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Reduction of 3-phosphoglycerate to
form glyceraldehyde-3-phosphate
which can be used in formation of
carbon compounds that are
translocated.
Saturation level.
sometimes called
photosynthetic
capacity.
Photosynthetic efficiency:
Increase in photosynthesis
per increase in irradiance
0
0
Compensation point
The irradiance at which CO uptake is zero
2
Measured light response curve of Abies amabilis first year foliage.
Shade foliage with low maximum value and low compensation point.
Observed assimilation rates
(µmolCO2/m2s) of Tsuga
heterophylla and Abies amabilis in
response to periods of 10 minutes
high light (1500µmol/m2s PPFD),
with 5 minutes intervals of
darkness (shaded parts in the
diagram) in between. Values
measured using 200 mol/s flow
rate.
Species differences in leaf photosynthesis
A has the highest
photosynthetic rate at
light saturation
Units: μmol/m2/s
micro mols of CO2 per square
meter foliage per second
Abies grandis
Western
hemlock 9.0 4.9 3.2 4.8
Tsuga
The problem of photorespiration
and the evolution of photosynthesis
When the enzyme Rubisco uses oxygen to breakdown
carbohydrate to CO2 rather than using CO2 to synthesize
carbohydrate
O2
P P P P
C-C-C-C-C C-C-C + C-C
RuBisCO
3-phospho 2-phospho
Ribulose 1, 5-biphosphate
glycerate glycerate
Enzyme
CO2
The inhibition of
photosynthesis by O2 was first
73 ppm CO2
noticed by the German plant
physiologist, Otto Warburg, in
1920, and called the "Warburg
effect".
Xylem
Phloem
Carbon
skeleton
compounds
return to
parenchyma
C4 acids synthesized in the
parenchyma move to the bundle
sheath
Crassulacean Acid Metabolism (CAM)
First discovered in succulents of the
Crassulacea: e.g.,sedums
Optimum 30-40oC
15-25oC 35oC
Temperature