ZOOLOGY : CHAPTER 1 amazing variety of color patterns,
Zoology habitats, and body forms. (a) This
➢ comes from the Greek words zōion, dogtooth cichlid (Cynotilapia afra) is meaning “animal”, and logos, meaning “the native to Lake Malawi in Africa. The study of” which makes Zoology the study of female of the species broods, animals. It is one of the broadest fields in developing eggs in her mouth to all of science because of the immense protect them from predators. (b) The variety of animals and the complexity of the fontosa (Cyphontilapia fontosa) is processes occurring within animals. native to Lake Tanganyika in Africa. SPECIALIZATIONS IN ZOOLOGY: A Scale-Eating Cichlid. Scale-eaters ∙ ANATOMY -Study of the structure of (Perissodus microlepis) entire organisms and their parts ∙ attack from behind as they feed on ∙ CYTOLOGY -Study of the structure scales of prey fish. Two body forms and function of cells are maintained in the population. In ∙ COMPARATIVE GENOMICS AND one form, the mouth is BIOINFORMATICS - Study of the asymmetrically curved to the right structure, function, and evolution of and attacks the prey’s left side. The the genetic composition of groups of second form has the mouth curved animals using computer-based to the left and attacks the prey’s computational methods right side. Both right- and left-jawed ∙ ECOLOGY -Study of the interaction of forms are maintained in the organisms with their environment ∙ population and prey do not become EMBRYOLOGY - Study of the wary of being attacked from one development of an animal from the side. Perissodus microlepis is fertilized egg to birth or hatching endemic (found only in) to Lake ∙ GENETICS - Study of the Tanganyika. A male with its brood of mechanisms of transmission of young is shown here Lakes Victoria, traits from parents to offspring Kivu Tanganyika, and Malawi ∙ HISTOLOGY - Study of tissues ∙ ∙ These lakes have cichlid populations Molecular biology - Study of that have been traced by zoologists subcellular details of structure and to an ancestry that is approximately function 200,000 years old. Cichlid ∙ PARASITOLOGY - Study of animals populations originated in Lake Kivu that live in or on other organisms at and Lake Tanganyika and then the expense of the host spread to the other lakes. ∙ PHYSIOLOGY- Study of the function of ZOOLOGY: AN EVOLUTIONARY PROCESS organisms and their parts EVOLUTIONARY PROCESS - the process that produces common or rare ∙ SYSTEMATICS - Study of the characteristics among species that result in classification of, and the the diversity of life. evolutionary interrelationships ∙ Animals share a common among, evolutionary past and evolutionary SPECIALIZATIONS IN ZOOLOGY BY forces that influenced their history. TAXONOMIC CATEGORIES ORGANIC EVOLUTION ∙ ENTOMOLOGY - Study of insects ∙ ∙ is change in the genetic makeup of HERPETOLOGY - Study of populations of organisms over time. ∙ amphibians and reptiles the source of animal diversity, and it ∙ ICHTHYOLOGY - Study of fishes ∙ explains family relationships within MAMMALOGY -Study of mammals ∙ animal groups. ORNITHOLOGY - Study of birds ∙ Charles Darwin published a book entitled, PROTOZOOLOGY -Study of protozoa "On the Origin of Species" in 1859, stating Cichlids a convincing evidence of evolution. He ∙ Cichlids of Africa exist in an proposed a mechanism that could explain evolutionary change: NATURAL 4. ERASMUS -DARWIN SELECTION. ∙ A physician and the grandfather of NATURAL SELECTION Charles Darwin ∙ the process by which organisms better ∙ He was interested in questions of suited for adaptation to their origin and change. environment survive, while those ∙ He -believed in the common that are poorly suited to their ancestry of all organisms. environment do not. 5. JEAN BAPTISTE LAMARCK ∙ Selection may occur as a result of ∙ His theory was based on a widely differences in survival, in fertility, in accepted theory of inheritance that rate of development, in mating organisms develop new organs, or success, or in any other aspect of modify existing organs, as needs the life cycle. arise. TYPES OF EVOLUTIONARY PROCESSES ∙ Similarly, he believed that disuse ∙ GENETIC DRIFT - change in resulted in the degeneration of frequency of an existing gene variant organs. The “need” was dictated by in the population due to random environmental change and that chance. change involved movement toward ∙ GENETIC MUTATION - alteration in perfection. the sequence of the genome of DNA. ∙ ∙ The idea that change in a species is GENE FLOW - transfer of genetic directed by need logically led material from one population to Lamarck to the conclusion that another population. species could not become extinct— Other proponents of evolutionary they simply evolved into different perspective before Charles Darwin ∙ species. The idea of evolution did not ANIMAL MORPHOLOGY, PHYSIOLOGY, originate with Charles Darwin. AND FUNCTION 1. -EMPEDOCLES ∙ Understanding how the diversity of ∙ He believed that all organisms, animal structure and function including humans, went through arose is one of the many challenges the same evolutionary process faced by zoologists. ∙ For him, past natural selection is EVOLUTIONARY RELATIONSHIPS ∙ Are responsible for what we are today. all relationships between two different living organisms, including organisms that are related through the humans, went through the same global process of evolutionary process evolution. 2. ARISTOTLE ∙ Evolution not only explains why ∙ He describe evolution as a“ladder of animals appear and function as they life,” where simple organisms do, but also explains family gradually change to more elaborate relationships within the animal forms kingdom. 