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Topic 2: Ecosystems and ecology

2.1 Species and populations


Species, Habitat, Niche
Ecosystem → a community of interdependent organisms (biotic) and their interactions
with the physical environment (abiotic) they inhabit.
Species → a group of organisms sharing common characteristics that can interbreed
and produce offspring that also produce young.
Limitations: doesn’t classify extinct populations, doesn’t account for asexual
organisms
Habitat → environment in which a species lives, where an organism can find food,
shelter, protection, mates.
Niche → the set of biotic and abiotic conditions and resources that an organism or
population responds to (not just habitat, but how an organism interacts with others). No
two species can have the same niche.
Fundamental Niche: full range of conditions/resources an organism can survive
and reproduce in.
Realised Niche: actual conditions a species exists in due to biotic interactions.
→ different niches can still share the same habitat due to space, behaviour
patterns etc

Abiotic Factors
The nonliving, physical factors that influence organisms and ecosystems.
Eg. temperature, sunlight, pH, precipitation, soil, landscape/topography.
→ there are upper and lower level limits beyond which a population cannot
survive
(optimal range → physiological stress → intolerance)
Population Interactions
Predation → when one animal or plant hunts and eats another organism.
→ predator-prey relationships are controlled by negative feedback
mechanisms
→ as prey increases, after time so do predators. Increase in predators
reduces number of prey.
→ predation benefits prey - removes old/sick individuals, leaving superior
breeding pool
Eg. lemming and snowy owl, gray wolf and moose
Herbivory → where an animal feeds on a plant - the animal is known as a herbivore.
→ the plants as a food source affect the carrying capacity of the
environment for the herbivore
Eg. hippopotamus grazing on vegetation / zooplankton feeding on phytoplankton
Parasitism → where one organism benefits at the expense of another (the host), type of
symbiotic relationship
→ endoparasites live inside hosts, ecto parasites live on the surface of
hosts
Eg. tapeworms (endoparasites) and ticks/mites (ectoparasites)
Eg. plants such as the Rafflesia flower via root systems
Mutualism → another form of symbiosis where both species benefit
Eg. coral reefs: zooxanthellae live within coral animal (polyp; they
photosynthesise to produce food for themselves and the polyp; in exchange they are
protected.
Disease → aka. pathogen - can be bacteria, virus, fungi - reduces carrying capacity of
infected organism.
Eg. Dutch elm disease, caused by fungus clogging vascular tissues in tree,
preventing water movement.
Competition → demand by individuals for limited environmental resources
→ can be intraspecific (within a species) or interspecific (between diff
species)
→ the degree to which niches (inter) overlap determines the level of
competitive exclusion
Population Growth
Population → group of organisms in the same species living in the same area at the
same time, capable of interbreeding.
→ abundance of resources affects population structure over time.

S Population Curve (aka sigmoid curve)


Rapid initial growth, then slowing as carrying capacity is
reached, where population fluctuates around K.
→ divided into lag phase, exponential growth phase,
transitional phase, stationary phase

J Population Curve
Increasingly rapid exponential growth with no signs of slowing,
exhibited by organisms that produce rapidly.
→ controlled by favourable abiotic components, results in a
population crash

Limiting Factors → slow population growth as carrying capacity is reached


Density-Dependent Factors - lower birth rate/raise death rate as population
grows
Density-Independent Factors - affect a population regardless of density, abiotic
factors

2.2 Communities and ecosystems


Communities and Ecosystems
Community → all populations living and interacting in a common habitat at a specific
time (only biotic)
→ involves many interactions among species, communities with higher
diversities are more stable and resilient to disturbances.
Ecosystem → a community and the physical environment it interacts with (biotic
interacting w/ abiotic)
→ divided into terrestrial, marine, and freshwater, defined uniquely by
various abiotic factors
Photosynthesis and Respiration
→ all organisms respire, only producers photosynthesise
Photosynthesis → process by which a plant converts light energy from the Sun into
usable chemical energy stored in organic matter. Produces the raw material for
biomass.

carbon dioxide + water → glucose +


oxygen

Inputs: sunlight as energy, CO2, H2O


Outputs: glucose (foundation for other organic molecules)
Transformations: light energy → chemical energy stored in biomass

Respiration → conversion of organic matter to carbon dioxide and water in all living
organisms, releasing energy for life processes. “Wasted” energy is lost as heat,
increasing the entropy in the ecosystem while allowing organisms to maintain low
entropy.

glucose + oxygen → carbon dioxide + water

Inputs: organic matter (glucose) and oxygen


Outputs: release of energy to maintain order (counteract entropy) and heat
Transformations: stored chemical energy → kinetic energy and heat energy

Feeding Relationships
Producers → autotrophs - organisms that convert abiotic components (sunlight) into
living matter. Support the ecosystem through constant input of energy and new
biomass.
→ convert sunlight energy to chemical energy using photosynthetic
pigments
Consumers → heterotrophs - organisms that eat other organisms to obtain energy and
matter
→ pass energy and biomass from producers through to apex predators
Decomposers → break down tissue of dead organic matter and release nutrients for
reabsorption by producers
→ improves the ability of soil to retain nutrients - the organic byproducts of
decomposed matter contribute to the humus in soil
→ essential for cycling matter within ecosystems, esp. in carbon and
nitrogen cycles

Trophic Levels, Food Chains, Food Webs


Flows of energy and matter can be shown through food chains. Interconnected chains
form food webs.
→ the position an organism occupies is known as its trophic level. Producers
form the first trophic level.
→ organisms can occupy different trophic levels depending on which food chain
it is in
→ decomposers feed at every level of the food chain
Producer → Primary consumer → Secondary consumer → Tertiary consumer
Autotroph → Herbivore → Omnivore/carnivore → Carnivore

Efficiency of Energy Transfers Through an Ecosystem


Only ~10% of energy is transferred to the next trophic level - trophic efficiency = 10%.
→ 2nd law of thermodynamics states energy transformations inefficient, so
energy is lost at each level
→ most energy is lost as heat energy through respiration
→ each trophic level has a smaller energy pool from which it can withdraw
energy, thus limiting the amount of trophic levels in a food web

Pyramids of Numbers, Biomass, Productivity


Ecological Pyramids → models showing the quantitative differences between the trophic
levels of an ecosystem, measured for a given area and time. Shows feeding
relationships in a community.

