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The shape of the body is short, thick, and very compact. The
composition of the individual segments resembles that found in
Chilopoda rather than that of Chilognatha.

The antennae are very long, longer than in any of the Chilopods, and
are composed of a great number of very small joints. The mouth
parts show a greater length and slenderness than do those of the
other Orders mentioned as yet. They consist of—

1. An upper lip partly free, but fused at the sides with the rest of the head. The
upper lip is in three parts, as in the Chilopoda, but with the middle part very small
and the lateral pieces large.

2. A pair of jaws or mandibles. These are provided not only with teeth, as in the
other Myriapods, but also with a sort of comb of stiff bristles.

3 and 4. Two pairs of maxillae or foot jaws distinguished by their length and
slenderness.

5. The poison claws long, slender, and not sharply curved. The bases of the
poison claws hardly fused together and short.

The respiratory system in the Schizotarsia differs from that in all

other Myriapods in the fact before mentioned, that they breathe by
means of lungs and not by tracheae. There are, as before
mentioned, eight dorsal scales in these animals; each dorsal scale
except the last bears one of the peculiar organs which I have called
lungs. At the hinder end of the scale there is a slit which leads into
an air sac, from which a number of short tubes project into the blood
in the space round the heart and serve to aerate it before it enters
the heart. The heart, therefore, sends aerated blood to the organs,
while in the tracheal-breathing Myriapods the blood is aerated in the
organs themselves by means of tracheae.

The poison claws are followed by segments bearing fifteen pairs of

true ambulatory legs. These are covered by eight large dorsal plates,
increasing in size from before to the middle of the body, the middle
plate being the largest, and then diminishing in size.

The nervous system resembles that of the Chilopoda, but there is a

special pair of nerves which supply the sense organ, which has been
mentioned as peculiar to the Order. The ventral nerve cord shows a
very clear division into two, the ganglia of the two cords being almost
entirely separate. The first few ganglia are fused, as has been
mentioned in the Chilopoda.

The digestive tube resembles that of the Chilopoda. The legs are
very long and slender, and the joints are beset with bristles. Both
sexes have small hook-like appendages at the sides of the genital
openings.

The eyes have already been mentioned as being more highly

developed than in other classes, in correspondence with the more
active habits of the animal. The generative organs open at the hind
end of the body, as in Chilopoda.

The heart is highly developed, quite as much so as the Chilopod

heart, the alary muscles being strong and broad, and the arteries
being quite as perfect as those in any Myriapod. The muscular coats
which govern the pulsations by their contractions are powerful and
well developed.

Order IV. Symphyla.

We next come to one of the last two Orders which have been
recently added to the Myriapoda. These little animals have a great
resemblance to the Thysanura among the Insects, and especially to
Campodea among the Thysanura. It will be well, therefore, to begin
our account with a few of the reasons which have induced naturalists
to include them among the Myriapods rather than among the
Thysanura.

1. Campodea has three pairs of mouth appendages, while Scolopendrella has only
two.

2. Scolopendrella has broad plates covering the back, not only on the anterior
(thoracic) segments, but on the whole body.

3. The terminal appendages of Scolopendrella differ from those in Campodea.

4. Scolopendrella has a sense organ which is absent in Campodea.

5. Campodea breathes by means of three stigmata in the anterior part of the body.
The stigmata of Scolopendrella are hard to see, and are not in the same position.

6. Scolopendrella has twelve pairs of legs, and Campodea, like all Insects, has
only three.

I will now go on to an account of their anatomy. The body is small

and slender, and is covered with a delicate shell or exoskeleton of
chitin, which is so thin as to be almost transparent.

The antennae are long, and are composed of many joints of equal
size.

The mouth parts consist of—

1. An upper lip.
2. A pair of mandibles.
3. A pair of maxillae.

The segments are not all of equal size. Some are larger than others.
The larger and smaller segments are arranged alternately, and the
smaller do not bear legs. As before stated, there are twelve leg-
bearing segments.
At the end of the body there are two hook-like appendages which are
pierced by a canal, through which is poured the secretion of a pair of
glands. Near the sides of these appendages are a pair of sense
organs, consisting of long hairs connected with nerves.

The digestive canal is a long straight tube passing through the length
of the body. In the middle it is much enlarged, so as to form a
stomach with a glandular coat. Posterior to the stomach the digestive
tube receives the contents of two Malpighian tubes which act as
kidneys.

The tracheal system consists of a single pair of stigmata on the

under surface of the head, and the tracheae connected with them.

Order V. Pauropoda.

