae is considered a family on its own following GUIMARAES (1977).
A key to most species have already been published (PERIS & MARILUIS, 1984; MA­
RILUIS, 1987). The group is not considered here.

1 (2) Metathoracic spiraele with a single large lappet with a dorsal opening. Remi­
gium (stem-vein) setulose or bare aboye. Vein m evenly rounded bend. Female
postabdomen not forming telescopic ovipositor. Exclusively Neotropical.. ..
............................................................................. Mesembrinellinae
2 (1) Metathoracic spiraele with the usual structure in two lappets.
3 (8) Remigium setulose aboye.
4 (3) Remigium setulose on both sides, dorsal and ventral (this last character absent
in several Toxotarsus with almost bare arista). Arista plumosa, pilose or almost
bare, Body metallic coloured. Hind coxae pilose behind. Occiput without bare
surface on upper parto First ventrite with borders overlapping the correspond­
ing tergites. Species fram South America, Los Andes regions or influenced areas
................................................................................... Toxotarsinae

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A PRELIMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 81

5 (4) Remigium only setulose on dorsal parto

6 (7) First ventrite not overlapping lateral edges of the corresponding tergites, at least
posteriorly. Upper occiput without abare non-pollinose area. AII other char­
acters vary according to tribe (see later) Chrysomyinae
7 (6) First ventrite wide, shield forming, with borders overlapping lateral edges of
corresponding tergites. Occiput with upper area not pollinose and rather con­
cave. Greater ampullae not pilose, with only microscopic tomentum. Only two
sternopleurals. Thoracic squamae bare dorsally. AII warm regions of Old World
and Australia, not in America (only one species introduced in Bermuda)
....................................................................................... Rhiniinae
8 (3) Remigium not setulose aboye (with arare exception in Pollenia, which has black
ground colour). Subalar bullae always bare Other groups

2.1. Subfamily Toxotarsinae

A group endemic to the Andes region of South America. DEAR (1979) studied
them in an excelent paper, but his classification was modified by MARILUIS & PE­
RIS (1984) after observations of living specimens in the field. The key pressented is
based on this last paper, where is also a key to species.

2.1.1. Key to genera

1 (8) prst aer bristles presento Third antennal segment longer than second.
2 (3) Usually big flies, 12 to 16 mm in length. Male basitarsus thickened apically.
Cerci fused and paralobi reduced. Male body and legs with dense and long pile.
Female with fifth tergite when viewed in profile concave and with strong bristles.
Alar squamae brown with pile of sorne colour, the thoracic squamae also brown
with marginal pile white. Sternopleurals O + 1, 1 + 1 or 2 + 1. aer setae
present but with variability Neta
3 (2) Usually smaller flies, 7 to 11 mm in length. Male basitarsus normal. Cerci not
fused and paralobi of usual size. Body and legs no long pile. Female fith tergite
not as described aboye. Squamae with no such colouration.
4 (7) post aer bristles presento Sternopleurals 1 + 1, or 2 + 1.
prst is present near the suture. Both sexes with a sub median v seta on mid-tibia.
5 (6) post aer placed at equallevel as corresponding de. Thoracic squamae strongly
darkened. Arista plumose on at least basal half, the longest hairs longer than
width of third antennal joint. Vestiture of presutural area of notum wavy, more
so in males Sareonesiopsis
6 (5) post aer setas at different level of the de setas. Thoracic squamae white or
yellowish. Arista plumose on less then basal half, the largest hairs not exceed­
ing the width of third antennal joint. Presutural vestiture bristly, straight. .
........................................................................... Chlorobraehycoma
7 (4) post acr absent. Sternopleurals 1 + 1. prst ia absent. Mid-tibia with a subme­
dian v bristle in females, lacking in males. Arista plumose on basal half only,
the longest hairs subequal widht of third antennal joint. Sareonesia

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82 s. V. PERIS

°
8 (1) No acr present. Sternopleurals + 1. Arista bare or only short-pilosa, plumose
or pectinated. Squamae wellowish white. Third antennal, segment subequal to
the second Toxotaraus
a (d) Remigium ventrally bare. Arist practically bare or short-pectinated. Legs
dark, brawn or black.
b (c) Thoracic squama not lobulated, diverging fram thorax. Ariste with pile
shorter then maximum width of arista. R5 open at wing margin.......
......................................................................... .Kuschelomyia
c (b) Thoracic squama lobulated, not diverging fram thorax. Pile of arista
longer than maximum width of ariste. R5 closed at wing border or shortly
petiolate Toxotarsus S.str.
d (a) Remigium setulose ventrally. Pile of arista as long as the maximum arista!
width. Leges yellow. Thoracie squama not lobulate. R5 open at border
......................................................................... Callyntropyga

2.1.2. List of genera

Chlorobrachycoma TOWNSEND, 1918, Ins. Ins. Menstr. 6: 155 (Type-species: Chlo­
robrachycoma splendida TOWNSEND, 1918, orig. design.)
= Sarconesiomima LOPES & ALBUQUERQUE, 1955, Rev. Chi1. Ent. 4: 104 (Type­
speeies: Sarconesiomima bicolor LOPES & ALBUQUERQUE, 1955, orig. designo
= Sarcophaga dichroa SCHINER, 1868)
= Sarconesia LOPES & ALBUQUERQUE, 1915 Le.: 67 (type-speeies: Sarconesia versi­
color BIGOT, 1857, orig. design.)
Neta SHANNON, 1926, Proc. Entomol. Soco Wash. 28: 123 (Type-species: Phrysso­
poda splendens MACQUART, 1851, orig. designo = Musca chilensis WALKER,
1837)
= Callyntropus ENDERLE1N, 1830, Dtsch. Entomol. Zts. 20: 70 (Type-species:
Phryssopoda splendens MACQUART, 1851, orig. designo (=Musca chilensis WAL­
KER, 1837)
Sarconesia B1GOT, 1859, Ann. Soco Entomol. Fr. 6: 301 (Type-speeies: Sarcophaga
chlorogaster WIEDEMANN, 1830, designo SHANNON, 1926.
Sarconesiopsis TOWNSEND, 1918 Ins. Ins. Menstr. 6: 155 (Type-species: Sarconesiop­
sis caerulea TOWNSEND, 1918, orig. designo (= Calliphora magellanica LE OU1­
LLOU, 1842).
= Rorairomusca TOWNSEND, 1935, Rev. Entomol. Rio S: 59 (Type-speeies: Rorai­
romusca roraima TOWNSEND, 1935, orig. design.)
Toxotarsus MACQUART, 1851, Mem. Soco Scí. Agr. Lille (1850) 211 (Type-species:
Toxotarsus rufipa/pis MACQUART, 1851, monotypy.
= Trixoneura SHANNON, 1925, Proc. Entomol. Soco Wash. 28: 121 (Type-speeies:
Agria fuscipennis MACQUART, 1843, orig. designo (= Sarcophaga nigrocyanea
WALKER, 1837).
= Callyntropyga ENDERLEIN, 1940, Nat. Hist. Juan Fernandez (Zool.) 3 (5): 644.
(Type-species: Callyntropyga se/kirki ENDERLE1N, 1948, orig. designo
(= Stomoxys humeralis WALKER, 1837).
= Kuschelomyia LOPES, 1961, Rev. Brasil. Biol. 21: 455 (Type-species: Kuschelom­
yia ambrosiana LOPES, 1961, orig. design.)