Pyramid of Numbers → records number of individuals at each trophic level in an


ecosystem
→ numbers tend to decrease along food chain and pyramid tends
to narrow at apex
→ can be inverted when size of individuals at lower trophic levels is
large (eg. tree)
Advantages: easy method of comparing changes in population numbers over
different times
Disadvantages: numbers too great to be accurate, doesn’t represent energy flow
accurately
Pyramid of Biomass → the standing stock or storage of each trophic level
→ measured in g m-2 (grams per m2) or j m-2 (joules per m2)
→ can depend on seasonal variations as they are a snapshot in
certain time
Advantages: overcomes problems w/ pyramid of numbers
Disadvantages: only uses sample populations, organisms must be killed to
measure dry mass, subject to seasonal variations, gives no indication of productivity
over time
Pyramid of Productivity → shows flow of energy through trophic levels, the rate at which
biomass is being generated.
→ measured in g m-2 yr-1 (energy per unit area per unit time)
→ shows amount of energy available as food to next trophic level,
more useful in measuring a system’s changes over time
→ always show a decrease, as transfer of energy is inefficient
Advantages: shows rate of productivity of a system
Relevance of Thermodynamics
First Law → energy cannot be created or destroyed, only transformed.
→ Solar energy is absorbed by plants and converted to stored chemical
energy. Chemical energy is used and lost as heat energy.
Second Law → energy transformations result in a loss of energy, leaving less energy to
maintain order, thus increasing entropy over time.
→ Energy is transferred when trophic levels feed off one another, thus
more energy is lost through each trophic level.
Bioaccumulation and Biomagnification
Bioaccumulation → build up of persistent / non-biodegradable pollutants within an
organism/trophic level because it cannot be broken down.
Biomagnification → increase in concentration of persistent / non-biodegradable pollutant
along a food chain.
***biomagnification does not occur due to higher trophic levels eating more - the
respiration of biodegradable biomass leaves non-biodegradable toxin in higher
concentration.

Impact of Pollutant → DDT, a non-biodegradable pollutant used as a pesticide by


farmers.
→ producers take in DDT, organisms in 2nd trophic level retain the
pesticide in their body tissue from the producers (bioaccumulation) as it is unable to be
broken down
→ process continues, with more DDT accumulated at each level. Top
carnivores are ultimate accumulators, and are the most vulnerable to ecosystem
disruption due to their small population and high doses of toxins they receive.
2.3 Flows of energy and matter
Transfer and Transformation of Energy
As solar radiation (insolation) enters the atmosphere, some energy becomes
unavailable for ecosystems as the energy is absorbed by inorganic matter / reflected
back into the atmosphere.
→ 51% of available energy from sun does not reach producers
→ 49% absorbed by ground - only 0.06% of all radiation is captured by
chloroplasts
Producers convert light energy to chemical energy, which is then available for other
organisms to use; all energy is lost from an ecosystem in the form of heat →
Ecological efficiency = energy used for growth (biomass development) x 100
Energy supplied
Energy Pathways →
1. Light → chemical
2. Transfer of chemical from one trophic level to another
3. Conversion of visible light and UV to heat energy
4. Re-radiation of heat energy to atmosphere

Primary and Secondary Productivity


Primary Productivity (PP) → the gain by producers in energy/biomass per unit area per
unit time.
→ depends on amount of sunlight, availability of factors needed for growth, ability
of producers to use energy to create organic molecules
→ highest growth occurs in optimal conditions (warm, high nutrients, water etc
eg. tropical rainforests)
Gross Primary Productivity (GPP) → mass of glucose created by photosynthesis
per area per time.
Net Primary Productivity (NPP) → gain by producers in energy/biomass after
accounting for loss of energy through respiration. Represents potential energy available
for next level of consumers. NPP = GPP - R.

Secondary Productivity (SP) → the biomass gained by heterotrophs through feeding


and absorption, measured in unit mass/energy per unit area per unit time.
→ depends on amount of food present and efficiency of energy conversion to
new biomass
Gross Secondary Productivity (GSP) → total biomass assimilated by
consumers.
GSP = FOOD EATEN - FAECAL LOSS.
Net Secondary Productivity (NSP) → gain by consumers in energy/biomass after
allowing for respiration. Represents amount of potential energy available for next trophic
level. Aka. assimilation.
NSP = GSP - R

Maximum Sustainable Yields


Rate of increase in natural capital that can be exploited without depleting original stock.
→ equivalent to NP (whether NPP or NSP) of a system - net productivity is
amount of energy stored as new biomass; removal of biomass above max sustainable
yield reduces natural capital and is unsustainable.
Nutrient Cycles
→ Energy flows; matter cycles between abiotic and biotic environments in an
ecosystem.
Factors Affecting Nutrient Cycles → soil erosion, runoff, amount of rainfall,
decomposition, plant density etc.
→ Nutrients can be stored in organic (plants and animals) or inorganic (rocks etc.)
matter.
→ Macronutrients are needed in large quantities, eg. carbon, nitrogen, oxygen,
hydrogen

Carbon Cycle →
Carbon - essential in ecosystems, forms a key component of all biological molecules
(proteins, fats etc)
→ can be stored in trees, fossil fuels, limestone for long periods of time as well
as organic matter
Storages: organic → organisms (plants + animals)
inorganic → atmosphere, soil, oceans, fossil fuels
Flows: transfers → herbivores x producers, carnivores x herbivores, decomposers x
dead organic matter
→ CO2 in atmosphere dissolves into oceans
transformations → photosynthesis (carbon dioxide turned into glucose +
oxygen)
→ respiration (organic matter turned into carbon dioxide)
→ combustion (biomass is turned into carbon dioxide)
→ fossilisation (dead organic matter turned into fossil fuels via
pressure and decay)

Nitrogen Cycle →
Nitrogen - building block for amino acids and DNA
→ the most abundant gas in atmosphere (80%) but largely inaccessible due to its
stability
→ can only be converted by certain organisms
Storages: organic → organisms
Inorganic → soil, fossil fuels, bodies of water, atmosphere
Flows: transfers → herbivores x producers, carnivores x herbivores, decomposers x
dead organic matter
→ plants absorbing nitrates through roots, metabolic waste products from
organism (excretion)
transformations → fixation of nitrogen from atmosphere by lighting and
nitrifying bacteria
→ nitrifying bacteria transforms ammonium ions into nitrite →
nitrate
→ denitrifying bacteria transforms nitrates back to nitrogen
→ decomposers break organic nitrogen (proteins) into ammonia
→ nitrogen from nitrates used by plants to make amino acids and
protein (assimilation)
Impact of Human Activities on Energy Flows and Matter Cycles
→ combustion of fossil fuels, urbanisation, agriculture, deforestation impact both energy
and matter flows
Energy Flows
→ industrial revolution increased use of fossil fuels, allowing humans access to energy
trapped in oil, coal etc.
→ amount of energy available to humans increased, increasing agricultural
output
→ however, changes in ‘energy budget’ lead to climate change, reduction of
natural capital etc
→ combustion of fossil fuels alters the way light energy interacts w/ surface of
planet and atmosphere
→ increased CO2 → increasing temp → reduction in ice → less reflected
sun energy → more GHG
→ pollution → increased trapping of solar radiation → more heat
Matter Cycles
→ timber harvesting interferes w/ nutrient cycling through decomposition
→ removal of trees = canopy cannot intercept rainfall and rich floor litter is
washed away
→ trees often cleared to grow oil palm (for food, domestic products and biofuel),
thus increasing need for fertiliser in nutrient poor rainforest soil to produce yields
→ fertilisers contain nitrates, leading to contamination of nearby bodies of water
(eutrophication)
→ harvested crops are transported, along w/ sequestered nitrogen, altering
storages
→ burning fossil fuels reduces storages of non-renewable energy and increases storage
of carbon in atmosphere

