PALEONTOLOGY (GEOL

1108)
THEME ASSIGNED: Invertebrates
TITLE:Origin, phylogeny, faunal composition, and
stratigraphical significance
of the Albian engonoceratidae (pulchelliaceae,
ammonitina) of Peru
BY: Emmanuel Roberta, Luc Georges Bulotb
Presented by:
Morocho Marlon, Ortega Nathalia
SCHOOL OF EARTH SCIENCES, ENERGY AND ENVIRONMENT
YACHAY TECH UNIVERSITY
Dr. Jorge Toro Álava, PhD.
Lecturer of Geol 1100
Urcuquí, August 16, 2022.

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Content
• Background - General presentation
• Geographic location of the study
• General geology
• The Geo-Problem – Objective to be solved
• Methodology
• Findings – Results
• Conclusions
• References

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 Background
• Recent study of the Albian ammonite fauna from
Peru provides a major revision of the systematic
position and taxonomy of the family
Engonoceratidae, which we place in the superfamily
Pulchelliaceae.
• Peruvian specimens show a monophyletic evolution
marked by four morphological and stratigraphical
stages, from unornamented Platiknemiceras to
evolute Glottoceras and Engonoceras.
• The highest diversity (15 species) in the upper part
of the Lower Albian.

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 Geographic location of the study
• During the Albian,this
basin recorded a major
marine transgression,
characterized by three
transgressive pulses:

• LowerAlbian:Pariahuanca
• Anoxic sediments of the
Pariatambo
• Shelf carbonates of the
Pulluicana
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Geographic location of the study

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 General Geology
• The Peruvian part of the Andean basin is a back-
arc zone protected by a magmatic arc to the west
and characterized by shallow-water depths.

• In the North Andean Basin: 550 specimens

representative of the genera Parengonoceras,
Glottoceras, Engonoceras, and Platiknemiceras.
Understanding of the faunal assemblages,
biochronostratigraphic significance, ontogeny, and
phylogeny of the Peruvian Engonoceratidae.

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GeoProblem- Objective

•Family engonoceratidae (Hoplitaceae or

pulchelliaceae?)
•Peruvian and Andean engonoceratids
• Origin
• Peruvian ancestor Parengonoceras haasi
Benavides Ca ́ceres
• Phylogeny
• Major characters and faunal composition
• Genus Glottoceras Hyatt
• Other genera
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Methodology
• Recollecting the major amount of samples
in the study zone.

• Classification of different samples

depending of:
• Family engonoceratidae
• Peruvian and Andean engonoceratids

• Analizing the samples by a correct

interpreation of the paleoenviroment
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Results
Family engonoceratidae
Hoplitaceae or pulchelliaceae?
• Luppov and Drushchits (1958), integration of
Engonoceratidae (including subfamilies
Neolobitinae, Engonoceratinae, and
Knemiceratinae) and the families Sphenodiscidae
and Placenticeratidae.

• Similarity of the discoidal sections of the primitive

Platiknemiceras and some Engonoceras to the
Barremian–Aptian Subpulchellia and close forms.
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Results
Family engonoceratidae
Taxonomic content
• Engonoceratidae included Knemiceras Bo ̈hm,
Engonoceras Neumayr and Uhlig, Neolobites
Fischer, and Protengonoceras Hyatt.
• Enonoceratidae includes Knemiceras Bo ̈hm,
Parengonoceras Spath, Glottoceras Hyatt,
Hypengonoceras Spath, Platiknemiceras
Bataller, Engonoceras Neumayr and Uhlig,
Protengonoceras Hyatt, Metengonoceras
Hyatt, and Neolobites Fischer.
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Results
Family engonoceratidae
Origin and biostratigraphical significance

• The oldest representatives of the family are

Parengonoceras mahmoudii, Parengonoceras
algerianum, and Parengonoceras hachourii of the
‘Monts du Mellegue’

• From east to west, they have been recorded from

present Japan and India to new world basins
such as the Caribbean and Andean
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Results
Family engonoceratidae
Origin and biostratigraphical significance

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Results
Peruvian and Andean engonoceratids
Origin
• It belongs to the first Albian ammonite association
of Peru, represents the umbilicostatus biohorizon,
considered as equivalent pro parte to the
European

• Beyond its original area (Baños del Inca),

biostratigraphy identifies this specimen as the
oldest engonoceratid in the Andean basin

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Results
Peruvian and Andean engonoceratids
Origin

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Results
Peruvian and Andean engonoceratids
Origin

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Results
Peruvian and Andean engonoceratids
Origin

Puente Zonanga

• The first clear representative of the Peruvian

Engonoceratidae is the primitive Parengonoceras
haasi Benavides (Puente Zonanga)
• The general morphology of specimen 99-Z3Eb-01

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Results
Peruvian and Andean engonoceratids
Origin

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References

• Robert, E., & Bulot, L. G. (2004). Origin, phylogeny,

faunal composition, and stratigraphical significance
of the Albian engonoceratidae (pulchelliaceae,
ammonitina) of Peru. Journal of South American
Earth Sciences, 17(1), 11-23.

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