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A PRELIMINARY KEY TO THE WORLD GENERA üF THE SUBFAMILIES OF CALLIPHORIDAE 83

3. SUBFAMILY CHRYSOMYINAE

The group is not easy to delimitate, except using negative characters. Only two
of them seem to apply and those are in the key; all others may appears in sorne genera
but are lacking in others. These flies inhabit all the world over but the bulk of them
are in the tropical areas.
It is customary to divide them into two groups, which are given the rank of tribes.
Both are also difficult to separate by well clear cut characters, but biogeographical
distribution may be a help and so is included in the following key.

3.1. Key fo tribes

1 (2) Hind coxas without hairs posteriorly. Species of dark colours (except Phormia
and Protophormia bright metallic), dark green or blue and violet metallic or
wholly black, with head and legs also black or dark ground colour. Mesonotum
usually without vittae. Greater empullae bare. Thoracic squama setulose aboye
or not setulose. Northern species, in the Nearctic reaching south as far as Me­
xico, D.F., in the Palaearctic from Boreal to the Mediterranean and Japan
...................................................................................... Phormiini

2 (1) Hind coxae with posterior hairs aboye the femur articulation. Flies usually
brightly coloured, metallic green or blue, often with golden or copper reflec­
tions. Sometimes the legs yellow, and mesonotum with distinct vittae. Greater
ampullae setulose or pilose in the Old World representatives, but not so in the
native genera of the Southern Continents (Australia and South America), ex­
ception are the species of Chrysomya present everywhere. The members of this
tribe inhabit all warm regions of the World, especially Tropical.. .
........ .. . . Chrysomyini

4. TRIBE CHRYSOMYINI

In this tribe difficulties of nomenclature have appeared in sorne Neotropical

genera. These have been dealt with by DEAR (1985: 118) who reached sorne con­
clusions that are here followed (see original paper for details). AIso the proposals
made by PONT (1983) to the ICZN have been incorporated.
The largest genus is Chrysomya, which is also the one which has accumulated
a larger synonymy list. This is due to the variety of its members and the vast area
in which they are living (recently extended to South America). BEZZI (1927) dealt
with sorne of them, considered by him as different groupings, mainly as subgenera,
and erected Achoetandrus for albiceps: Psi/ostoma was for him also a subgenus on
account of its testaceous legs and part of abdomen. Microcalliphora was accepted
as a valid genus. This classification has not obtained much acceptance. SEGUY (1928)
presented a key to aggregates inside Chrysomya based on the phallus structure, but
not with great succes either. TOWNSEND (1935) also admitted a number of different
genera, mostly not accepted by subsequent authors. Certainly there is something of
true in SEGUY'S arrangement. The structure of the phallus in albiceps-rufifades (and

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84 S. V. PERIS

whatever the status of both will be) is certainly very different from that of regalis
or chloropyga. However the name Chrysomya as nominating aH these species has
gained wide acceptance and here is kept in this traditional sense.
Microcalliphora has been situated in the key as a group. ActuaHy it is so, but
I mention it specially due to the fact that it sometimes appears as a genus in literature,
and with this I try to indicate that it is scarcely separable. Therefore Chrysomya main­
tains aH the synonymy.
This genus Chrysomya has recently a special atention among American and
European for its rapid invasion of that continent (see PERIS, 1987; BAUMGARTNER,
1988, for further references).

4.1. Key to genera

In all of them the propleura, prosternum and pteropleura are pilose.

(4) Greater ampullae covered with long, wristly hairs or largely pilose. TypicaHy
forms from Old World, although Chrysomya has invaded almost all South and
North America.
2 (3) acr, ia, ph and prst setae present and weH distinct. AH warm zones of the world
...................................................... ............................... Chrysomya
a (b) Eyes of male close each other, front almost linear aboye. Size from normal
(that of a common fly) to big Chrysomya S.str.
b (a) Eyes of male widely separated, frons alleast half the eye width, very little
more in female. SmaH species, about 5 mm length. From South Pacific
.............................................................. Microcalliphora-group
3 (2) acr, ia, ph and prst setae not differentiated from surrounding vestiture. Scutel­
lum with six long and 1-3 discals. Greater ampullae with long pale pile. In late­
ral view head with three dark spots: at lower part of parafrontal; middle of
parafacial: and anteriorly on the occipital dilatation. Parafacials black setulose
aboye, in lower part with white setulae. Thoracic squama lobulated. Only from
Madagascar Chrysopyrellia
4 (1) Greater ampullae bare or only short pilose, no as aboye.
5 (6) General colour testaceous on pleural parts, legs and base of abdomen: the
mesonotum, scutellum and last abdominal segments dark with blue reflections,
on the abdomen this dark colour extending lateraHy and also forming a median
longitudinalline. Dorsum not vittate. Thoracic squamae somewhat lobulate,
although posteriorly rounded, entirely pilose aboye. Greater ampullae bare,
only with microscopic tomentum. Australia Eucompsomyia
6 (5) Not so. General body colour metallic, although parts of it may be yellow, includ­
ing the legs. Thorax often with a clear vittated pattern. Thoracic squama bare
or setulose only at its base, behind the alar squama, lobulated or not. Genera
typically from South America, sorne reaching the U .S.A.
7 (8) Palpi short and reduced, filiform, not reaching the epistomal border. Parafacials
setulose. Thoracic squamae lobulate. prst acr and ia absent. ... Cochliomyia
8 (7) Palpi normal, club-shaped and clearly reaching the epistomal border.