3. GEORGES-LOUIS BUFFON ANIMAL NAME CLASSIFICATION ∙ ∙ He observed structural variations in Binomial Nomenclature - a system of particular organs of related classifying and naming animals into animals. two parts: ∙ He attributed change in organisms to a. genus name - origin the action of the environment. b. species epithet - name ∙ He believed in a special creation of - ex. panthera tigris ( adopted from the species and considered change as plant naming system proposed by Carl Von being degenerate—for example, he Linnae) described apes as degenerate - ECOLOGICAL PERSPECTIVE humans Population ∙ Global overpopulation is at the root of virtually all other environmental single-celled organisms that don't have problems. a nucleus and membrane ∙ As the human population grows, the bound organelles. disparity between the wealthiest and Under prokaryotes, there are 3 domains of poorest nations is likely to increase. ● World life which are Arachea, Eubacteria and resources Eukaryota. ∙ Human overpopulation is stressing ∙ Eukaryotic cells (true nucleus). This world resources. type of cell on the other hand has a ∙ new technologies continue to nucleus that contains DNA and increase food production, most food membrane bound organelles. is produced in industrialized Additionally, the difference of countries that already have a high prokaryotic to eukaryotic is that the per-capita food consumption. latter has microfilaments and ∙ Deforestation of large areas of the microtubules in the cytoskeleton world results from continued whose function is to give shape and demand for forest products, fuel, allows the cell to move. and agricultural land. ∙ This trend contributes to climate WHY ARE MOST CELLS SMALL? DUE change by increasing atmospheric TO SURFACE AREA AND VOLUME carbon dioxide from burning forests RATIO. and impairing the ability of the ∙ Balloon Analogy earth to return carbon to organic ∙ Carnival Analogy matter through photosynthesis. ∙ Forest preservation would result in CELL MEMBRANES STRUCTURE OF the identification of new species of CELL MEMBRANES plants and animals that could be ∙ Functions of Cell Membranes ∙ Cell important human resources. membranes are composed of a double WILD LIFE ALERT layer of proteins and ∙ Extinction has been the fate of most phospholipids which are fluid. plant and animal species. It is a ∙ To protect the machineries inside ∙ natural process that will continue. Helps the cell to maintain its shape ∙ The threat to the welfare of wild Regulates the entering and leaving of plants and animals has increased materials dramatically—mostly as a result of STRUCTURES habitat destruction PHOSPHOLIPIDS - has two ends ∙ An endangered species is in which are polar and nonpolar ends. imminent danger of extinction Comparing it to a sandwich, the throughout its range (where it lives). fillings will be the phospholipid tail ∙ A threatened species is likely to (polar) which attracts each other become endangered in the near and is hydrophobic. While the future. nonpolar ends attracts water CHAPTER 2: CELLS, TISSUES, ORGANS, CHOLESTEROL - embedded along AND ORGAN SYSTEMS OF ANIMALS with phospholipid tails. Helps the WHAT ARE CELLS? membrane to be less permeable to ∙ Cell is the basic unit of life. water soluble substances. Prevents ∙ All living things are made up of phospholipids from sticking to each cells. other to stabilize the figure. There are two types of cells: prokaryotic PROTEINS - has two types of cells and eukaryotic cells. associated proteins which are peripheral (external)whose function is to interact with other proteins ∙ Prokaryotic cells (before nucleus) are and integral (internal) membrane proteins are inserted into the membrane and most pass through gradient across the wall. the membrane. ACTIVE TRANSPORT: ENERGY Other types of proteins and their REQUIRED function as follows: according to Specific carrier proteins in the plasma Bailey (2019) membrane bind with molecules or ions to 1.Structural proteins - gives the cell help them cross the membrane against a shape and support concentration gradient. Cellular energy is 2. Receptor proteins- communicates required. with external environment CARRIER PROTEINS FOR ACTIVE 3. Transport proteins- transport TRANSPORT molecules across membranes by Uniporters- if they transport a single type of diffusion (ex. Globular proteins) molecule or ion 4. Glycoproteins - paired with a Symporters- if they transport two molecules carbohydrate. It also has glycocalyx or ions in the same direction which is the cell coat made up of Antiporters- if they transport two molecules surface carbohydrates, proteins, or ions in the opposite direction and lipids. BULK TRANSPORT MOVEMENT ACROSS MEMBRANES 2 Large molecules cannot be transported Major ways of cell membrane transport through the plasma membrane by the Active transport- requires energy. processes described in the previous Passive transport- use no energy. sections. Endocytosis and exocytosis SIMPLE DIFFUSION together provide bulk transport into and No cell energy is needed. Molecules move out of the cell, respectively. (The term “down” a concentration gradient. Molecules “bulk” is used because many molecules are spread out randomly from areas of higher moved at the same time.) concentration to areas of lower ENDOCYTOSIS concentration until they are distributed The bulk movement of material into a cell evenly in a state of dynamic equilibrium. by the formation of a vesicle. FACILITATED DIFFUSION PINOCYTOSIS Carrier (transport) proteins in a plasma The plasma membrane encloses small membrane temporarily bind with molecules amounts of fluid droplets (in a vesicle) and and help them pass across the membrane. takes them into the cell. Other proteins form channels through PHAGOCYTOSIS which molecules move across the The plasma membrane forms a vesicle membrane. around a solid particle or other cell and TRANSPORT PROTEINS draws it into the phagocytic cell. Molecules can move into and out of cells RECEPTOR-MEDIATED ENDOCYTOSIS through integrated protein channels in the Extracellular molecules bind with specific plasma membrane without using energy. receptor proteins on a plasma membrane, OSMOSIS causing the membrane to invaginate and Water molecules diffuse across selectively draw molecules into the cell. permeable membranes from areas of higher EXOCYTOSIS concentration to areas of lower The bulk movement of material out of a cell. concentration. A vesicle (with particles) fuses with the HYPERTONIC SOLUTION plasma membrane and expels particles or A solution that has a higher solute fluids from the cell across the plasma concentration than another solution. membrane. The reverse of endocytosis. HYPOTONIC SOLUTION CYTOPLASM, ORGANELLES, AND A solution that has a lower solute CELLULAR COMPONENTS concentration than another solution. Organelles effectively compartmentalize a FILTRATION cell’s activities, improving efficiency and Essentially protein-free plasma moves protecting cell contents from harsh across capillary walls due to a pressure chemicals. It also enable cells to secrete various substances, derive energy from Eukaryotic cells contain a variety of nutrients, degrade debris and waste