2.4 Biomes, zonation and succession


Biomes
Biome → collections of ecosystems that share distinctive abiotic factors, species and
climatic conditions.
→ influenced by rainfall, insolation and temperature
→ water and insolation needed for photosynthesis, which determines
productivity
→ water needed for transpiration and cell turgidity
→ temperature affects rate at which photosynthesis progresses (chemical
reaction)
→ 5 classes: terrestrial (forest, desert, grassland, tundra), marine and freshwater
(aquatic)
→ each biome has unique limiting factors, biodiversity and limiting factors
Tricellular Model of Atmospheric Circulation → explains differences in temp +
precipitation and how they influence the structure and productivity of different biomes.
→ latitude and atmospheric circulation are the primary factors affecting
insolation, temp, and precip.
→ the higher the latitude, the colder the temperatures
→ areas around equator receive the most insolation / unit area of earth
→ polar areas have more atmosphere to pass through, = more loss of
energy and cooler temps
→ Hot air heated at equator,
rises to form Hadley cell.
→ as air rises, it cools and
condenses, forming tropical
storms (explains tropical
rainforest at equator)

→ cooled air spreads and


descends - descending
creates high pressure and dry
areas at 30° (explains desert
biome)

→ air travels towards pole as


warm winds, when met with
cooler polar air at 60° it rises,
condenses and forms
precipitation (explains
temperate forest biome)
Different Biomes
→ distribution, structure, biodiversity, productivity (climate = temperature, precipitation,
insolation only)
DTPIPBS
Tropical Rainforest
→ distribution: band around equator, within tropics of cancer and capricorn
→ temperature: high and consistent yearly (~26°C)
→ precipitation: high (2500mm yr-1 +)
→ insolation: high, little to no seasonal variation. Provides year round growing season.
→ productivity: comprises ~ 40% of NPP for terrestrial ecosystems. High
photosynthesis and NPP caused by low latitude and ample direct sunlight.
→ biodiversity: high, up to 480 species / hectare, estimated half of world’s species in
rainforest canopy. High diversity due to high climate factors year round.
→ structure: stratified tree canopy, many niches. Only ~1% of light on canopy reaches
forest floor, canopy has highest NPP.
→ soil low in nutrients, majority stored in trees. High rates of decay maintain rates of
growth.
→ heavy rains can result in nutrients being washed away, which limits PP.
→ canopy usually protects soils from rainfall, but logging causes soils to be eroded
rapidly
***→ high light intensity → high temps → high NPP → high resources → high
complexity of habitats → high biodiversity ***

Temperate Forest
→ distribution: between 40° - 60°N of equator
→ temperature: cold winters, warm summers
→ precipitation: between 500-1500mm yr-1, determines whether temperate forests or
grasslands develop
→ insolation: varies according to tilt of Earth, limits growing season
→ productivity: lower compared to rainforests due to power temps and rainfall. Second
highest NPP in all biomes.
→ biodiversity: lower than rainforests, forests usually dominated by one species (90%
of forests may consist of only 6 species)
→ structure: less stratification and layering, less dense canopy, reduces species
diversity and complexity of niches
→ two types of trees (evergreen, deciduous - deciduous lose their leaves in winter)
→ forest floor leaf layer increases insulation and nutrients when it decays in warm
temperatures

Deserts
→ distribution: 30° N and S
→ temperature: high during day (45-49°C), low at night (10-0°C)
→ precipitation: low - 250mm yr-1, often very uneven
→ insolation: high (air is dry after leaving tropics)
→ productivity: lack of water limits photosynthesis and NPP, results in sparse
vegetation
→ biodiversity: xerophytic species (adapted to fluctuations in temp and scarcity of
water), reptiles most common vertebrates due to cold-blooded metabolism, cacti reduce
surface area for transpiration via spines
→ structure: soil can be rich in nutrients as there is no leaching, decomposition is low
due to lack of water

Tundra
→ distribution: high altitudes, the north polar region
→ temperature: low for majority of year, -50°C, warmer during 6 week period.
→ precipitation: low, water mainly stored in ice
→ insolation: short days, limited sunlight; almost 24h of sunlight during summer. Life
increases during summer.
→ productivity: very low due to variable light intensity, rainfall and temperatures
affecting race of photosynthesis
→ biodiversity: low, very few species adapted to cold conditions - large animals to
reduce heat loss
→ structure: low temperatures lead to low cycling of minerals → peat bogs form in
carbon sinks

Effect of Climate Change On Biome Distribution


→ increases in CO2 and other GHG increases mean global temperature, affecting
rainfall patterns
→ climate change alters biome distribution

Spatial and Temporal Changes in Communities


→ spatial changes occur along environmental gradients due to changes in altitude,
latitude, distance from sea
→ temporal changes occur as a community develops from early to later stages

Zonation
→ the arrangement or pattern of communities in bands in response to a change in
environmental factors over distance (eg. altitude, latitude, distance from shore)
Case Study: Rocky Shores
→ organisms high on shore exposed to air for long periods of time, have adapted to
withstand changes in salt concentration and temperature
→ organisms low on shore are covered by seawater, experience less variation in
temperature and salt concentration, with greater wave stress
Succession
→ change over time in an ecosystem involving pioneer / intermediate / climax
communities
→ each distinct community in the succession is a seral stage
→ succession explains how ecosystems develop from a bare substrate over time
→ lithosere (bare rock)
→ fresh water (hydrosere)
→ dry habitat (xerosere)

pioneer community: first stage of ecological succession - species able to withstand


difficult conditions
climax community: final stage of succession, more stable than earlier stages, in
equilibrium
primary succession: occurs on previously uncolonised substrate (eg. rock)
secondary succession: occurs in places where a previous community has been
destroyed. Faster than primary succession due to soil and seed bank.