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A PRELIMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 85

9 (10) Parafacial setulose. prst dc absent. Thoracic squama pilose on basal dorsal
surface. Occipital dilatation entirely yellow pilose Compsomyiops
10 (9) Parafacialia bare. prst dc and ia presento Thoracic squama not lobulate. Oc­
cipital dilatation with golden yellow pile and usually black setulose on its
upper anterior part.
11 (12) Thoracic squama with dorsal basal setulae. Wings without middle dark
suffusions, only the basal parte of wing brownish. Abdomen dark, the same
colour as the rest of the body. Most exterior ph seta presento Sternopleurals
2 + 1. 2 prst acr present. Para/ucilia
12 (11) Thoracic squama dorsally bare. Wings with dark parts, not hyaline. Sterno­
pleurals I + 1. Most exterior ph seta presento
13 (14) Wings with anterior suffusions, one from half the sc cell to the and of r/
vein, another about the apical half or r2-r3 vein. Abdomen with sorne yello­
wish colour on its basal part. Prosternum golden pilose. At least 2 post acr.
Thoracic squama diverging from middle line of body. Male eyes with unequal
facets Hemilucilia
14 (13) Wings uniformly grey brown more so along costal border. Abdomen with
basal part brown, other segments like thorax. Prosternum pile gold brown.
Only one post acr distinct. Thoracic squamae with posterior margin rounded
and subparallel to thorax. Males usually with large facets on antero-upper
part even often separated from the smaller ones by a line situated at the level
of the lunula Ch/oroprocta

4.2. List of genera

Ch/oroprocta WULP, 1896, Bio/. C. Amer., Dipt. 2: 296 (Type-species: Ch/. semivi­
ridis WULP, 1896, monotypy (= Chrysomyia idoidea ROBINEAU-DESVOIDY,
1830).
= Ca/litrogopsis TOWNSEND, 1935, Rev. Entomo/. Rio S: 70 (Type-species: Calli­
trogopsis dorsalis TOWNSEND, 1835, orig. designo (= Chrysomyia idioidea
ROBINEAU-DESVOIDY, 1830).
Chrysomya ROBINEAU-DESVOIDY, 1830 Myod.: 244 (Type-species: Chrysomya re­
galis ROBINEAU-DESVOIDY, 1830, designo COQUILLETT, 1910; placed in Offi­
cial List, see NATSUCK & al. Bull. Zoo/. Nom. 42 (2): 393-394.
= Chrysomyia AGASSIZ, 1847, Nomencl. Zoo/.: 8S, emend. pro Chrysomya.
= Compsomyia RONDANI, 1875, Ann. Mus. Genova 7: 425 (Type-species: Musca dux
ESCHCHOLTZ, 1822, designo COQUILLETT, 1910 (=Musca megacepha/a FABRI­
CIUS, 1794).
=Pycnosoma BRAUER & BERGENSSAMM, 1894, Denkschr. Akad. Wiss. Wien S6:
623 (Type-species: Musca marginalis WIEDEMANN, 1830, orig. design.)
(= Chrysomya regalis ROBINEAU-DESVOIDY, 1830).
=Paracompsomyia HOUGH, 1898, Proc. Acad. Nat. Sci. Philadelphia 50: 184 (Type­
species: Paracompsomyia nigripennis HOUGH, 1898, monotypy, = Chrysomya
regalis ROBINEAU-DESVOIDY, 1830).

Bol. R. Soco Esp. Hist. Nat. (Sec. Biol.), 88 (1-4), 1992.

----~-------- ~------------
86 S. V. PERIS

= Microcalliphora TOWNSEND, 1916, Proc. U.S. Nat. Mus. 49: 618 (Type-species:
Lucilia varipes MACQUART, 1851, orig. design.)
= Achoetandrus BEZZI, 1927 (sbg. Chrysomyia), Bull. Entomol. Res. 17: 235 (Type­
species: Musca albiceps WIEDEMANN, 1819, orig. design.)
=Psilostoma SURCOUF, 1919, Arch. Mus. Hist. Nat. Paris (5) 6 (1914): 58 (Type­
species: Ochromyia incisuralis MACQUART, 1850, monotypy).
= Hemichrysomyia SÉGUY, 1926, Bull. Soco Entomol. Fr. (1926): 303 (Type-species:
Ochromyia incisuralis MACQUART, 1850, monotypy).
=Somomyia SÉGUY, 1927, E. E. Dipt. 4: 8 (nec RONDANI, 1841 nec BERTOLONI,
1861 (Type-species: see PONT, 1983, Bull. Zool. Nom. 40: 110).
= Cyaneosomyia SÉGUY, 1928, E.E. Dipt. 4: 112 (Type-species: Cyaneosomyia phao­
nis SÉGUY, 1928, orig. design.)
= Pycnosomops TOWNSEND, 1934, Ent. News 45: 277 (Type-species: Musca puto­
ria WIEDEMANN, 1830, orig. designo (=Musca chlorophyga WIEDEMANN, 1818).
= Ceylonomyia FAN, 1965, Key Comm. Synan. Flies China: 196 (Type-species:
Chrysomyia (Microcalliphora) nigripes AUBERTIN, 1932, orig. design.)
Chrysopyrellia SÉGUY, 1920, E. E. Dipt. 4: 3 (Type-species: Chrysopyrellia saph­
yrea SÉGUY, 1927, orig. design.)
Cochliomyia TOWNSEND, 1915, J. Wash. Acad, Sci. 5: 646 (Type-species: Musca ma­
cellaria FABRICIUS, 1775, orig. design.)
= Callitroga BRAUER, 1883, Denskchr. Akad. Wiss. Wien 47: 274 (Type-species:
Musca macellaria FABRICIUS, 1775, orig. designo (Supressed genus: MELVILLE,
1984 Bull. Zool. Nom. 41 (2): 128).
=Protochrysomyia PIERCE, 1945, Bull. S. Calif. Acad. Sci. 44: 8; Pleistocene CA.
(Type-species: Protochrysomyia howardae PIERCE, 1945, orig. design.)
Compsomyiops TOWNSEND, 1918, /ns. /ns. Menstr. 6: 153 (Type-species: Callipho­
ra fulvipes MACQUART, 1843, orig. designo (= Chrysomya fulvicrura ROBINEAU­
DESVOIDY, 1830).
= Myiolucilia HALL, 1948, Blowfl. N. Amer.: 109 (Type-species: Musca lyrcea WAL­
KER, 1849, orig. designo (= Chrysomya fulvipes ROBINEAU-DESVOIDY, 1830).
Eucompsomyia MALLOCH, 1927, Pric. Linn. Soco N. W.S. 52: 325 (Type-species:
Eucompsomyia semimetallica MALLOCH, 1927, orig. design.)
Hemilucilia BRAUER, 1895, Sto Wiss. Akad. Wien. 104, Abt.l: 598 (Type-species:
Musca segmentaria FABRICIUS, 1805, orig. design.)
=Mya RONDANI, 1850, Nuov. Ann. Sc. Nat. Bologna 2 (3): 175 (Type-species: Mya
semidiaphana RONDANI, 1850, designo PONT, 1983), preocc. Mya LiNNAEUS,
1758).
Paralucilia BRAUER & BERGENSTAMM, 1891, Denschr. Akad. Wiss. Wien 58: 391
(87) (Type-species: Calliphora fulvipes MACQUART, 1843, sensu BLANCHARD
nec MACQUART, misidentif. (= Somomyia fulvinota DIGOT, 1877)).