enzyme-bearing, membrane enclosed materials, and reproduce. vesicles called microbodies. The Cytoplasm of a cell has two distinct parts: distribution of enzymes into microbodies is (1) The cytomembrane (or endomembrane) one main way eukaryotic cells organize system consists of well-defined structures, their metabolism. such as the endoplasmic reticulum, Golgi - One specific type of microbody is the apparatus, vacuoles, and vesicles. (2) The peroxisome. Peroxisomes contain fluid cytosol suspends the structures of the enzymes that catalyze the removal of cytomembrane system and contains various electrons and associated hydrogen dissolved molecules. atoms from hydrogen peroxide. Ribosomes: Protein Workbenches Mitochondria: Power Generators Ribosomes are non-membrane-bound Mitochondria are double-membrane bound structures that are the sites for protein organelles that are spherical to elongated in synthesis. They contain almost equal shape. A small space separates the outer amounts of protein and a special kind of membrane from the inner membrane. The ribonucleic acid called ribosomal RNA inner membrane folds and doubles in on (rRNA). itself to form incomplete partitions called – ribosomal RNA (rRNA) is the molecule in cristae. And cristae increase the surface the cells that forms part of the protein area available for the chemical reactions synthesizing organelle known as a ribosome that trap usable energy for the cell. and that is exported to the cytoplasm to The space between the cristae is the matrix. help translate the information in messenger The matrix contains ribosomes, circular RNA (mRNA) into protein. DNA, and other material. Endoplasmic Reticulum: Production and Because they convert energy to a usable Transport form, mitochondria are frequently called the The endoplasmic reticulum (ER) is a “power generators” of the cell. Mitochondria complex, membrane-bound labyrinth of usually multiply when a cell needs flattened sheets, sacs, and tubules that additional high-energy molecules. branches and spreads throughout the Cytoskeleton: Microtubules, cytoplasm. The ER is continuous from the Intermediate Filaments, nuclear envelope to the plasma membrane and Microfilaments and is a series of channels that helps In most cells, the microtubules, various materials to circulate throughout intermediate filaments, and microfilaments the cytoplasm. form the flexible cellular framework called Golgi Apparatus: Packaging, Sorting, and the cytoskeleton (“cell skeleton”). Three Export Major Classes of Protein Fibers Making The Golgi apparatus or complex (named for Up the Cytoskeleton of Eukaryotic Cell Camillo Golgi, who discovered it in 1898) is ● Microtubules - are hollow, slender, a collection of membranes associated cylindrical structures in animal physically and functionally with the ER in cells. Each microtubule is made of the cytoplasm. It is composed of flattened spiraling subunits of globular stacks of membrane-bound cisternae proteins called tubulin subunits. - The Golgi apparatus sorts packages, ● Intermediate filaments - are a and secretes proteins and lipids. chemically heterogeneous group of Lysosomes: Digestion and Degradation protein fibers, the specific proteins of Lysosomes (Gr. lyso, dissolving 1 soma, which can vary with cell type. body) are membrane-bound spherical ● Microfilaments (actin filaments) - organelles that contain enzymes called acid are solid strings of protein (actin) hydrolases, which are capable of digesting molecules. organic molecules Cilia and Flagella: Movement Microbodies: A Diverse Category of Cilia (sing., cilium; L. eyelashes) and Organelles flagella (sing., flagellum; L. small whips) are elongated appendages on the surface of cytoskeleton. some cells by which the cells, including Vacuoles: Cell Maintenance many unicellular organisms, propel Vacuoles (L. vaccus, empty space) are themselves. In stationary cells, cilia or membranous sacs that are part of the flagella move material over the cell’s cytomembrane system. surface. Vaults: Mysterious Symmetrical Shells ● Axoneme or Axial filament Vaults are cytoplasmic ribonucleoproteins - Consists of nine pairs of shaped like octagonal barrels. Their name microtubules arranged in a is derived from their multiple arches that circle around two central look like vaulted cathedral ceilings. tubules. This is called a 9 1 Exosome: Signaling Structures 2 pattern of microtubules. Exosomes - Each microtubule pair (a - contain cell-specific payloads of doublet) also has pairs of proteins, lipids, and genetic material dynein (protein) arms that are transported to other cells projecting toward a and tissues, where they alter many neighboring doublet and cellular functions. spokes extending toward the THE NUCLEUS: INFORMATION CENTER central pair of microtubules. Nucleus ● Basal body - When viewing a cell image, the - Made up of microtubules nucleus is one of the most visible and structurally identical to elements. the centriole. - a nucleus is a membrane-enclosed - controls the growth of organelle within a cell that holds the microtubules in cilia or chromosomes. flagella. The microtubules in - An array of holes, or pores, in the the basal body form a 9 1 0 nuclear membrane allow for the pattern: nine sets of three selective passage of particular with none in the middle. molecules (such as proteins and Internal Structure of Cilia and Flagella - nucleic acids) into and out of the In cross section, the arms extend from each nucleus. microtubule doublet toward a neighboring - It houses all of the eukaryotic cell's doublet, and spokes extend toward the genome and acts as a center for central paired microtubules. The dynein controlling cellular activities arms push against the adjacent Nuclear Envelope microtubule doublet to bring about - The nuclear envelope separates the movement. nucleus' contents from the Centrioles and Microtubule-Organizing cytoplasm and serves as the Centers microtubule-organizing centers nucleus' structural framework. - The specialized nonmembranous - The nuclear membranes, acting as regions of cytoplasm near the barriers that prevent the free nucleus. passage of molecules between the - These centers of dense material give nucleus and the cytoplasm, rise to a large number of maintain the nucleus as a distinct microtubules with different biochemical compartment. functions in the cytoskeleton. ● The nuclear membrane consist of: - Centrioles Outer nuclear membrane - is