The Pauropoda, which form the fifth Order of Myriapods, are as yet
very imperfectly known. Pauropus was discovered by Sir John
Lubbock, and its discovery was announced by him in 1866. He found
this little Centipede in his kitchen garden among some Thysanura,
and at first considered it as a larval form, but continued observation
showed that it was a mature creature. He described it as a small,
white, bustling, intelligent little creature about 1⁄25 inch in length.

The antennae are very curious and highly characteristic of the Order.
They resemble those of Crustacea rather than those of Myriapoda.
Each antenna is composed in the following manner. First there is a
shaft of four joints. From the fourth joint of this shaft spring two
branches; one of these two branches is narrower than the other, and
ends in a long thin bristle composed of a great number of joints. The
other and broader branch bears two such bristles, and between them
a small pear-shaped or globular body, the function of which is
unknown.
The mouth parts consist of two minute pairs of appendages, the
anterior pair toothed and the posterior pointed. The body is rather
narrower in front; the segment behind the head has one pair of legs,
the second, third, fourth, and fifth behind the head two each. The
posterior legs are the longest; the genital organs open at the base of
the second pair of legs, between these and the third pair. The
manner of breathing is as yet unknown, tracheae not having been
discovered.

Pauropus at first looks most like a Chilopod, but differs from that
Order—

1. In the form of the antennae.

2. In the absence of poison claws and in the form of the mouth parts.

3. The opening of the generative organs being in the front part of the body.

It differs from Chilognatha in the following respects:—

1. The legs are not of equal length, the posterior legs being the longest, as in
Chilopods.

2. The mouth parts differ from those of Chilognaths almost as much as from those
of Chilopods.

3. The form of the antennae.

Only a few Pauropoda have been discovered as yet.

Embryology.

The preceding account of the anatomy of Myriapods would be

incomplete without some reference to the wonderful manner in which
the different organs of the body are built up; the whole of the
complex organism proceeding by a gradual and regulated process of
development from a simple cell called the ovum derived from the
female body, and united with a cell from the male body (called the
spermatozoon). I hope to be able to give my readers some idea of
the interest which the pursuit of the difficult study of embryology
adds to anatomy, by offering us a key to the interpretation of the
relations between our knowledge of the forms at present living on the
earth and those which, we learn from Palaeontology, have inhabited
our planet in past ages.

Fig. 36.—Young ovum of Julus terrestris: nucl, nucleolus; nu, nucleus;

R, first appearance of yolk; F, follicle cells.

Like all living creatures with which we are acquainted, the starting-
point of Myriapod life is the ovum, as it is called. This ovum is a cell
resembling the cells of which the body of all living animals are built
up, and which may be compared to the bricks of which a building is
composed. This cell or ovum is a small sphere of living transparent
substance called protoplasm, and it is nucleated—that is, it contains
a small spot of denser protoplasm called the nucleus, and within that
a still smaller spot of still more dense protoplasm called the
nucleolus. In the process of impregnation the ovum unites with the
male cell, and the cell so formed is called the impregnated ovum.
This ovum has the property of dividing into two cells, each
resembling the parent cell from which it is derived; each of these
cells has, like the parent cell, the same property of dividing into two
more, and so on. Thus from this continual process of division or
reproduction of every living cell, the materials are provided for the
building up of the body.

The regularity of the process of the division of the ovum, or, as it is

called, segmentation of the ovum, is interfered with by the presence
of food yolk. The cells formed by the process of cell division just
described need nourishment, and this nourishment is supplied to
them by the food yolk formed in the body of the ovum before the
process of segmentation begins. It is easy to understand that this
yolk, which is not alive like the cells, cannot divide like them, and
therefore the segmentation of the ovum in Myriapods is irregular, as
it is called.

Fig. 37.—Later stage: nu, nucleolus; c.p, nucleus; y.sp, yolk spherules;
ch, shell.

I will now go back a little and describe what happens to the ovum
before the process of segmentation is complete. It increases in size
and forms the supply of food yolk which is to provide the nutriment of
the ovum. Then after impregnation the egg-shell is formed round it,
and it becomes what we know as the egg. This egg is not a perfect
sphere, but is oval (in most Myriapods) in shape. The egg is laid, and
the process of segmentation begins shortly after it is laid, as has
already been described.