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A PRELlMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILlES OF CALLIPHORlDAE 87

5. TRlBE PHORMIlNI

An exclusively Holarctic group. Mainly zoosaprophagous, but sorne are myiasic,

either sanguivorous in bird nestlings (Proloealliphora) or subcutaneous (Trypoea­
lliphora). Only six genera are known which may be distinguished by the following
key, mainly traslation from GONZÁLEZ-MoRA & PERIS (1966).

5.1. Key lo genera

1 (8) Arista rather long plumose, its longest rays subequal to or exceeding the width
of the third antennal segment. Anterior spiracles of ussual size, i.e. similar to
those shown in Calliphora. Females with pollinose parafrontals.
2 (5) Thoracic squama white or black setulose (requires careful observation). Pro­
pleural depression setulose. prsl acr weak, compared with the de and hardly
differentiated from surrounding vestiture. Saprophagous forms.
3 (4) Thoracic squama internally lobulated, white and usually finely white setulose,
aer short but usually distinct, 3-4 + 2-3, only the prse strong. Anterior thoracic
spiracle pale orange-yellow, contrasting with surrounding areas of thorax. Pa­
rafacial setulae not descending further the level of half the length of third an­
tennal segment. Thoracic notum slightly convex before scutellum and with fi­
ne white grey pollinosity but not forming vittae in front of suture. Body meta­
llic green in colour Phormia
4 (3) Thoracic squama not lobulated, its internal border diverging from thorax, dark
and black setulose. acr 0-1 + 0-2, all as fine as surrounding vestiture. 2-3 +
1 de bristles, Anterior thoracic spiracle black-brown, not contrasting with sur­
roundings areas of thorax. Dorsum of thorax with extremely sparse pollinosity,
not forming vittae. Parafacial setulae reaching (frontal view) the level of half
the length of third antennal segment, Mesonotum flat before scutellurn. General
color of body rnetallic blue. Male with a reclinate frontal bristle .
................................................................................. Prolophormia
5 (2) Thoracic squama not setulose or lobulated, of variable colour. acr as strong
and distinct as the de, of usual strength, 2-3 + 3-4. Pollinosity of dorsurn rather
dense and forrning vittae, at least in front of suture area. Postalar depression
alrnost bare, sornetirnes only sorne setulae on upper falf. Parafacial setulae
(frontal view) extending downward below level of tip the third antennal joint.
Usually noturn sornewhat flat before scutellurn. Color of body dark rnetallic,
greenish or bluish. Myiasic producers.
6 (7) Basicosta yellow or brownish yellow. Both squarnae frorn yellow to yellowish
brown. Male: Distiphallus with en apical dorsal sclerotization. Fernale without
a reclinate frontal bristle Trypoealliphora
7 (6) Basicosta of variable colour, but rnainly dark. Both squarnae also of variable
coloration, predorninantly light coloured. Male: distiphallus without dorsal api­
cal sclerotization. Fernale with a reclinate frontal bristle ..
............................................................................... Proloealliphora

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88 s. V. PERIS

8 (l) Arista short-pi10se, and on1y so on the thicker part of arista. prst acr distinct.
Prathoracic spirac1e very 1arge, reaching fram near first coxae to the upper
part of notopleural suture. Head slight1y pratruding at vibrissal leve!. Body
b1ue-green, not pollinose. Parafranta1s non-pollinose. Boreal holarctic .
...................................................... .................................. Boreellus
The sixth genus known is Phormiata, known on1y fram one species in a unique speci-
men from Pamir (Tadjik). 1 have not seen it. In this key it seems to run to Boreellus,
with which is compared by GRUNIN (1971) for the arista short-pi10se and the 1ack
of prst acr. It differs by its head more 1arge1y pratruded at the base of antenna;
spirac1es of unusua1 size and a1so the 1ack of dc and is brist1es. The body of Phor-
miata is said to be shining b1ack with no metallic tinge.

5.2. List of genera

Boreellus ALDRICH & SHANNON, 1923, /ns. /ns. Menstr. 11: 107 (Type-species: Bo-
reellus aristatus ALDRICH & SHANNON, 1923, monotypy (=Sarcophaga atriceps
ZETTERSTEDT, 1845).
=Mallochomyia TOWNSEND, 1926 /ns. /ns. Menstr. 14: 25 (Type-species: Mallo-
chomyia johansenni TOWNSEND, 1926 (= Sarcophaga atriceps ZETTERSTEDT,
1845).
Phormia ROBINEAU-DESVOIDY, 1830 Myod.: 465 (Type-species: Musca regina MEI-
GEN, 1824 (design. ROBINEAU-DESVOIDY, 1849).
= Euphormia TOWNSEND, 1919, Proc. U.S. Nat. Mus. 56: 542 (Type-species: Mus-
ca regina MEIGEN, 1824, designo orig.)
Phormiata GRUNIN, 1971, Entom. Obozr. 50: 444 (Type-species: Phormiata phor-
miata GRUNIN, 1971, designo orig.)
Protocalliphora HOUGH, 1899, Entomo!. News 10: 66 (Type-species: Musca azurea
FALLÉN, 1817, orig. design.)
= Avihospita HENDEL, 1901, Wien. Entomo!. Ztg., 20: 30 (Type-species: Musca azu-
rea FALLÉN, 1817, orig. design.)
= Apau!ina HALL, 1948 B!owf!. N Amer.: 179 (Type-species: Protocalliphora avium
SHANNON & DOBROSKY, 1924, orig. design.)
= Orneocalliphora PEUS, 1960, (sbg. Protocalliphora) Dtsch. Entomo!. Z. (NF.) 7:
198 (Type-species: Musca chrysorroea MEIGEN, 1896, orig. design.)
= Philornis ENDERLEIN, 1936, Tierw. Mitte!eur. 6,2: 210 (Type-species: Musca azu-
rea FALLÉN, 1917, by present designation), nec MEINERT, 1899,
misidentification.
Protophormia TOWNSEND, 1908, Smiths. Misc. Coll. 51, n? 1.803: 123 (Type-species:
Phormia terranovae ROBINEAU-DESVOIDY, 1830, orig. design.)
Trypocalliphora PEUS, 1960, Dtsch. Entomo!. Zeit. (NE.) 7: 199 (Type-species: A vi-
hospita braueri HENDEL, 1901, orig. design.)