composed of nine triplet - Inner nuclear membrane microtubules that radiate from the - Perinuclear membrane center like the spokes of a wheel. - are - Nuclear Pores duplicated before cell division, are - Nuclear Lamina involved with chromosome - Nuclear pore complexes (NPCs) movement, and help organize the serve as the gateway of the cell nucleus. These macromolecular organelles that are required to assemblies form selective aqueous complete all of the basic processes translocation channels permitting necessary for survival and the free diffusion of small molecules, regeneration. Animals with cell-level as well as receptor-mediated organization have a loose transport of large cargoes arrangement of cells. Chromosomes: Genetic Containers (kahit What is Structural Organization in a elements na lang na may chromosomes Cell? sa ppt) - The cell contains all the organelles - DNA, or deoxyribonucleic acid, is which are required for a cell to located in the nucleus of almost survive. All the organelles are placed every cell in the human body and form a structural organization - Histones serve to bundle in the cell. chromosomal DNA inside tiny Cell Organelles and their Functions 1. nuclei. These are positively charged Nucleus: Storage of genetic material i.e proteins that bind to negatively DNA. charged DNA to form complexes 2. Endoplasmic Reticulum: They help known as nucleosomes. in the synthesis of protein and Nucleolus: Pre Assembly Point for lipids. Ribosomes 3. Golgi Apparatus: It mainly helps in - The formation of ribosomes involves tagging and transportation. the assembly of the ribosomal 4. Ribosome: They synthesize the precursor RNA with both ribosomal protein with the help of mRNA. proteins and 5S rRNA 5. Mitochondria: Mitochondria - RNA polymerase II transcribes the generate ATP, and because of that genes that encode ribosomal mitochondria are known as the proteins outside of the nucleolus, Powerhouse of cells. producing mRNAs that are 6. Lysosome: Helps in digestion within translated on cytoplasmic the cell via decomposing. ribosomes. Ribosomal proteins are 7. Cytoplasm: Cytoplasm is a thick then transported from the cytoplasm substance in which all the cell to the nucleolus and combined with organelles are embedded. rRNAs to produce pre ribosomal 8. Plasma Membrane: It is the outer particles. Although 5S rRNA genes layer of the cell, and helps in are also transcribed outside of the maintaining the equilibrium of the nucleolus, this time by RNA cell polymerase III, 5S rRNAs are Tissue Level of Organizations assembled into pre ribosomal - Tissue is the most critical layer in particles within the nucleolus. the protection of the organ, and Levels of Organization in Animals - A tissues let only the necessary one cell is the primary unit of life and the into the organ while keeping it safe. - fundamental unit of the animal. - A sort of Tissues are collections of tissue is formed when a large number of comparable cells from the same cells (of the same type) congregate in one origin that collaborate to carry out a location. defined function. - Tissues join together to produce Human basic tissues are classed as follows: organs Epithelial Tissue - it is found all over the - Organ systems emerge later on. - To body. They cover all body surfaces, line function properly, many organs body cavities and hollow organs, and make collaborate. up the majority of gland tissue. They serve Cellular Level of Organizations a number of purposes, including defense, - The cell is the most basic unit of life. - secretion, absorption, excretion, filtration, The cell contains all of the cell diffusion, and sensory reception. Types of Epithelial Tissue across the body, where both support 1. Simple squamous - Transport by and suppleness are required. Blood diffusion and where minimal vessels, nerves, and muscles are all protection is required wrapped in loose connective tissue. 2. Simple Cuboidal Epithelium - They, along with adipose tissues, create Absorption and secretion the layer of fat beneath the skin, linking 3. Simple Columnar Epithelium - muscles and other Protection, absorption, mucus structures to the skin. secretion and movement in a ● Adipose Tissue specific direction ● Areolar Tissue 4. Stratified Squamous Epithelium - ● Reticular Connective Tissue Protection 2. Dense Connective Tissue - 5. Stratified Cuboidal - Protection of Fibroblast cells and fibers are ducts of various glands densely packed in dense connective 6. Stratified Columnar - Protection tissue. Their primary role is to and secretion transfer and support mechanical 7. Pseudostratified Columnar - forces. They are less pliable than Protection, secretion and movement loose connective of mucous ● Dense regular tissue 8. Transitional epithelia or ● Tendons and Ligaments urothelium - Stretch readily to ● Dense irregular tissue accommodate the different volume 3. Specialized Connective Tissue - of liquids Aside from these, there are 9. Keratinised - Protection against supportive connective tissues, such abrasion as cartilage and bone, that aid in Connective Tissue - As the name implies, maintaining proper posture and connective tissues support and link various supporting internal organs. Blood tissues and organs of the body. They are and lymph are fluid connective abundantly spread throughout the body. tissues that circulate throughout They are derived from the mesoderm. the body and aid in organ - Connective tissues contain three interaction and communication. types of fibers: collagen, elastic ● Cartilage - Cartilage is largely and reticular present throughout embryonic ● Collagen fibers are the most development and serves as a common and are formed of the supporting skeleton. fibrous protein collagen. Collagen ● Bones - Bone is the hardest fibers are strong and flexible, with connective tissue and aids in the tensile strengths comparable to maintenance of the body's structure steel. and posture, as well as the ● Elastic fibers can be stretched like protection of interior organs. a rubber band to form a network. ● Blood - Blood is made up of various They are composed of the protein cells present in the plasma. The elastin. When the force is blood contains red blood cells withdrawn, they return to their (RBCs), white blood cells (WBCs) normal shape and size. and platelets. ● Reticulate fibers consist of collagen ● Lymph - it drains into the and glycoproteins. They are thin and bloodstream and