When it has been laid for about 36 hours, if we take an egg and,
after proper preparation, cut it into thin slices known to microscopists
by the name of sections, and examine it by means of the
microscope, we shall see that segmentation has resulted in this. Just
beneath the egg-shell there is a thin layer of cells, one cell thick,
which completely surrounds the egg. Inside this coat of cells is the
food yolk, with a few cells scattered about in it at rare intervals,
something like the raisins in a plum-pudding.
With the next process the formation of the young Myriapod may be
said to begin. A strip along the length of the oval-shaped egg is
thickened, and this thick mass of cells represents the future ventral
surface of the animal. The rest of the thin layer of cells already
mentioned just below the shell will form the shell or exoskeleton of
the future animal. The thick strip of cells at the ventral surface has by
this time split into layers, so that, resorting to our microscope again,
a section through the short axis of the oval-shaped egg—a
transverse section—will show us—

1. The egg-shell.

2. A layer of cells completely surrounding the egg, thin everywhere but on the
ventral surface. This layer is known to embryologists as the epiblast. The thick part
of the epiblast on the ventral surface gives rise to the nervous system.

3 and 4. Two layers of cells connected in the middle, along the line of the thick
strip, but separate elsewhere, and not extending round the whole of the inside.
These layers constitute what is known as the mesoblast, and give rise to the
muscles and most of the internal organs.

5. The scattered cells in the yolk. They are known as the hypoblast and give rise to
the digestive canal.

After this point is reached the formation of the organs begins. The
segments are formed in order from before backwards. First the head,
then the next segment, and so on. When the number of segments
with which the animal will be hatched are formed, another process
begins, and the tail end of the animal, which can already be
distinguished, is bent towards the head. This is a process that takes
place in many animals besides Myriapods, and is called the
formation of the ventral flexure. Shortly after this the animal bursts
the shell and comes into the outer world. The various processes may
be understood by reference to the Figs. 36, 37, 38, 39, which are
successive stages in the development of a Chilognath. Figs. 37, 38,
are thin slices through the shorter diameter of the egg, which, as
before mentioned, is an oval in shape. Fig. 39 is a section through
the longer diameter of an egg in a more advanced stage of
development, in fact just about to burst the shell. The body of the
future animal is marked by constrictions, the future segments. Some
of the organs are already formed, as the brain and the digestive
tube, the openings of which will form the mouth (st) and the anus
(pr).

Fig. 38.—Transverse section through next stage: mk, keel-like mass of

cells from which the mesoblast is produced; ec, epiblast. (From
Heathcote, Post. Emb. Dev. of Julus terrestris; Phil. Trans. vol.
179, 1888, B.)

Fig. 39.—Longitudinal section through later stage: Segs. 2, 3, etc.,

segments; Ceph. Seg, head; mes, mesoblast; en, hypoblast; st,
future mouth; pr, future anus; mesen, gut; mem.ex, as in Fig. 41.
(From Heathcote, Post. Emb. Dev. of Julus terrestris.)

Myriapods are hatched at different stages of development. The

Chilognatha have only three appendages, which are so little
developed that they are only small shapeless stumps, while the
Chilopoda have the full number of legs in some cases; in others only
a small number of legs, but yet more than the three pairs of legs of
the Chilognatha, and fully developed instead of stump-like. The eyes
are usually developed late in the life of the young animal. The
bursting of the egg-shell is assisted in some Myriapods by a special
kind of spike on the back part of the head.
The Fig. 40 shows a young Chilognath which has just burst the shell
and come into the outer world. It is still surrounded with a membrane
which has been formed by its skin or epiblast within the egg. One
eye-spot has been formed.

Fig. 40.—Young Julus terrestris just hatched.

Fig. 41 shows a longitudinal section through the young Chilognath

shown in Fig. 40, and the next (Fig. 42) a transverse section through
the same. In comparing the two Figs. 41 and 42 it must be
remembered that they are sections in different planes through the
animal shown in Fig. 40, and therefore they only show a small
portion, a thin slice, of the organs.

Fig. 41.—Longitudinal section through late stage: Sup.oe.gl, First

appearance of brain; st, mouth; pr, anus; mesen, gut; n, nerve
cord; n.gang, nerve ganglion; mem.ex, membrane surrounding
the animal; v.f, ventral flexure; mes, mesoblast cells. (Heathcote,
Post. Emb. Dev. of Julus terrestris.)

The first appearance of the mouth appendages has been already

mentioned, and these are shown in Fig. 43, where the small stumps
that later on change to jaws are shown. The figure shows the head
of a young Chilognath seen from the lower side, and the second pair
of stumps fuse together later on and produce the broad plate already
mentioned as the characteristic mouth appendage of the Order.
 Fig. 42.—G, gut; Malp.T, Malpighian tube; N.C, nerve cord; Tr.I, deep
invagination by which the tracheae are formed; y.s, yolk spherules
still present; L, first appearance of legs; S.S, part of mesoblast.
(Heathcote, Post. Emb. Dev. of Julus terrestris.)