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 A PRELIMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 89

6. SUBFAMILY RHINIINAE

Since my revision of the group (PERIS, 1952) only a few genera have been pro-
posed and sorne are only new concepts applied to species or species-groups, there
are also sorne minor changes in nomenclature, these have been accepted and also
the modifications in generic concepts as far as they seem reasonable. The following
notes refer to all those changes and help to explain the additions and corrections
to the key.
Arrhinidia remains an unknown genus, and nothing else but the original de-
scription is known. It may not even be a Calliphorid, but nothing else can be said
with certainty. A note about it is the key in the proper place.
The division of Idiella (sensu PERIS, 1952) into two genera: Fainia for the Afro-
tropical species, and Idiella for the Indoaustralien, was already performed in the aboye
mentioned papero Actually there are differences in male genitalia, as well as in distribu-
tion, and so both genera are here accepted and so included in the key. This onIy needed
minor modifications.
The concepts of Rhinia and Stomorhina have also changed. In my previous paper
(1952) I was following MALLOCH (1926) in using the character of mesopleural bristles,
which is not as universal as I formerly thought; a lucid discussion of this is presented
by OEAR (1977: 790). It would be useful to reconstruct the species key based on the
new concepts.
Albaredaya is genuily a distinct genus, described almost simultaneously by PE-
RIS (1986) and ZUMPT (1957). I have again examined the types and can reassure that
the description of the antennal arista given in my paper was accurate. Comparison
between the two descriptions could leave sorne doubts concerning this character.
Trichoberia was placed in Rhyncomya in the 1952 paper, an opinion challenged
by ZUMPT (19587. Actually I now think that to place it in a different genus may be
more appropiated. I saw in 1952 only specimens in poor conditions and the study
of fresh ones my justify its location in the genus proposed by TOWNSEND in 1932.
Perisiella was made a new genus by ZUMPT (1958). It seems to me a matter of
opinion whether to retain it or to include it in Rhyncomya, forming a distinct species-
group as already I did (PERIS, 1952). In the key I accept it with my thanks to my
colleague ZUMPT for his naming.
Villeneuviella was not included by me in 1952. I considered it to be a Rhynchoes-
trini, as defined by SEGUY (1928), and so did ZUMPT (1956) in his revision of Pa-
learctic Calliphoridae. PONT (1980) made it a junior synonymy of Rhyncomya; and
certainly following 1952 key it runs to this genus and even near the deserticolous
forms of it. But it now appears to me that it desertes better its rank as a full genus.
Accordingly there is a new couplet in the key for its distinction.
Cosmina and Idiopsis are here maintained as a single genus despite the opinion
of ZUMPT (1956). He based its distinction mainly on the presence or absence of hairs
on the propleural depression. It is irrelevant to discuss here what a genus is, but really
I do not think that there is a clear cut distinction between specific and generic charac-
terso The reason why any ones are chosen as generic is only because they are common
to a number of species we think of as related, either by a mental hypothesis or other
consideration (1 would not be even against practical convenience as a reason). I think
that Cosmina and Idiopsis are close; their specific status of sorne of their species
is rather cumbersome and the whole groups needs a new revision. Unfortunately the

Bol. R. Soco Esp. Hist. Nat. (Sec. Biol.), 88 (1-4), 1992.

----------------------------------;------
90 S. V. PERlS

type material is scattered. For the time being 1 prefer to maintain aH the group together
awaiting new data. Therefore 1 retain Cosmina in my former wider sense.
Lastly there is Strongyloneura. For a long time and by many authors this name
was used for the present Isomyia (see sorne comments on this name in PERIS, 1956).
KURAHASHI (1967-on) seems to leave things as arranged by ZUMPT. However 1 think
that these species near to prasina are worthy of separation by their morphological
characters and even for practical reason. Isomyia is a large genus and it may be worthy
to make aggregates for it but without denaturalising it, Stronglyloneura is perhaps
the only one accomplising this requirement. From a biogeographic view it has also
a rather fixed pattern, around Japan and related islands. So 1 maintain in'this key
the separation of Strongyloneura as a good and natural genus.

6.1. Key to genera

1 (14) Arista pectinate, with rays only on dorsal part of arista and rather long and
distinct, if sorne cilia are present on the underside (Albaredaya) they are reduced
to small cilia of a different arrangement, structure and aspect. aer and de
present only in front the scutellum. Propleural depression bare. Suprascuamal
ridge bare.
NOTE: Arrhinidia is known only from its original description and has not been redescribed. Its
arista is mentioned as pectinated and the propleral depression as bare. Both characters isolate
it from al! other genera here included.

2 (3) Transverse m-m strongly angled inwards forming almost a right angle. Band
of m strongly curved. R5 closed and short petiolated, burseshaped. Cell R3
narrow at its middle. Fore tibia without medianpv seta. Upper occiput strongly
concave, its lower part convexo Front in both sexes wider than width of ocellar
triangle. Afrotropical.. Vanemdenia
3 (2) Transverse m-m not angled inwards, more or less sigmoid. Band of m not
strongly curved, rounded or angulate. R3 of similar width along its entire
length. Occiput rather flat. Frons of male narrower than width of ocellar
triangle.
4 (5) No ph seta more external than the prst. Longest rays of the arista not exceed-
ing one half the width of third antennal joint. Flies of stout build. prstg bristle
very fine or absent. Fore tibia with a submedian pV. R5 cell closed. Indo-
australian Chlororhin ia
5 (4) One ph sete more external than the prst. Longest rays of arista longer than
the width of third antennal segment.
6 (13) prstg seta absent.
7 (10) Hind tibie with 2-3 ad setae as large as the tibial diameter, no other longer
setae differentiated. Mesopleura with only two posterior bristles. R5 cell open,
sometimes very narrowly so. Flies rather rather slender with abdomen of almost
subparallel sides and yellow coloured. Male: First abdominal tergite without
marginal hairs.
8 (9) Fore tibia without a submedian pv seta. Femore of red or redish colour.
Pollinose areas without piliferous spots. Sternopleure with or without yellow

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A PRELIMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 91