carries absorbed form a delicate network. They join fat that cannot reach the connective tissues to neighboring bloodstream directly tissues. Nervous Tissues Types of Connective Tissues Consists of diverse cells such as red blood 1. Loose Connective Tissue - Loose cells, white blood cells, and neurons. connective tissues can be found all Specific cells are for protection, support and nourishment such as: proposed that eukaryotes formed 1. Blood - consists of RBC, WBC, and when large, non nucleated cells platelets that are suspended in engulfed smaller and simpler cells. ∙ plasma in blood vessels. Main Endosymbiont is an organism that can function is to carry oxygen, live only inside another nutrients, hormones, minerals, etc. 2. organism, forming a relationship Nervous Tissue- where neurons can be that benefits both partners. found more specifically in the ∙ Symbiosis is an intimate brain and spinal cord whose association between two organisms function is to transfer electrical of different species. The merging of signals from senses. these different species to produce 3. Skeletal muscle tissue - made up evolutionarily new forms is called of striated muscle cells that have symbiogenesis striped appearance, long and ∙ More recently, DNA evidence cylindrical. Can be found in bones indicates that both Archaea and that are connected to skeletal Eubacteria contributed to the origin muscle. Responsible for movements of eukaryotic cells. About 2.5 billion and locomotions. years ago, bacteria and 4. Smooth muscle tissue - non striated cyanobacteria occurred together in muscle and cells are closed to each water environments. Over millions other to form a sheet. of years, the cyanobacteria pumped Spindle shaped cells and each oxygen into the primitive contains a nucleus. atmosphere as a by-product of Muscle Tissue photosynthesis. As a result, those responsible for movement and there are 3 cellular organisms that could classifications of muscle tissue which are tolerate free oxygen began to cardiac muscle, skeletal muscle and flourish. smooth muscle. The contraction of muscle ● One way for a large cell, such as an tissue allows different body parts and archaeon, to survive in an oxygen rich organs to move. environment would be to engulf an Cardiac muscle- keeps the heart from aerobic (oxygen-utilizing) pumping with involuntary movement bacterium in an inward-budding Skeletal muscle- for joints, movement of vesicle of its plasma membrane. bones, chewing, swallowing, urination and ● The membrane of the enveloped even maintaining body temperature Smooth vesicle became the outer membrane Muscle - can be found throughout the body of the mitochondrion. except heart and bones. ● Archaean cells that picked up ORGANS & ORGAN SYSTEMS cyanobacteria or a photosynthetic Organs bacterium obtained the forerunners (an independent part of the body) are the of chloroplasts and became the functional units of an animal’s body that ancestors of the green plants. are made up of more than one type of ● The exact mechanism for the tissue. Examples include the heart, lungs, evolution of the eukaryotic cell will liver, spleen, and kidneys. never be known with certainty. Organ system ● Some probably evolved into is an association of organs that together multicellular organisms in which performs an overall function various cells become specialized into EVOLUTIONARY INSIGHTS tissues, which in turn led to the Endosymbiont theory potential for many different ∙ was first proposed by Lynn functions. Margulis (1938–2011), a biologist INTEGUMENTARY SYSTEM who worked at the University of - It is the largest organ (skin) of the Massachusetts at Amherst. She body that forms a physical barrier between the external environment and the internal the neural impulse environment that it serves to protect and THREE TYPES OF VERTEBRATE maintain. NEURONS THREE LAYERS OF THE SKIN - Sensory neurons: act as receptors of EPIDERMIS stimuli or are activated by receptors. - - It is the most superficial layer of Interneurons: receive signals from the the skin and provides the first barrier of sensory neurons and transmit them to protection from the invasion of substances motor neurons. Interneurons are located into the body. entirely within the central nervous system. - DERMIS Motor neurons: send the - It is the connective tissue processed information via a signal to the meshwork of collagenous, reticular, and body’s effectors (e.g., muscles), causing elastic fibers beneath the epidermis. them to contract, or to glands, causing HYPODERMIS them to secrete. - It consists of loose connective TWO MAIN DIVISIONS OF NERVOUS tissue, adipose tissue, and nerve endings, SYSTEM which separate the skin from deeper CENTRAL NERVOUS SYSTEM tissues. - The central nervous system is THE SYSTEM HAS FOUR MAJOR composed of the brain and spinal cord and FUNCTIONS: is the site of information processing. 1. Regulation of body temperature PERIPHERAL NERVOUS SYSTEM 2. Excretion of waste materials - The peripheral nervous system is 3. Vitamin D3 formation by the action composed of all the nerves of the body of ultraviolet radiation from sunlight on outside the brain and spinal cord. These a cholesterol derivative in the nerves are commonly divided into two skin. groups: sensory (afferent) nerves, which 4. Reception of environmental stimuli, transmit information to the central nervous such as pain, temperature, and system; and motor (efferent) nerves, which pressure. carry commands away from the central NERVOUS SYSTEM nervous system. - an extensive network of specialized THREE AREAS OF THE BRAIN cells that carry information to and from all HIND BRAIN parts of the body - The hindbrain is continuous with NEURONS the spinal cord and includes the medulla - Neurons are specialized to produce signals oblongata, cerebellum, and pons. that can be communicated over short to MEDULLA OBLONGATA relatively long distances, from one part of - The medulla oblongata is the an animal’s body to another. enlargement where the spinal cord enters Neurons have two important properties: (1) the brain. It contains reflex centers for excitability, the ability to respond to breathing, swallowing, cardiovascular stimuli, and (2) conductivity, the ability function, and gastric secretion. to conduct a signal. CEREBELLUM PARTS OF NEURONS - The cerebellum is an outgrowth of the - Soma: the cell body of the neuron, medulla oblongata. It coordinates motor responsible for maintaining the life of the activity associated with limb movement, cell maintaining posture, and spatial - Dendrites: branch-like structures orientation. that receive messages from other neurons - PONS Axon: a long, tube-like structure that - The pons is a bridge of transverse carries the neural message to other cells nerve tracts from the cerebrum of the - Myelin: fatty substances produced forebrain to both sides of the cerebellum. It by certain glial cells that coat the axons of also contains tracts that connect the neurons to insulate, protect, and speed up forebrain and spinal cord in all vertebrates and functions in the regulation of muscle contractions breathing. 