Fig. 43.—Under surface of the head of a young Julus terrestris: pro.m,

rudimentary jaws; Deut.m, rudimentary mouth plate; an,
antennae.

After the animal is hatched it has still, in the case of most Myriapods
(those which are not hatched with all the segments complete), to
undergo a further development, and in particular the eyes are still
unformed. The process of development of the eye has only been
followed out as yet in the Chilognatha, and in only one form, Julus,
and is so curious that a short account may be of interest here. The
development of the eye begins (in Julus) on the fourth day after
hatching, and continues until the animal is full grown. A single
ocellus or eye-spot appears first, and the rest are added one by one
until the full number are reached.

The first appearances connected with the formation of the eye take
place in the cellular layer just beneath the chitinous exoskeleton.
This layer, called the hypodermis, plays an important part in the
organisation of the animal. It forms the inner layer of what we may
call the skin of the animal, and the cells of which it is composed
secrete the chitin of which the shell or exoskeleton of the animal is
composed, and which is moulted every year.

The first process in the formation of the eye-spot is the thickening of

the hypodermis beneath the chitin, just in the place where the eye
will come. At the same time the cells of this thickened mass of
hypodermis secrete a quantity of pigment of a dark red brown colour.
Next the cells of the thick mass of hypodermis begin to separate
from one another in such a way that a vesicle is formed. This vesicle
is hollow inside, and the thick walls are formed from the cells of the
thickened hypodermic mass. This can be seen from Fig. 44, which
represents a section through an ocellus when it is partly formed.
From this vesicle the eye is formed.

The wall of the vesicle nearest the exoskeleton gives rise to the lens
of the eye, while the other walls of the vesicle form the retinal parts
of the eye. The cells from the brain grow out and form the optic
nerve connecting the retina with the brain. The whole eye spot is
covered internally by a thin membrane, formed not from the
hypodermis but by cells from the inside of the body (mesoblast
cells).

Fig. 44.—Section through eye when first forming: Hyp, hypodermis; Ln,
lens; F.W.V, front wall of optic vesicle; b.w.v, back wall of vesicle;
cap, capsule.

In the Chilognatha, the first Order of Myriapods, the young animal

leaves the egg with three pairs of appendages; the first have already
the form of antennae, the second will form the jaws, but have not yet
taken their proper form, while the third pair will fuse together and
alter their shape so as to form the curious plate that has already
been mentioned as forming the second pair of mouth appendages.
Behind the mouth appendages will come the first three pairs of legs.
The whole young animal on leaving the egg is enveloped in two
membranes. These membranes are secreted by the outside layer of
cells in the same way that the shell or exoskeleton of the animal will
be eventually formed, and represent the first two moults of the
animal, which continues to moult its shell every year throughout life.

Of the Chilopoda, the second Order of Myriapods, all the families

leave the egg-shell with the full number of legs, with the exception of
the Lithobiidae, which have seven pairs of legs including the poison-
claws. The Schizotarsia, the third Order, also have seven pairs of
legs when hatched.

The legs make their appearance not one by one but in batches (in
Julus terrestris in batches of five). The addition of legs and segments
to the body takes place, not at the end of the body, but between the
end segment and the penultimate.

This is a short sketch of the gradual development of the Myriapoda

from the ovum to the fully-grown animal. It is, I am aware, a short
and insufficient account of all the beautiful processes by which the
different organs take their rise, but space is insufficient here, and too
much detail would be out of place in a work of this nature, which only
aims at giving an outline sketch of the group, which shall be
intelligible to the general reader who has not made a special study of
such matters. Before leaving the subject, however, I must mention a
few of the points of interest which are to be learned from the
examination of the course of development which has been sketched
here. One of the greatest puzzles in the natural history of the Order
Chilognatha has always been the double segments, as they are
called; that is, in fact, the possession of two pairs of legs to each
segment, which is, as we have already said, a distinguishing
characteristic of the Order. As we have seen, the Chilognatha at an
early stage of existence do not possess this characteristic, which is
only peculiar to the adult and half-grown forms. Now what does this
mean? Does each double segment in the full-grown Millepede
represent two segments which have become fused together, or is
each double segment, so called, a real segment resembling the
segments present in the other Orders (for instance, Chilopoda),
which has grown an extra pair of legs? Both these views have been
advocated by distinguished naturalists. Neither of them is, in my
opinion, quite right when viewed in the light cast on the subject by
recent investigations into the life history of the Chilognatha.