→ one species changes the habitat they colonise and make it more suitable for new
species
→ lichens, moss etc. are good pioneer species as they photosynthesise and are
effective at absorbing water; they need no soil to survive. When they decompose they
form simple soil for other intermediate species.
→ newer species more able to trap light for photosynthesis and outcompete
previous species
→ final stage is climax community, species of larger biomass increase,
decomposers break down soil for other species etc.
1. Bare, inorganic surface
2. Colonisation by lichens, dead organic material results
3. Further weathering, beginnings of soil formation
4. Growth of small herbaceous plants, competition between pioneer species
5. Larger plants grow in more nutrient rich soil
6. Climax community dominated by shrubs and trees

Case Study: Primary Succession on Shingle Ridge


→ lichens and mosses pioneer species that photosynthesise and trap water on nutrient-
poor shingle
→ pioneer species trap particles blowing by and weather the rock
→ decomposition of pioneers results in a thin layer of soil
→ red fescue colonise area, roots trap soil and prevent erosion, pioneers begin to be
outcompeted
→ xerophytic plants eg. sea kale prevent water loss, nitrogen-fixing plants eg. rest
harrow increase soil nutrients
→ decomposition continues and allows growth of larger plants eg. shrub community of
bramble
→ climax community of temperate forest (oak/sycamore) develops, shrubs are replaced
by shade-adapted species like ferns

Case Study: Secondary Succession in Yellowstone, 1988


Fires in Yellowstone National Park wiped out many aspects of the park’s forest - some
fires burned soil and ground biomass, some burned the canopy. Fires burned for
several months.
→ recovery began almost immediately with herbaceous fireweed as a pioneer species
→ lodgepole pines, though a climax species, are serotinous, allowing fast regeneration
in burned areas
→ comprising 80% of park’s forests, they only release seeds when high
temperatures eg. fires create favourable open canopies for seedling establishment
→ aspen, wildflowers had an increase in productivity as nutrients were released from
forest litter during burning
→ soil depth only charred to 14mm, leaving diverse root systems unharmed
→ minimal overall loss of wildlife from fires, only browsers eg. moose populations
declining
→ other k-strategists eg. elk, bison, deer rebounded due to rapid plant growth,
birds able to find ants and worms easily in newly uncovered soil
→ as pine, larkspur, aspen and other climax species increased, other animals began
migrating to the area
Changes Through Succession → productivity, mineral cycles, diversity etc. all change
during succession.
GPP → pioneer communities have low GPP because of the low density in producers.
Climax communities have high GPP as there is an increased consumer community.
NPP → high in pioneer communities as community respiration is low (low # organisms).
High NPP means biomass is continuing to accumulate. Approaches zero in climax
community as GPP is balanced by increased respiration
Production/Respiration Ratio: when production = respiration, P/R = 1
when P/R is greater than 1, biomass increases
when P/R is less than 1, biomass depletes
Pioneer communities have low GPP and high NPP due to lowered rates of respiration,
P/R is greater than 1.
Climax communities have high GPP but increased respiration, therefore low NPP, P/R
approaches 1.
Diversity → pioneer communities have low biomass, low species diversity and few
niches. Climax community have complex niches and more biomass; the increase in
niches leads to habitat and species diversity. Complex interactions result in a steady-
state equilibrium
Mineral Cycling → pioneer communities have open systems, carbon/nitrogen cycles
easily. Climax communities have closed systems as the role of decomposition in cycling
nutrients from soil to biomass increases.

Climax Communities → a community of organisms that is in steady-state equilibrium


with natural environmental conditions. It is the endpoint of ecological succession.
→ greater biomass, high species, habitat and genetic diversity
→ favourable soil conditions and structure (deeper, greater water
retention/aeration)
→ more k-strategist organisms (taller plants etc)
→ greater community complexity, resilience and stability
→ climax communities are more stable as greater energy pathway and biodiversity
means species can turn to alternate food sources in the event of a shock; nutrient
cycles are self sustaining.

r and K-strategist Species → species can be determined by how rapidly they produce,
the degree of parental care, and the type of environments they are most suited to.
*species that lie in between are C-strategists.

r-strategists K-strategists

→ opportunistic, fast rates (r) of increase, → slow growing organisms limited by carrying
inhabit rapidly changing + unpredictable capacity (K). inhabit stable environments/later
environments (ie. pioneer communities), seral stages (ie. climax communities). Offspring
produce many small offspring that mature mature slowly and species is vulnerable to high
quickly. Little to no parental care. death rates.

 Colonisers  Dominant species


 Highly adaptable  Specialist, susceptible to change
 Rapid growth/development  Slow growth/development
 Short lifespans  Long lifespans
 Small size, many offspring  Large size, few offspring
 Early, high reproduction  Delayed reproduction
 Type III survivorship  Type I or II survivorship
 Suited to pioneer communities  Suited to climax communities
 Continuous J-curve population  S-population curve

r- and K- Selection Theory → 1. In disturbed habitats, natural selection favours


individuals with high reproductive rates over those with slower reproductive rates, but
better competitive ability, as they can respond quickly. 2. In predictable environments,
species that maximise natural resources and produce few young are more favoured.
Survivorship Curves → show changes in survivorship of a species’s lifespan
→ r-strategists produce large numbers of offspring
to colonise new habitats quickly and make use of
limited resources
→ most individuals die very young, those that
survive live long

→ K-strategists produce small numbers of


offspring to increase their survival rate and live in
climax communities.
→ almost all individuals survive for potential
lifespan and die roughly simultaneously
Impact of Human Activities On Succession → interrupted succession = plagioclimax.
Human disturbance can halt the process of succession and divert it so a different stable
state other than a climax community is reached.
→ activity modifies the ecosystem (eg. use of fire, grazing, agriculture,
deforestation, overfishing); depending on the resilience of an ecosystem changes may
be more or less permanent.
Eg. deforestation of tropical rainforest biome
→ increased demand for meat demands land for cattle ranching and agriculture,
90% of the reason why land is deforested in the Amazon. Results in habitat destruction
and loss of climax community.
→ replacement with agricultural system affects global biodiversity, weather,
sedimentation patterns
→ CO2 released returns to atmosphere