2. Certain hormones are species-specific MIDBRAIN with respect to their function; conversely, - The midbrain was originally a some hormones produced in one species center for coordinating reflex responses to may be completely functional in another visual input. As the brain evolved, it took species. on added functions relating to tactile Pituitary Gland (Hypophisis) (touch) and auditory (hearing) input, but it ● located directly below the did not change in size. The roof of the mid hypothalamus brain, called the optic tectum, is a ● has two lobes thickened region of gray matter that ∙ Adenohypophisis (Anterior integrates visual and auditory signals. lobe) FOREBRAIN ∙ Neurohypophisis (Posterior - The vertebrate forebrain has lobe) changed a great deal during vertebrate ● the pituitary in many vertebrates evolution. The forebrain has two main (except humans, birds and parts: the diencephalon and telencephalon. cetaceans) also has a functional The diencephalon lies just in front of the intermediate lobes (pars intermedia) midbrain and contains the pineal gland, that is responsible for the coloration pituitary gland, hypothalamus, and of their body surface thalamus. Neurohypophysis - The thalamus relays all sensory ● does not manufacture any information to higher brain centers. - The hormones hypothalamus lies below the thalamus ● neurosecretory cells of the and regulates many functions, such as hypothalamus synthesize and body temperature, sexual drive, secretes 2 hormones carbohydrate metabolism, hunger, and Adenohypophysis thirst. ● the true endocrine portion of the - The pineal gland controls some pituitary body rhythms. The pituitary is a major ● it synthesizes 6 polypeptide endocrine gland hormones Endocrine System Thyroid Gland A hormone (Gr. hormaein, to set in motion ● Is located in the neck, anterior to or to spur on) is a specialized chemical the trachea messenger that an endocrine gland or ● secretes thyroxine and tissue produces and secretes. Hormones triiodothyronine - which are circulate through body fluids and affect the responsible for the overall growth, metabolic activity of a target cell or tissue development and metabolic rate of in a specific way. the organism 3 Major Regions that secrete neuropeptides ● also secretes calcitonin - which 1. Pineal Gland (Epithalamus) - secrete controls the extracellular levels of hormones for pigmentation and calcium ions hormones that inhibit reproductive Parathyroid Gland development (both are stimulated by ● tiny, pea-sized glands embedded in light) the thyroid lobes 2. Preoptic Nuclei (Hypothalamus) - ● secrete parathormone - which secretes neuropeptides that control regulates the concentrations of different functions such as growth, calcium and phosphate ions in the sleep, locomotion blood 3. Urophysis (oura - tail + physis - Thymus Gland growth) - produces neuropeptides ● located near the heart that help control water and ion ● secretes peptide hormones: balance, balance and smooth ∙ Thymopoietin ∙ Alpha1 into the bloodstream after a ∙ Beta4 meal and inhibits ∙ Thymosin somatostatin secretion, all of which are essential for normal gallbladder contraction, and development of the immune system the secretion of pancreatic Adrenal Glands digestive enzymes. ● located on top of the kidneys Gonads ● each gland consists of the: ● ovaries and testes secrete hormones ∙ medulla (inner portion) that help regulate reproductive ∙ cortex (outer portion that functions surrounds the medulla) Testes Adrenal Cortex ● secretes testosterone secretes 3 classes of steroid hormones: ● ∙ necessary for the growth and Glucocorticoids (cortisol)- regulate maintenance of the male sex metabolism and concentration of organs blood sugar ∙ It promotes the development ● Mineralocorticoids (aldosterone) - and maintenance of sexual maintains concentration of solutes behavior, and in humans, it entering the blood stream and plays stimulates the growth of major role in maintaining facial and pubic hair, as well homeostasis of extracellular solutes as the enlargement of the ● Sex Hormones (Androgens and larynx, which deepens the Estrogens) voice. Adrenal Medulla Ovary ● under neutral control ● secrets 4 major classes of ovarian ● secretes epinephrine (adrenaline) hormones that help regulate female and norepinephrine (noradrenaline) ∙ reproductive functions. helps control heart rate and Skeletal system carbohydrate metabolism As organisms evolved from the ancestral ∙ has a major role in the fight - protists to the multicellular animals, body flight reaction size increased dramatically and systems Pancreas involved in movement and support evolved ● functions as an exocrine gland to as well. secrete digestive enzymes and an 3 Types of Skeleton endocrine gland 1. Rigid Endoskeleton ● The endocrine portion of the 2. Rigid Exoskeleton pancreas makes up only about 1% 3. Hydrostatic Skeleton of the gland. This portion These skeletal systems also function in synthesizes, stores, and secretes animal movement that requires muscles hormones from 200,000 - 2,000,00 working in opposition (antagonism) to each clusters of cells called pancreatic other. islets. Exoskeleton ● these islets contains 4 special ● exo - outside + skeleton groups of cells called: ● This type of skeleton also have ∙ Alpha - produces glucagon locomotor functions because they ∙ Beta - produces insulin provide sites for muscle attachment ∙ Delta - somatostin and counterforces for muscle (hypothalamic growth movements. hormone inhibiting factor ● It also supports and protects the body. that also inhibits glucagon Exoskeletons, however, limit an animal's and insulin secretion) growth, hence, these animals occasionally ∙ F cells - secrete a pancreatic polypeptide that is released shed, as arthropods