pollen. Male with cerci fused and fifth sternite with pilose lateral prominences.
Afrotropical Fainia
9 (8) Fore tibia with a submedian pv seta. Femora dark, blackish. Sternopleura
always yellow pollinose, with or without piliferous spots. Male: Cerci free,
not fused and fifth sternite of the usual form, without such pilose promi-
nences. Indo-australian /diel/a
10 (7) Hind tibia with a file of ad subequal setulae longer than the surrounding
setulae of tibiae, including in the row 2-3 longer ones. Fore tibia with a sub-
median pv seta. Flies of variable size, usually small, but of more stout ap-
pearance, with oval abdomen which is often, at least partially, dark.
11 (17) R5 cell petiolate. Sternopleura glossy, not pollinose. Mesopleura with no pi-
liferous spots. Mesopleural setae reduced to the upper posterior ones present.
Legs yellow. Male: Abdominal tergite I + II usually with long posterior
marginal hairs. Cerci arched and forceps-like, fifth sternite with rather stout
spinules on the apical part of the lateral arms. Tropical and Subtropical Old
W orld , , , Rhinia
12 (11) R5 cell open or closed, if petiolate the stenopleura thicly pollinose. Male cerci
of usual form, not forceps-like and fifth sternite no spinulose on lateral
arms. Warm areas of Old World Stomorhina
13 (6) prst bristle present. Fore tibia with a short submedian pv seta. Mesopleural
posterior bristles complete. Arista long-pectinate on dorsal part, at high
magnification and good light conditions, sorne short white ciliation visible
on ventral part. Crossvein m-m rather sigmoid. Band of m very smooth and
closing R5 cell at wing border. Occipital dilatation and lower occiput lemon
yellow and not distinctly pollinose. Mesonotum, including humeri, metallic
green, pleural areas and legs yellow. Propleural depression bare. Rather slan-
der flies. Madagascar. .A/baredaya
14 (1) Arista pubescent or plumose, not pectinate and with hairs or rays, similar
in both sides, dorsal and ventral.
15 (16) prst bristle absent (only present in a Stegosoma species, easily distinguished
by its glossy yellow integument and stout-bodied aspect) (In Trichoberia the
prstg bristle is also lacking although it is difficult to see due to the surroun-
ding thick pile, but this same character facilitates its identification). Fore tibia
with a small submedian pv seta, very small in Trichoberia, being scarcely
the length of tibial diameter.
16 (15) Body covered with thick yellowish-white pile, giving a bombyliid-like appear-
ance. Hypopleural setae wanting, replaced by a tuft of similar pale yellow
hairing. Head swollen-Iooking and translucent pale yellow, with broad pa-
rafrontals and parafacials, the wholle head globose. Frontal setae fine and
delicate accompanied by fine black and yellow setulae, these extending on
to parafacials where they are short and yellow. The face strongly widening
at level of the middle the third antennal joint. Legs and palpi yellow. Afro-
tropicaL Trichoberia
17 (15) Not so, more usual froms. Hypopleural setae black and present in near vertical
row.

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 92 S. V. PERIS

18 (21) No ph seta positioned more externally than the prst. Yellow coloured fIies
and with the R5 cell open.
19 (20) Arista bare or almost so, Glossy yellow fIies, of short and stout body. Notal
setae reduced to the prse pairs. Afrotropical.. Stegosoma
20 (19) Arista long plumose. Slender fIies, not glossy. Oriental. Borbororhinia
21 (18) One ph seta positioned more externally that the prst. At least the prst de
distinct. Arista practically bare.
22 (23) Suprasquamal ridge posteriorly setulose. R5 cell open. Pteropleural seta ab-
sent. Palpi wide, somewhat leaf-shaped, about three times as long as wide.
Afrotropical Pseudorhyneomyia
23 (22) Suprasquamal ridge bare. At least one pteropleural seta.
24 (25) R5 cell open. At least one pair of prst de. Palpi scarcely widened at apex,
with almost subparallel sides and length about four times their width. Afro-
tropical Zumba
26 (15) prstg seta present, rarely absent and then the fore tibia without a submediam
pv seta.
27 (28) Suprasquamal ridge setulose on its posterior parto aer and de rows well de-
veloped. R5 call open. Arista long pilose, the longest hairs slightly ascend-
ing one half the width of third antennal joint. Propleural depression bare.
Fore tibia with a submedian pv seta. Afrotropical.. ..... .... Euehyneomyia
28 (27) Suprasquamal ridge bare its whole legth.
29 (36) Arista not plumose, the longest hairs never exceed half the width of the third
antennal segmento Fore tibia with a submedian pv seta.
30 (31) Arista long pilose, the largest hiras clearly exceed twice the half width of
the third antennal segment, Epistoma protruding between vibrissae in profile.
Propleural depression bare. Parafacials with a black shining spot: Face and
parafrontals also darkened on sorne parts. Thorax metallic and rather dense
pollinose with darker spots at bases of setulae, a brillian stripe, no pollinose,
from humeri to wing-base at each side. Wings with the anterior border dark-
ened. Afrotropical. Perisiel/a
31 (30) Arista bare or short pilose, the longest hairs not so long, or there is another
combination of characters. Fore tibie with a submedian pV.
32 (35) Mouth-parts aparently functional, of the usual size and with a normal haus-
tellum with distinct labellae. Palpi more or less elongata and distinct.
33 (34) Propleural depression pilose. Indo-australian Metal/ea
34 (33) Propleural depression bare. South Palaearctic, Afrotropical & Oriental. ..
...................................................... ........................... Rhyneomya
35 (32) Mouth-parts reduced, represented by a pair of globular structures, oval
swol1en, which are probably the palpi, not at al1 visible from outside in profile;

Bo/. R. Soco Esp. Hist. Nat. (Sec. Bio/.), 88 (1-4), 1992.

---~---~--- ~
A PRELIMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 93

between them is a short central subconical structure, shorter than the palpi
and which is probably the reduced haustellum. Body vestiture short and spar-
se, not clear prst bristies, only sorne prse setae slightly differentiated. Third
antennal segment almost the same length as the second. Arista practically
bare and rather short, only a little longer than the combined length or the
second and third antennal segments. Male with wide frons, about three ti-
mes the width of the ocellar triangle. The frontal setae as short as the ac-
companing setulae, which are sparse, extending onto the upper part of para-
facials. Interfrontalia not distinghishable by its texture or color from para-
frontalia which are only separated by a little distinct groove. Deserts of Old
World Villaneuviella
36 (29) Arista plumose, the longest rays at least subequal to the width of the third
antennal segment.
37 (38) Fore tibiae without a submedian pv seta. prstg seta very fine or absent. A
ph seta situated laterally the prst. Oriental. Malayomyza
38 (37) Fore tibia with a submedian pv seta.
39 (42) No ph seta lateral of the prst.
40 (41) Mesopleura without setae on its upper border near notopleura. Band of m
smooth and regularly curved, not angulate. Parafrontals and parafacials
narrower than width of the third antennal joint. Interfrontalia in both sexes
wider than the ocellar triangle. prst aer and de absent. Female without distinct
orbital setae. Oriental. Sumatria
41 (40) Mesopleura with strong setae on its upper border near notopleura. Band of
m widely angulate. Parafrontals and parafacials clearly wider than third
antennal joint wide. Interfrontalia in both sexes narrower than width of oce-
llar triangle. prst aer and de well developed and distinct. Female with two
orbital proclinate setae. Oriental and E. Africa Thoraeites
42 (39) One ph seta more lateral of the prst.
43 (44) prst aer absent or indistinct; post de and aer usually also indistinct except
the prse ones; rarely one or two prst aer longer than surrouding vestiture
but then the propleural depression thickly pilose. South Palaearctic, Afro-
tropical & OrientaL Cosmina (including /diopsis)
44 (43) prst aer well developed at least in one pair clearly distinct, the de also distinct.
Propleural depression bare.
45 (46) Mesopleura with a group of setae on its upper par near notopleura. Bend
of m regularly curved or angulate. Male with hypopygium of the usual normal
size, scarcely visible when the abdomen is viewed im profile. The last sternite
in female not protruding posteriorly. Tropics of the Old WorId ..
...................................................... ................................ /somyia
46 (45) Mesopleura without such setae. Bend of m smooth. Male hypopygium large,
as large as the pregenital segment and together these are larger than the rest
of the abdomen. Female with last sternite widely esposed by corresponding