2.5 Investigating ecosystems


Ecosystems can be better understood to the investigation and quantification of their
components.
Standardised methods and studies allow ecosystems to be modelled, monitored and
evaluated over time.
Identifying Organisms in Ecosystems → can use a dichotomous key, a stepwise tool
for identification of unfamiliar organisms.
Limitations: keys examine physical rather than behavioural characteristics
→often use technical terms only understood by experts
→ may not be a key for the type of organism available
→ some features of organisms are difficult to identify in wild
Measuring Abiotic Components of Ecosystem
MARINE → salinity, pH, dissolved oxygen, wave action, temperature
FRESH WATER → turbidity, dissolved oxygen, pH, flow velocity, temperature
TERRESTRIAL → light intensity, slope, wind speed, temperature, mineral content,
drainage, soil moisture
**measurements must be repeated to increase reliability of data. Errors in sampling may
result in an inaccurate representation of an environment.**
Light → measured using a light meter, meter must be held at a fixed height and at the
same angle.
Limitations: cloud cover/other changes in light means value must be taken at
same time of day and same atmospheric conditions.
Temperature → measured using electronic thermometer with probes, soil / air / water.
Data loggers allow long term fluctuations to be taken into account.
Limitations: if thermometer depth is not consistent, problems arise
pH → using pH meter or data logger. Values change depending on fresh/salt water,
probe must be cleaned between readings.
Turbidity → measured using secchi disc lowered into water - cloudy water = high
turbidity, clear water = low turbidity. Indirectly corresponds to rate of photosynthesis.
Limitations: Sun glare on water, subjective nature of eyesight.
Measuring Biotic Components of Ecosystem
Estimating Abundance of Organisms
motile → pitfall traps, nets, light interception traps, small mammal traps
non-motile → quadrats, point frames
Abundance → the relative representation of a species in an ecosystem. Can be done by
directly counting # organisms (non-motile) or indirectly estimating using Lincoln index
(motile).

Lincoln Index → estimates total population size of a motile animal - ‘capture-mark-


release-recapture’ technique.
[indirect]
N1 = number in first sample
N2 = number in second sample
Nm = number caught in second sample that were marked
Limitations: animals may move in/out of sample area, seasonal variations may
affect population size. Density of a species’ population may differ in different habitats.

Direct Methods For Estimating Motile Animal Abundance → actual counts+sampling to


give a relative abundance of different animals in a sample.
Limitations: sample size and collection methods must be standardised, some
animals may remain hidden / unsampled.
Eg. canopy fogging to knock insects into collection trays, number can be
extrapolated
Quadrats [Non-motile] → limits the sampling area when measuring non-motile
organisms. A square frame.
→ random sampling: if habitat is same throughout, quadrats should be located at
random.
→ stratified random sampling: if two/more areas of habitat, results from both
areas should be obtained.
→ systematic sampling: if area occurs along environmental gradient, quadrats
should be placed at set distances along a transect. Continuous sampling occurs
across the whole length of the transect.
Limitations: often subjective - mistakes easy to make in identifying, difficult to
measure plant colonies

Population density → (total number of a species in all quadrats)


(area of one quadrat) x (total number of quadrats)
Percentage cover → estimate of an area in a quadrat covered by the organism in
question
Percentage frequency → number of actual occurrences
Number of possible occurrences
Abundance Scales → DAFOR - Dominant, Abundant, Frequent, Occasional, Rare

Estimating Biomass of Trophic Levels


Biomass → a measurement of dry mass (mass - water content), indicates the total
energy within a living organism. The greater the mass, the greater the amount of
energy. Measured in g / m-2.
→ mass of one/the average of a few organisms x total number of organisms
Limitations: involves killing living organisms. Biomass of root systems difficult to
measure.
Species Richness and Diversity
Richness → number of species in a community
Diversity → a species’ relative abundance in a given area.
Simpson’s Diversity Index
D → diversity

N → total number of organisms of all species found


n → number of individuals of a particular species
→ The higher the value of D, the greater species diversity and the more stable the
ecosystem/population. Only useful when comparing similar habitats or ecosystems, as
diversity is relative, not absolute.
Measuring Changes in Ecosystems
Changes Along Environmental Gradient
→ grid quadrat, point quadrats
→ line transect, belt transect (continuous or interrupted)
→ transects should be controlled for atmospheric conditions and repeated to ensure
reliability.

Changes Due to Human Activity → eg. landfills, eutrophication, oil spills, change in land
use, overexploitation
→ can use: diversity index, measure of abiotic variables, soil erosion (pH etc),
lincoln index on various disturbed and undisturbed sites
Topic 3: Biodiversity and conservation
3.1 An introduction to biodiversity
Biodiversity → the variety of life on earth (ie. species, habitat and genetic diversity).
→ often used to evaluate the health and complexity of an ecological area
Species Diversity → the variety of species per unit area - includes # species present
and their relative abundance.
→ higher the species diversity = greater ability to support different
niches
→ measured in richness (number of species) and evenness (relative
proportions)
→ community with high evenness has a similar abundance of all
species; low evenness means one or few dominant species, indicating lower
complexity.
→ Simpson’s Diversity (D) can be used to compare communities:
→ low D can indicate pollution, eutrophication, recent disturbances
→ high D suggests a stable and mature site
Habitat Diversity → the range of different habitats in an ecosystem or biome, indicating
niche variety
Genetic Diversity → the range of genetic material present in a population of a species
→ genes: sections of DNA found in the nucleus of all cells
→ gene pool: different types of gene found within every individual of a
species. A large gene pool leads to high genetic diversity; low genetic diversity makes
species prone to extinction.

Overview of Biodiversity → conservation of habitat diversity leads to conservation of


species and genetic diversity - diverse habitats have a diverse variety of species, which
tend to have different genes

Conservation of Biodiversity → conservation aims...


 to protect habitats, ecosystems and species from human disturbances
 to slow the rate of extinction caused by unsustainable exploitation of natural
resources
 to maintain biotic interactions between species
→ the quantification of biodiversity is crucial so high biodiversity can be identified,
explored and conserved
→ human activity causes disturbance that can remove an ecosystem from steady-state
equilibrium
→ measuring biodiversity vital to identify endemic species and their habitats that should
be protected

3.2 Origins of biodiversity


Biodiversity Arising From Evolutionary Processes
Evolution → the cumulative, gradual change in the genetic composition of a species
over many successive generations, giving rise to a species different from the common
ancestor.
→ evidence for evolution can be found in fossil records
Natural Selection → an evolutionary driving force - the selection of beneficial biological
variations best suited to survival in a given environment.
1. Species over-reproduce
2. Genetic variation (genetic diversity / mutation) occurs randomly in a species
3. Natural variation makes some individuals fitter for survival than others
4. Selection occurs as fitter individuals have a survival advantage and are more
likely to live to reproduce
5. Offspring of fitter individuals more likely to inherit the advantageous gene,
adaptation more likely to be passed to subsequent generations
→ mutations that give advantages are selected for - the individual will have better
competitive advantages and will survive long enough to pass on the trait
→ mutations that give disadvantages are selected against - mutations that make and
individual less suited to the environment make survival more difficult
→ variation arises randomly and can either be beneficial, damaging or have no impact
on survival

Isolation and Formation of New Species


Reproductive Isolation → must occur between populations so genes cannot be
exchanged between them. If the environments of isolated populations are different,
natural selection will form new species.
Speciation → the formation of a new species when populations of a species become
isolated and evolve differently.
Geographic Isolation → a physical barrier that causes a population to become
separated. Without this, new species cannot form as genes from populations continue
to mix.
1. Two populations of one species interbreed - gene flow occurs
2. Populations separated by geographical barrier and cannot interbreed; species
develops own variations as gene flow is interrupted
3. Two separate species develop in response to different selection pressures -
environmental changes produce new challenges to species and natural selection
takes place
4. Even without a barrier, species are now genetically distinct and can no longer
interbreed
→ during the ice ages, a fall in sea levels (decrease in temperature and water locked in
glaciers) lead to a land bridge forming between Alaska and Siberia. When sea levels
rose, the areas became isolated again.