do when they molt. Endoskeleton A few functional differences among ● endo - within + skeleton invertebrate ● This type of skeleton is enclosed by CIRCULATORY SYSTEM other body tissues and is used for ∙ A circulatory or cardiovascular protection and support. system is a specialized system in Cartilage which a muscular, pumping heart ● It is a specialized type of connective moves the fluid medium called tissue that provides a site for either hemolymph or blood in a muscular attachment, aids in specific direction determined by the movement at joints, provides presence of unidirectional blood support, and transmits the force of vessels. muscular contraction from one part LYMPHATIC SYSTEM of the body to another during - The lymphatic system, or lymphoid movement. system, is an organ system in vertebrates ● It consists of cells (chondrocytes), that is part of the immune system, and fibers and a cellular matrix. complementary to the circulatory system. It Bone consists of a large network of lymphatic ● Or osseous tissue is a connective vessels, lymph nodes, lymphoid organs, tissue that provides a point of lymphoid tissues and lymph. attachment for muscles and LYMPHATICS: are thin-walled vessels with transmits the force of muscular valves that ensure the one-way flow of contraction form one part of the lymph. body to another during movement. LYMPH- is the extracellular fluid that ● Bones of the skeleton support the accumulates in the lymph vessels. internal organs of many animals, RESPIRATORY SYSTEM store reserve calcium and phosphate - the system in living organisms that takes and manufacture red blood up oxygen and discharges carbon dioxide in cells.skeleton order to satisfy energy requirements. MUSCULAR SYSTEM Muscular tissue is the driving force, the Respiratory Surfaces power behind movement in most Protists and animals have five main types of invertebrates and vertebrates. respiratory systems (surfaces): The ability to move—one of the most (1) simple diffusion across plasma defining properties of animals—often has membranes, life-and-death significance. (2) tracheae, Animals may have one or more of the (3) cutaneous (integument or body following types of muscle tissue: Smooth, surface),exchange, Cardiac, and Skeletal muscles. The (4) gills, and contractile cells of these tissues are called (5) lungs. muscle fibers. Bimodal breathing is the ability of an Smooth muscle is also called involuntary organism to exchange respiratory gases muscle because higher brain centers do not simultaneously with both air and water. control its contractions. Smooth muscle Cutaneous Exchange fibers have a single nucleus, are spindle Some vertebrates that have lungs or gills, shaped, and are arranged in a parallel such as some aquatic turtles, salamanders pattern to form sheets with lungs, snakes, fishes, and Cardiac muscle fibers are involuntary, mammals,use cutaneous respiration or have a single nucleus, are striated (have integumentary exchange to supplement gas dark and light bands), and are branched exchange. However, cutaneous exchange is Skeletal muscle, also a striated muscle, is most highly developed in frogs, toads, a voluntary muscle because the nervous lungless salamanders, and newts. Gills system consciously controls its Gills are respiratory organs that have either contractions. a thin, moist, vascularized layer of epidermis to permit gas exchange across whether an animal feeds on plants thin gill membranes, or a very thin layer of (herbivore) or animals (carnivore), or both epidermis over highly vascularized dermis. (omnivore), and on how it obtains its food. Larval forms of a few fishes and amphibians SALIVARY GLANDS (“Specialized have external gills projecting from their Exocrine Glands”) bodies. Adult fishes have internal gills. - any of the organs that secrete saliva, a Lungs substance that moistens and softens food, A lung is an internal sac-shaped respiratory into the oral cavity of vertebrates. Salivary organ. The typical lung of a terrestrial glands may be predominantly serous, vertebrate comprises one or more internal mucous, or mixed in secretion. Mucus is a blind pouches into which air is either drawn thick, clear, and somewhat slimy or forced. substance. Lung Ventilation ESOPHAGI (“Moving Food to the Ventilation is based on several physiological Stomach”) ( LONG OR SHORT ) principles that - Storing food in the crop ensures an apply to all air-breathing animals with almost continuous supply of food to the lungs: stomach and intestine for digestion. This Gas-Exchange Portion structure allows these to reduce the Small tubes called alveolar ducts connect frequency of feeding and still maintain a the respiratory bronchioles to grapelike high metabolic rate. outpouchings called alveoli. STOMACHS (“Holding Stations”) Gas Transport - The stomach is an ancestral As animals became larger and acquired vertebrate structure that evolved as higher metabolic rates, simple diffusion vertebrates began to feed on larger became increasingly inadequate as a means organisms that were caught at less frequent of delivering oxygen to the tissues. intervals and required storage. Respiratory pigments help the various GIZZARDS (“Mechanical Grinders”) transport systems satisfy this increased - The gizzard has a number of oxygen demand. In addition to oxygen important functions, such as aiding transport, respiratory pigments may also digestion by particle size reduction, function in short-term oxygen storage. chemical degradation of nutrients and DIVERSITY IN DIGESTIVE STRUCTURES: regulation of feed flow, and responds VERTEBRATES rapidly to changes in the coarseness of the - The complete vertebrate digestive tract diet. (gut tube) is highly specialized in both RUMENS (“Eat Now, Digest Later”) - This structure and function for the digestion of a method of digestion has evolved in animals wide variety of foods. The basic structures of that need to eat large amounts of food the gut tube include the oral cavity (buccal relatively quickly, but can chew the food at cavity or mouth), pharynx, esophagus, a more comfortable or safer location. CECA stomach, small intestine, large intestine, (“Blind Pouches”) rectum, and anus/cloaca. In addition, three - cecum, also spelled caecum, pouch or important glandular systems are associated large tubelike structure in the lower with the digestive tract: (1) the salivary abdominal cavity that receives undigested glands; (2) the