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94 S. V. PERIS

tergites and its posterior border somewhat protruding outwards. Japan, Ryuk-
yu and Taiwan ... Strongyloneura
o ••••• o ••••• o . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . •

6.2. List of genera

Albaredaya PERlS, 1956, Eos 32: 238 (Type-species: Albaredaya malgache PERlS,
1956, orig. design.)
= Cosminiella ZUMPT, 1957, Nat. Malgache 9: 113 (Type-species: Cosminiella ma-
dagascariensis ZUMPT, 1957, orig. designo (=Albaredaya malgache PERlS, 1956).
Arrhinidia BRAUER & BENGESTAMM, 1891, Denskrschr. Akad. Wien 58: 390 (Type-
species: Rhynchomyia aberrans SCHINER, 1868, monotypy).
Borbororhinia TOWNSEND, 1917, Rec. Ind. Mus. 13 (4), n? 2: 188 (Type-species: Bor-
bororhinia pubescens TOWNSEND, 1917, orig. designo (= Idia bivittata W ALKER,
1857).
=Alikangia VlLLENEUVE, 1927, Rev. Zool. Afr. 15: 389 (Type-species: Alikangia
pulchella VlLLENEUVE, monotypy (= [dia bivittata WALKER, 1857).
Chlororhinia TOWNSEND, 1917, Rec. Indo Mus. 13 (4) n? 12: 191 (Type-species: Chlo-
rorhinia viridis TOWNSEND, 1917 (orig. design.) (=Musca exempta WLAKER,
1857).
Cosmina ROBlNEAU-DESVOlDY, 1830, Myod.: 433 (Type-species: Cosminafuscípe-
nis ROBlNEAU-DESVOlDY, 1830, designo BRAUER & BERQENSTAMM, 1889).
= Seseromyia RONDANl, 1863, Arch. Zool. Modena 3: 32 (Type-species: Idia punc-
tulata WIEDEMANN, 1819 (orig. design.)
= Synamphoneura BIGOT, 1886, Bull. Soco Entomol. Fr. 1886: 14 (Type-species:
Synamphoneura cuprina BIGOT, 1886, monotypy).
= Idiopsis BRAUER & BERGENSTAMM, 1889 Denskchr. Akad. Wiss. Wien 6: 85 (Type-
species: Idiopsis prasina BRAUER & BERGENSTAMM, 1889, monotypy).
=Eusynamphoneura TOWNSEND, 1917, Rec. Ind. Mus. 13, 4 n? 12: 189 (Type-
species: Idia seriepunctata LOEW, 1852, orig. design.)
=Synamphoneuropsis TOWNSEND, 1917, Rec. Ind. Mus. 13, 4 n? 12: 199 (Type-
species: Synamphoneuropsis viridis TOWNSEND, 1917, orig. design.)
= Synamphoneurops ZUMPT, 1956, Die Fliegen, USA 641: 116, errore pro
Synamphoneuropsis.
Eurhyncomyia MALLOCH, 1926, Ann. Mag. Nat. Hist. (9) 19: 513 (Type-species:
Rhyncomyia obtusa (BIGOT) sensu MALLOCH, 1925, orig. designo
(= Eurhyncomyia thoracica CURRAN, 1931 = Rhynchomyia diversicolor BlGOT,
1887) =Eurhynchomyia auct., errore.
Fainia ZUMPT, 1956, Explor. Parc Natur. Albert, Miss. Witte, 92: 83 (Type-species:
Idiella albitarsis MACQUART, 1846, orig. design.)
Idiella BRAUER & BERGENSTAMM, 1889, Denskchr. Akad. Wiss. Wien 56: 154 (Type-
species. Idiella mandarina WIEDEMANN, 1830), monotypy).

Isomyia W ALKER, 1860 (sbg. Musca), Proc. Linn. Soco Lond. 4: 134 (Type-Species:
Musca (Isomyia) delectans WALKER, 1860, monotypy.

Bol. R. Soco Esp. Hist. Nat. (Seco Biol.), 88 (1-4), 1992.

A PRELIMINARY KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 95

= The/ychaeta BRAUER & BERGENSTAMM, 1891; Denskchr. Akad. Wiss. Wien 58:
390 (Type-species: The/ychaeta ch/aybea BRAUER & BERGENSTAMM, 1891. mo-
notypy (=Musca viridaurea WIEDEMANN, 1819, nec Tachina viridaurea WIE-
DEMANN, 1842).
=Apol/enia BEZZI, 1911, Bol/. Lab. 200/. Portici 6: 79 (Type-species: Apol/enia nu-
diuscu/a (BIGOT) sensu BEZZII, 1911, orig. designo ( = Curtoneura tristis BIGOT,
1887).
=Anna MALLOCH, 1926, Ann. Mag. Nat. Hist. (9) 18: 520 (Type-species: Anna ca-
//iphoroides MALLOCH, 1926, orig. design.)
Strongy/oneuropsis TOWNSEND, 1927, Phi/ip. J. Sci. 34: 376 (Type-species: Strong-
y/oneuropsis ma/ayensis TOWNSEND, 1827, orig. design.)
=Pachycosmina SÉGUY, 1934, E.E. Dipt. 7: 18 (Type-species: Pachycosmina oes-
tracea SÉGUY, 1934, orig. design.)
= The/ychaetopsis SÉGUY, 1949 (sbg. Isomyia) Rev. Brasil. Bio/. 9: 115 (Type-species:
Strongy/oneura pseudo/ucilia MALLOCH, 1928, orig. design.)
= Gerschia LEHRER, 1970, Annot. 200/. Bot. Bratis/ava 61: 30 (Type-species: Isom-
yia eos ZUMPT, 1958, orig. design.)
=Strongy/oneura auct. pIur., nec BIGOT, 1886.
Ma/ayomyza MALLOCH, 1928, Ann. Mag. Nat. Hist. 1 (10): 491 (Type-species: Ma-
/ayomiza humeralis MALLOCH, 1928, orig. design.)
Metal/ea WULP, 1880, Tijds. Entomo/. 23: 174 (Type-species: Metal/ea notata WULP,
1880, monotypy).
= Meta//iopsis TOWNSEND, 1917, Rec. Ind. Mus. 13, (12) 193 (Type-species: Meta-
//ipsis setosa TOWNSEND, 1917, orig. design.)
= Ch/ororhynchomyia TOWNSEND, 1932, J. N. Y. Entom. Soco 40: 440 (Type-species:
Ch/ororhynchomyia clausa TOWNSEND, 1932, orig. design.)
Pararhynchomyia BECKER, 1910, Denskchr. Akad. Wiss. Wien 71: 142 (Type-species:
Pararhynchomyia cribriformis BECKER, 1910, monotypy).
Perisiel/a ZUMPT, 1958, Exp/or. Parc. Nat. A/bert. Miss. Witte 92: 187 (Type-species:
Rhynchomyia anchora WIEDEMANN, 1824, orig. design.)
Pseudorhyncomyia PERIS, 1952, An. Est. Exp. Au/a Dei 3: 50 (Type-species:
Rhynchomyia braunsi VILLENEUVE, 1920, orig. design.)
Rhinia ROBINEAU-DESVOIDY, 1830, Myod.: 421 (Type-species: Rhinia testacea
ROBINEAU-DESVOIDY, 1830, monotypy (=Idia apicalis WIDEMANN, 1830).
= Beccarimyia RONDANI, 1873, Ann. Mug. Genova 4: 287 (Type-species: Beccarimyia
g/ossina RONDANI, 1873, monotypy (= Idia apicalis WIEDEMANN, 1830).
Rhyncomya ROBINEAU-DESVOIDY, 1830, Myod.: 424 (Type-species: Musca felina
(FABRICIUS) ROBINEAU DESVOIDY, 1830, monotypy (=Musca ruficeps FABRI-
CIUS, 1805).
=Rhynchomyia & Rhynchomyia, Rhynchomyia, Rhyncomya, errare pIur.
?=Beria ROBINEAU-DESVOIDY, 1830, Myod.: 418 (Type-species: Beria inf/ata.
ROBINEAU-DESVOIDY, 1830, monotypy).
= Rhynchomyopsis TOWNSEND, 1917, Rec. Ind. Mus. 13-14 (12): 194 (Type-species:
Rhynchomyiopsis indica TOWNSEND, 1917, orig. design.)
= Trichometal/ea TOWNSEND, 1917, Rec. Ind. Mus. 13-14, (12): 194. Trichometa-
l/ea po//inosa TOWNSEND, 1917, orig. design.)