Plate Tectonics
Tectonic Plates → have moved throughout time, creating physical barriers and land
bridges that lead to gene pool isolation and speciation
Plate Tectonics: the movement of plates → move parallel to, be pushed under or
collide with each other.
→ during Palaeozoic and Mesozoic eras, land mass existed as supercontinent
Pangaea
→ Pangaea later split to form Gondwana and Laurasia
→ the different species found on different land masses can be explained
by the movement apart and formation of land bridges of these plates - plate movement
across Earth allows new habitats to form.

Plate Boundary Types


Divergent Boundary → constructive - new crust being formed
Convergent Boundary → destructive - crust being destroyed
Transform Boundary → conservative, friction is created

Continental-Continental Divergent Boundary


→ continental plates diverge and form rift valleys
→ deep lakes/seas can form in the gaps, the creation of new aquatic habitats drives
speciation. Magma from rift can create new land, giving more opportunities for evolution

Oceanic-Continental Convergent Boundary


→ subduction of denser oceanic crust beneath less dense continental crust
→ leads to new island arcs and mountain areas forming as magma rises from
subduction zone. Often volcanic.

Oceanic-Oceanic Convergent Boundary


→ oceanic crust subducted beneath oceanic crust
→ rising magma causes volcanic activity and new islands to form, providing new
habitats and driving speciation

Continental-Continental Convergent Boundary


→ continental plates collide and increase plate thickness
→ new mountain ranges are formed - habitats at different altitudes contribute to
biodiversity

Mass Extinctions
Mass Extinction → a period where at least 75% of total species on Earth were wiped out
at the same time. Species disappear in a geologically short time period due to abiotic
phenomena.
→ all mass extinctions have resulted in an increase in biodiversity - the
large-scale loss of species left new opportunities for surviving populations to undergo
adaptive radiation and fill different niches
Ordovician - Silurian Extinction
→ 439 million years ago, killed 86% of all species
→ causes: drop in sea levels as glaciers formed; rise in sea levels as glaciers melted
Late Devonian Extinction
→ 364 million years ago, killed 75% of all species
→ causes: global cooling followed by global warming
Permian - Triassic Extinction
→ 251 million years ago, killed 96% of all species
→ causes: debated - some believe flood volcanism destroyed algae and plants and
reduced oxygen in sea. Others believe tectonics and movement of Pangaea may have
lead to environmental changes on the landmass, decreasing the quantity of shallow
seas and exposing isolated areas and organisms to increased competition.
End Triassic Extinction
→ 199 million years ago, killed 80% of all species
→ causes: flood volcanism (lava) erupting from opening in Atlantic, leading to climate
change
Cretaceous Tertiary Extinction
→ 65 million years ago, killed 76% of all species
→ causes: impact of miles wide asteroid created crater in Gulf of Mexico - dust thrown
into atmosphere by impact could have reduced sunlight, limiting productivity and
dropping temperature. Plate tectonics and re-arrangement of world’s land masses could
have resulted in climatic changes that deteriorated habitats.

3.3 Threats to biodiversity


Number of Species on Earth → estimates vary considerably, as they are based on
models and limited classification data. As a result, many habitats and groups are
significantly under-recorded.
→ current consensus at ~9 million species
→ most described species are larger organisms, mostly animals,
easier to study
→ most diverse groups (insects, bacteria, fungi) are most difficult to
study
→ of 1.8 million described species, 1% are vertebrates, yet they are
the most studied
Rates of Species Loss → ~30,000 - 60,000 species a year, 100 - 100,000x greater than
background extinction rate
→ existing species must be identified and named in order to understand
extinction
→ humans contribute greatly to diversity loss, eg. mammals 1 every 200 years
should be lost, yet 90 species extinct in past 400 years.
Causes of Species Loss
Natural Causes → typically hazard events eg. volcanoes, ice ages, drought
Human Causes → habitat destruction, invasive species, pollution, overharvesting,
hunting all reduce diversity

Habitat Destruction → habitat degradation, fragmentation and loss


→ agricultural practices destroy native habitats and replace them with less
diverse monocultures
→ non-specific pesticides often used in agriculture and wipe out both native and
pest species
→ mining activities often destroy forests containing endangered species
→ plantation crops replace natural ecosystems
→ invasive species compete with endemic species, leading to extinction of native
species
→ pollution eg. chemicals, plastics, oil spills damages habitats and kills
organisms
→ overharvesting and hunting - animals are hunted for food, medicines etc
Threats to Tropical Biomes
Tropical Biomes → 5.9% of earth’s land surface, some of the most globally biodiverse
areas. Unsustainable exploitation of these areas results in loss of biodiversity and ability
to perform ecological services.
→ ~1.5 ha of tropical rainforest is lost every 4 seconds.
→ includes rainforests, coral reef, mangrove forests
→ complex structure and warm, stable climate increase productivity and allow many
niches to be supported.
→ perform crucial ecosystem functions eg. soil erosion prevention, controlling water
cycle and weather, carbon sequestering etc.

Human Activity and Disturbance


→ deforestation and forest degradation driven by increasing demands for timber, beef,
land for crops, and biofuels impact rainforests
→ palm oil plantations replace a diverse climax community with a monoculture
→ large timber removal means fast growing species block out light for slower K-
strategists
→ the rate of loss of biodiversity varies depending on the ecosystems present,
protection policies available, environmental viewpoints and stage of economic
development
Example: Oil Palm Plantations
→ second most traded vegetable crop, comprises 90% of exports in Malaysia and
Indonesia
→ projected 16.5 million hectares of oil palm in 2020 in Indonesia
→ 6.5 million hectares of oil palm in Borneo estimated to have destroyed 10 million ha
of rainforest

Conflict Between Exploitation, Sustainable Development and Conservation


→ ecosystem exploitation often prevalent in LICs due to their need to provide income
for local economies; HICs can preserve ecosystems as they do not rely on them for
income
→ argument to preserve biodiversity is different in LICs, where most tropical
biomes are found
→ for sustainable development to occur in LICs towards becoming MICs, balance
between using land for income and conservation must be struck using local and
governmental support.