liver, gallbladder, and bile food material from the small intestine and duct; and (3) the pancreas and pancreatic is considered the first region of the large duct intestine. TONGUES (“Specialized Mouthparts”) ( LIVERS AND GALLBLADDERS TYPES OF TONGUE ) (“Accessory Organs”) - A tongue or tonguelike structure - In those vertebrates with a develops in the floor of the oral cavity in gallbladder, it is closely associated with the many vertebrates. liver. The liver manufactures bile, which the TEETH (“Reflect the Nature of the Diet”) gallbladder then stores. Bile is a fluid - Teeth are specialized, depending on containing bile salts and bile pigments. Bile salts play an important role in the digestion common bile duct that enters the of fats, although they are not digestive duodenum. Pancreatic enzymes complete enzymes. They emulsify dietary fat, the digestion of carbohydrates and proteins breaking it into small globules and initiate the digestion of lipids. Role of (emulsification) on the surface of which the the Liver and Gallbladder in Digestion fat-digesting enzyme lipase can function. - The liver, the largest internal organ in PANCREATA (“Specialized Exocrine and the mammalian body, the liver, millions of Endocrine Glands”) specialized cells called hepatocytes take up - As an endocrine gland, it functions nutrients absorbed from the intestines and mostly to regulate blood sugar levels, release them into the bloodstream. secreting the hormones insulin, glucagon, Hepatocytes also manufacture the blood somatostatin and pancreatic polypeptide. proteins prothrombin and albumin. As a part of the digestive system, it Reproductive System functions as an exocrine gland secreting A reproductive system is a biological system pancreatic juice into the duodenum that allows an organism to produce through the pancreatic duct. offspring. It consists of organs, glands, and INTESTINES (“Breakdown and structures that are involved in the process Absorption”) of sexual reproduction. The reproductive - The configuration and divisions of the system is essential for the survival and small and large intestines vary greatly evolution of a species, as it allows for the among vertebrates. Intestines are closely creation of genetic diversity and adaptation related to the animal’s type of food, body to changing environments. The size, and levels of activity. reproductive system is also a source of ORAL CAVITY pleasure, intimacy, and identity for many - A pair of lips protects the oral cavity organisms, especially humans. (mouth). The lips are highly vascularized, Asexual Reproduction in Invertebrates Is skeletal muscle tissue with an abundance reproduction without the union of gametes of sensory nerve endings. Lips help retain or sex cells. Asexual reproduction is food as it is being chewed and play a role in common among the protozoa, as well as phonation (the modification of sound). among invertebrates, such as sponges, - The oral cavity contains the tongue jellyfishes, flatworms, and many annelids. and teeth Asexual reproduction is rare among the PHARYNX AND ESOPHAGUS other invertebrates. - The epiglottis temporarily seals off the FISSION opening (glottis) to the trachea so that Fission is the division of one cell, body, or swallowed food does not enter the trachea. body part into two Initiation of the swallowing reflex can be In this process, the cell pinches in two by voluntary, but most of the time it is an inward furrowing of the plasma involuntary. When swallowing begins, membrane. sequential, involuntary contractions of Protists and some animals (cnidarians and smooth muscles in the walls of the annelids) may reproduce by fission. esophagus (Gr. oisophagos, to carry food) BINARY FISSION propel the bolus or liquid to the stomach. Binary fission occurs when the division is Neither the pharynx nor the esophagus equal; each offspring contains contributes to digestion. approximately equal amounts of protoplasm Role of the Pancreas in Digestion - The and associated structures. Binary fission is pancreas (Gr. pan, all 1 kreas, flesh) is an common in protozoa; for some, it is their organ that lies just ventral to the stomach only means of reproduction. and has both endocrine and exocrine It can be longitudinal, transverse, functions. Exocrine cells in the pancreas asymmetrical, oblique secrete digestive enzymes into the MULTIPLE FISSION pancreatic duct, which merges with the Some flatworms and annelids reproduce by hepatic duct from the liver to form a forming numerous constrictions along the length of the body; a chain of daughter nourished by an organ called the individuals results. placenta. BUDDING The bud cells proliferate and form a cylindrical structure, which develops into a new animal, usually breaking away from the parent. If the buds remain attached to the parent, they form a colony. FRAGMENTATION Fragmentation is a type of asexual reproduction whereby a body part is lost and then regenerates into a new organism. PARTHENOGENESIS Parthenogenesis is a spontaneous activation of a mature egg, followed by normal egg divisions and subsequent embryonic development. In fact, mature eggs of species that do not undergo parthenogenesis can sometimes be activated to develop without fertilization by pricking them with a needle, by exposing them to high concentrations of calcium, or by altering their temperature. parthenogenetic eggs are not fertilized, they do not receive male chromosomes. The offspring would thus be expected to have only a haploid set of chromosomes. In some animals, however, meiotic division is suppressed, so the diploid number is conserved. In other animals, meiosis occurs, but an unusual mitotic division restores the embryo to the diploid state. EXTERNAL FERTILIZATION Many invertebrates (e.g., sponges and corals) simply release their gametes into the water in which they live (broadcast spawning), allowing external fertilization to occur. INTERNAL FERTILIZATION Other invertebrates (from flatworms to insects) utilize internal fertilization to transfer sperm from male to female and have structures that facilitate such transfer The monotremes are a group of highly specialized egg-laying predatory mammals, containing the platypus and echidnas. Marsupial A marsupial is a mammal that raises its newborn offspring inside an external pouch at the front or underside of their bodies. Placental A placental is a mammal that completes embryo development inside the mother,