Bol. R. Soco Esp. Hist. Nat. (Sec. Biol.), 88 (1-4), 1992.

96 S. V. PERIS

=Doljia SUSTER, 1955, Bul. Sto A cad. Repub. P. Rom. Bucharesti (Biol.) 5: 768
(Type-species: Doljia viridicauda SUSTER, 1855, monotypy (= Idia speciosa
LOEW, 1844).
= Sokotra LEHRER, 1970, Annot. Zool. Bot. Bratislava 61: 32 (Type-species:
Rhynchomyia varifrons BECKER, 1910, orig. design.)
Stegosoma LOEW, 1863, Wient. Entomo!. Mon. 7: 15 (Type-species: Stegosoma vin-
culata LOEW, 1863, monotypy).
Stomorhina RONDANI, 1861, Dipt. Ital. Prodr. 4: 9, n. nomo pro Idia WIEDEMANN,
1820, preocc.
=Idia WIEDEMANN, 1820, Nov. Dipt. Gen.: 21, preocc. HUBNER, 1809 (Type-
species: Musca lunata FABRICIUS, 1805, designo BRAUER & BERGENSTAMM,
1889).
= Stomatorrhinia BEZZI, 1906, Sts. Syst. Hym. Dipt. 6: 53, emend.
=Stomatorhinia BEZZI & STEIN, 1907, Kat. Pal. Dip. 3: 583, emend. pro Stomor-
hina RONDANI, 1861).
= Stomorhyna, lapsus.
= Idielliopsis TOWNSEND, 1917, Rec. Ind. Mus. 13,4 (12): 190 (Type-species: Idie-
lliopsis similis TOWNSEND, 1917, orig. designo ( = Idia xanthogaster WIEDEMANN,
1820).
=Euidiella TOWNSEND, 1917, Rec. Ind. Mus. 13,4 (12): 192 (Type-species: Musca
discolor FABRICIUS, 1805, orig. design.)
Strongyloneura BIGOT, 1886, Bull. Soco Entomol. Fr. (1886): 14 (Type-species):
Strongyloneura prasina BIGOT, 1886, monotypy).
= Chloroidia TOWNSEND, 1917, Rec. Ind. Mus. 12 (4) n? 12: 196 (Type-species: Chlo-
roidia flavifrons TOWNSEND, 1917, orig. design.)
Sumatria MALLOCH, 1926, Ann. Mag. Nat. Hist. (9) 18: 512 (Type-species: Suma-
tria latifrons MALLOCH, 1926, orig. design.)
=Eucosmina MALLOCH, 1928, Ann. Mag. Nat. Hist. (10) 1: 492 (Type-species:
Eucosmina vittigera MALLOCH, 1928, orig. design.)
=Alikangiella VILLENEUVE, 1927, Rev. Zool. Afr. 15 (3): 389 (Type-species: Ali-
kangiella flava VILLENEUVE, 1927, monotypy).
Thoracites BRAUER & BERGENSTAMM, 1891, Denkschr. Akad. Wiss. Wien. 58: 59
(Type-species: Musca abdominalis WIEDEMANN, 1805, monotypy).

Trichoberia TOWNSEND, 1932, f.N. Y. Entomol. Soco 40: 439 (Type-species: Tricho-
beria rufopilosa TOWNSEND, 1932, orig. design.)
Vanemdenia PERIS, 1951, Eos 27: 237 (Type-species: Vanemdenia africana PERIS,
1951, orig. design.)
Villeneuviella AUSTEN, 1914, Novit. Zool. 21: 272 (Type-species: Villeneuviella har-
terti AUSTEN, 1914, monotypy).
= Rhynchoestrus SÉGUY, 1926, E.E. Dipt. 3: 1 (Type-species: Rhynchoestrus weissi
SÉGUY, 1926, monotypy).
Zumba PERIS, 1951, Eos 27: 239 (Type-species: Zumba rhinoidea PERIS, 1951, orig.
design.)

Bo/. R. Soco Esp. Hist. Nat. (Sec. Bio/.), 88 (1-4), 1992.

A PRELIMINAR Y KEY TO THE WORLD GENERA OF THE SUBFAMILIES OF CALLIPHORIDAE 97

ACKNOWLEDGEMENTS

My thanks are due to Dr. David R. Ragge and P. C. Barnard, respectively

Deputy Keeper of Entomology and Head of Diptera of the Natural History Museum
of London for their help during my stay there. Also my thanks to Mr N. P. Wyatt
for his help in finding of needed specimens. Also to the authorities of my Universi-
dad Complutense for their aid for the visit to London, to Prof. V. Monserrat, Dr.
M. D. González-Mora, Javier M. Peris and Dr. H. Kurahashi for their help in many
ways.

Recibido el 14 de junio de 1990

Aceptado el 30 de octubre de 1991

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