Determining Conservation Status


The Red List → published by IUCN (international Union of Conservation of Nature),
seeking to conserve genetic diversity through awareness and providing a basis for
conservation decision at local and global levels.
→ to identify species requiring conservation
→ to identify species with conservation status concern
→ to catalogue species facing a high risk of global extinction
Factors Determining Red List Conservation Status
→ population size: smaller populations have low genetic diversity, reducing their ability
to adapt to changes
→ trophic level: top predators are highly sensitive to disturbances at lower trophic levels
→ reduction in population size: can indicate species is under threat
→ geographic range: species occupying a restricted habitat are likely to be wiped out
→ degree of specialisation: species with specific diet or habitat are threatened when
their habitat is threatened
→ distribution: species in a small area are more threatened than those distributed
widely
→ reproductive potential and behaviour: k-strategists take longer to recover from
reductions in population
→ degree of habitat fragmentation: leads to islands within ecosystems and increases
impact of edge effect
→ quality of habitat: poor quality habitats can support less species

Case Study: Extinct - Passenger Pigeon


Description → located in NA, once the most abundant bird, travelled in 3-5 billion flocks.
Known for migratory habitats and was adapted to reach speeds of 100 km/h.
Ecological Role → determined forest composition - forests dominated by white oaks as
pigeons ate seeds of read oaks in spring, leaving white oaks to germinate in fall
Pressures on Species → 19th century Europeans commercialised pigeon meat, mass
hunting began depleting populations in 1800 and intensified in 1890. Last bird was shot
in 1901.
Consequences of Extinction → increase in # of white footed mouse as competition for
food decreased, increase in mice linked to increase of lyme disease as they are hosts
Species Restoration Strategies → genetic engineering to revive species

Case Study: Critically Endangered - North Atlantic Right Whale


Description → most endangered whale in world, currently 300 or less estimated. A
migratory species.
Ecological Role → baleen whale, important predator of krill and other plankton, prevents
populations from skyrocketing.
Pressures on Species → slow moving nature made it an attractive target for 19th
century hunters - 10,000 whales killed. High blubber content yielded whale oil and made
carcasses float on surface of water. Current threats are vessel strikes and entanglement
in fishing equipment - extinction estimated in 190 years.
Species Restoration Strategies → International Convention for Regulation of Whaling in
1935 banned hunting of right whales, Marine Mammal protection Act established in
1972. Climate change control can assist in managing changes in right whale’s food
availability (zooplankton).

Case Study: Improved By Intervention - Gray Wolf


Description → very widely distributed mammals, found primarily in Northern
Hemisphere biomes. Live in packs of up to 36, have a nomadic phase and stationary
phase depending on when pups are reared.
Ecological Roles, Consequences of Extinction → apex predators and play a crucial role
in regulating prey populations. Remove weaker individuals to form a superior breeding
pool. Extinction would result in an increase in prey and depletion of primary producers.
Pressures On Species → hunting increased mid 20th century due to human fears of
livestock depleting, thousands of wolves were hunted and in some regions entirely
exterminated. Continued competition with humans for game species, as well as threat of
habitat fragmentation.
Species Restoration Strategies → hunting outlawed in 1970s, populations began to
increase after this year. Legal protection and recolonisation of former habitats stabilised
populations, now classified as ‘least concern’.

Case Study: Threats to Area of Biological Significance → Great Barrier Reef


→ extremely diverse coral reef stretching along 2300 km of coast with high productivity.
→ 1500 species of fish, 359 types of coral, 6 of the 7 threatened species of turtle
→ tourism and fishing generates $1 billion AUD per year
→ crucial to Aboriginal culture and spirituality
Human Threats to GBR
→ tourism: though contributes to local economy, coral is fragile and tourists often break
them off for souvenirs
→ overfishing: can disrupt balance of species in food chain; seafloor trawling leads to
unintentional capture of other species and damage to floor
→ land use: changes from subsistence agriculture to large scale farming needing
fertilisers and pesticides - runoff causes increase of nitrogen pollution by 3000%
→ deforestation: coastal mangrove removal leads to increase in sedimentation by
800% as mangroves typically filter sediment. Pollution clouds water and reduces
productivity.
→ global warming: increase in sea temperature has bleached 60% of the reef in
2002, causing loss of biodiversity
Natural Threats
→ climate change increases cyclones and El Niño which cause structural
damage to coral
→ crown-of-thorns starfish prey on polyps (increased by removal of predators by
humans)
Consequences
→ continued threats to reef can make damage irreversible
→ if system reaches tipping point, it will not be able to recover
→ the loss of biodiversity and ecosystem services (eg. defence against
sedimentation) leads to a reduction in the value of the natural capital

3.4 Conservation of biodiversity


Arguments to Preserve Biodiversity → can be aesthetic, ecological, economic,
ethical and social
→ goods are easier to quantify than indirect values
and services
Aesthetic Reasons → species/habitats are pleasant to look at
Ecological Reasons → habitats with endemic species must be preserved
→ higher biodiversity = more resilience & stability = continued
ecosystem services in future
→ species extinctions have knock-on effects throughout food webs
Economic Reasons → value of ecotourism, genetic resources & commercial
considerations of capital
→ genetic diversity can allow improvements in crops etc. to be made
- a genetic resource
→ commercial resources (capital as medicines etc. or successful
tourism)
Ethical Reasons → intrinsic value of a species - all have a right to survive, responsibility
to protect for future
Social Reasons → ecosystems provide homes, livelihoods and cultural cohesion for
indigenous peoples

Reasons to Conserve Rainforests (consider someone who relies on the forest vs. an
outsider)
→ aesthetic: habitats & species pleasant to look at
→ ecological: life support functions - stabilising soil erosion, regulating temp and hydro
cycles, sequestering carbon, maintaining atmospheric balance of CO2
→ economic: natural capital (fuel, food, medicine, ecotourism) can bring in income
→ ethical: forests have intrinsic value & a necessity to be protected for future
generations
→ social: spiritual, cultural, religious value to local indigenous communities

Conservation Organisations
→ can be international, governmental or non-governmental, with varying levels of
success when approaching conservation due to their use of media, their speed of
response, diplomatic constraints, finances and influence.
NGOs eg. Greenpeace, WWF IGOs eg. United Nations Environment
Programme

→ not run by, influenced or funded by → established through agreements to allow


governments global cooperation between governments
→ field based, gathering information to support → information from paid scientific research
their claims → less controversial, more conservative
→ radical to spread their message and be approach
heard
Use of media → cooperates with media to
Use of media → gain coverage through effectively communicate policies and decisions
protests & campaigns (charismatic species), to the public
putting pressure on governments
Speed of response → fast, members already Speed of response → slow (bureaucratic),
at consensus regarding course of action decisions directed by governments & require
Agenda → using public pressure and lobbying consensus
to influence government policies and Agenda → provides guidelines and
legislation implementing international conservation treaties
Funding → from private donations Funding → budget from national economies
Political pressures → environment focused, Political pressures → can be
working towards idealistic conservation politically/economically driven rather than
strategies environmental
→ both provide information to educate public on environmental issues, publishing reports and
data
→ both encourage partnerships between nations and organisations to conserve ecosystems
→ both monitor species and conservation areas at local, regional and global scales

International Conventions on Biodiversity


→ conventions aim to create collaboration between nations for biodiversity conservation
IUCN (International Union for Conservation of Nature)
→ founded in 1948, concerned with importance of conservation of resources for
sustainable development
→ established Red List and World Conservation Strategy with the UNEP and WWF
→ World Conservation Strategy:
→ stresses importance of making the users of natural resources the guardians of
those resources
1. maintaining essential life support systems and ecological processes
(climate, water, soils)
2. Preserving genetic diversity
3. Using species and ecosystems sustainably
Conservation Approaches
→ 3 main types: habitat conservation, species based conservation, mixed approach
Local vs. Global → when problems are global, international cooperation is often useful
and can motivate governments to take action and honour their commitments (eg. global
warming)
→ IGOs have the funding to mobilise and coordinate a united,
transboundary response
→ global summits and conventions play a vital role in setting targets and
shaping action (eg. 2000 Millennium Summit)
→ when problems are local, local populations should be involved in
providing solutions
In-Situ Conservation → the conservation of species in their natural habitat
→ endangered animals & their habitats are protected, conserving
many other species
Ex-Situ Conservation → the preservation of species outside their natural habitat
→ in botanic gardens, zoos, with captive breeding programmes
→ focuses on vulnerable species
→ aims to attract interest & public pressure in conservation, more
funding
Habitat Conservation
→ buffers to human influence, area, edge effects, shape, corridors must be considered
(BAESC)
Buffer Zones → successful areas are surrounded by buffers to minimise disturbances
from outside influences
Area → larger conservation areas preferable to several smaller ones
→ they include more habitats, promoting large population sizes especially among
large vertebrates
→ the best indication of reserve success is the population of individual species
→ several reserves allow habitats to guard against possible effects of fire etc.
that could threaten species
Edge Effects → changes in abiotic factors at the edge of a protected area (eg. temp,
humidity, wind)
→ edges attract species not found deeper within the reserve, leading to
competition and an overall reduction in biodiversity
→ larger habitats reduce the perimeter relative to the area, minimising the edge
effects
Shape → circles are the ideal shape as they have the lowest edge effects
→ long, thin reserves have large edge effects
→ depending on location of habitats, parks are usually irregular
Corridors → close, clumped conservation areas with corridors are better than
fragmented areas
→ animals can migrate, disperse and recolonise in the event of a disturbance
→ corridors allow genetic flow through migration and seasonal movements,
reducing barriers to movement like roads and car collisions
→ strengths: protects whole ecosystem and complex relationships, ensuring long-term
survival of species
→ allows research to take place in intact habitats, enhancing understanding of
biodiversity
→ preserves many niches, prevents hunting and other disturbances
→ ecotourism and education raises awareness, generating profits to fund
conservation programs
→ species that haven’t been discovered yes can still be protected
→ limitations: requires considerable funding and continuous protection to ensure
minimal disturbance
→ difficult to establish due to conflicting EVSs
→ areas can become islands and lose biodiversity due to reduced gene flow and
edge effects
Species Based Conservation
→ CITES, captive breeding, flagship species, keystone species
CITES (Convention On International Trade in Endangered Species
→ established in 1973, int’l agreement regulating trade in endangered species
→ while trade in plants & animals is worth billions, it reduces wild populations &
exhausts species
→ strengths: CITES currently protects 35,000 species, with countries becoming
voluntary members (monitoring trade, extracting fines to discourage trade) - works
transboundary
→ ensures the overall sustainability of international wildlife trade
→ appendix I (endangered & illegal trade)
→ appendix II (non-endangered & sustainable trade)
→ legally binding - participating countries must implement the convention
→ limitations: species can be re-classified as appendix II
→ penalties and fines may not be severe enough to deter smugglers
→ CITES lacks financial mechanisms for implementation at the national
level
→ interpretations vary between countries
→ does not replace national legislation; countries must make their own
laws to apply CITES
Captive Breeding & Reintroduction Programmes (Zoos)
→ facilities where animals are housed & breeding can take place
→ development level of country must be considered (can the programme be supported
in the long term)
→ in situ or ex situ - habitat threatened species must be ex situ, species with local
involvement can be in situ
→ ‘five freedoms’ - freedom from thirst, hunger, physical discomfort, injury & disease,
fear & distress, freedom to express normal behaviours
→ strengths: able to educate public about need for conservation by allowing them to
empathise with wildlife
→ captive breeding can be used to increase population sizes of
threatened species
→ offers a temporary protected area to maintain genetic diversity, animals
reintroduced later
→ allows research to be conducted
→ limitations: captive animals have trouble re-adapting to the wild
→ not all species breed easily in captivity
→ habitats are very different from natural environment, species isolated
from their typical niche - zoos don’t treat the ecosystem as holistic
→ ethical issues surrounding using captive animals for profit
→ popular species are not necessarily the ones at the most risk
Flagship Species
→ charismatic species designed to appeal to the public and protect other species in an
area
→ iconic species allow conservation to catch attention and raise necessary
funds for initiatives
→ strengths: money can be raised for conservation of other threatened but less
appealing species
→ preserving the habitat of flagship species preserves other organisms in
the same habitat
→ limitations: favours charismatic species at the expense of less publically attractive,
more endangered species
→ does not guarantee habitat conservation - species may be preserved in
zoos instead
Keystone Species
→ species vital for conserving the function of an ecosystem
→ limitations: species difficult to identify due to complexity of ecosystems
→ establishing protected areas rather than conserving individual species
preserves complex interrelationships
→ keystone eg. agouti in SA feeding on the Brazil nut tree breaks open nut pods,
burying seeds in forest floor and allowing them to germinate. Other organisms
dependent on trees for food + shelter.
Mixed Approach
→ combining both in-situ and ex-situ conservation (protected areas & zoos) is often
most effective
→ eg. Giant Panda Conservation in Beijing Zoo
→ flagship species
→ on appendix I of CITES
→ successful breeding established in 1960s through artificial insemination and
breeding
→ 56 conservation areas and nature reserves established
→ Chengdu Panda Base does both in and ex situ conservation, emphasising
wildlife research, captive breeding and educational tourism

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