A close examination into the minutiae of the growth of the different

organs has shown us that the double characters of the double
segments are more deeply seated than was imagined. The
circulatory system, the nerve cord, and the first traces of
segmentation in the mesoblast all show this double character, and
the only single part about the segment is the broad plate covering
the segment. Now in some of the most ancient of the fossil
Myriapods this broad plate shows traces of a division, as if it were in
reality two plates fused together. We have also to consider that the
life history of the Chilognatha allows us to believe that the peculiar
cylindrical shape of the body shown in the greatest degree in the
Julidae is attained by the unequal development of the dorsal and
ventral surfaces of the body; the ventral surface being compressed
together till it is extremely narrow, and the dorsal surface, as it were,
growing round it till the originally dorsal surface forms almost a
complete ring round the body. Taking all this into consideration, we
are justified, in my opinion, in concluding that each double segment
in the Chilognatha is not two segments fused together, nor a single
segment bearing two pairs of legs, but is two complete segments
perfect in all particulars, but united by a large dorsal plate which was
originally two plates which have been fused together, and which in
most Chilognatha surrounds almost the whole of two segments in
the form of a ring.
Again in the Chilopoda we see that a great distinctive feature that
separates them from the Chilognatha is the character of the ventral
nerve cord, the cord being double and not single, a character
connected with the fact that the bases of the legs are widely
separated from one another, and not closely approached to each
other, as in the Chilognatha. As we before said, a more minute
anatomical examination showed us that this difference was not so
great as appeared at first sight, the cord showing traces of a
duplication. Well, are these traces superficial, or do they represent a
state of affairs more or less similar to that in the Chilopoda?
Embryology helps us to answer this question also. In the early
stages of the Chilognatha we find that the nerve cord has exactly the
form of that in Chilopoda, showing us that the appearances in the
anatomy had led us to a right conclusion, and giving us a valuable
confirmation of our views. These two examples will serve to show
the kind of interest which attaches to embryology.

Palaeontology.

We have seen that embryology enables us to look at the structure of

the Myriapods from a new standpoint, and to correct and supplement
the knowledge gained from an examination of the adult animal. In
the same way a study of the forms of Myriapods which have become
extinct on the globe, and have been preserved to us in a fossil form,
gives a further opportunity of considering the relations of one form to
another, and again of the relations of our group to other groups of
animals now existing on the earth. Myriapod fossils have been found
in strata of great antiquity. The oldest of such fossils must have been
among the first land animals. The figure below shows a fossil
Myriapod found in America, belonging to the Order of the
Protosyngnatha which are only found in the Palaeozoic strata. It is a
good example of the manner in which Myriapods were protected by
bundles of bristles in the same way as the Polyxenus of the present
time.
The oldest fossil Myriapods which have been discovered at the
present time are two species which have been found in the Old Red
Sandstone in Scotland. To realise the antiquity of these Myriapods, it
will be worth while recalling the typical fossils found in the Old Red
Sandstone, so as to see what the contemporaries of these ancient
Myriapods were like. Among the plants there were Algae, Ferns, and
Conifers, belonging to the lower divisions of the plant tribe. Among
the animals there were Sponges, Corals, Starfish, Worms, Shell-fish,
and Fishes, but none of the more highly organised of the animal or
vegetable tribe had appeared on the earth. The Myriapods of the Old
Red Sandstone, as has been before said, differ considerably from
those of the present day, and as we proceed towards the species
found in the more recent strata we find them more and more like the
ones at present living, till we get to the Polyxenus and other species
found in amber, which are hardly to be distinguished from living
forms.

The next oldest fossil Myriapods are found in the coal measures,
when both the animal and vegetable kingdoms were represented by
more numerous and more specialised forms. The fossil fauna of this
period is characterised by the number of gigantic Amphibia, many
remains of which have been found. The great forests and the
abundant vegetation of this time must have been favourable to the
existence of our class, and accordingly we find no less than 32
species of fossil Myriapods. Of these most have been found in
America, some in Great Britain, and some in Germany. One well-
preserved fossil of Xylobius sigillariae was found by Dr. Dawson in
America in the stump of a tree in the remains of a fossil forest. The
eyes, head, and legs were plainly seen under the microscope. All
these fossils belong to the earliest or Palaeozoic period.
 Fig. 45.—Palaeocampa anthrax. (After Meek and Worth.) From Mazon
Creek, Illinois.

The figure below (Fig. 46) shows a fossil also from the coal
formations of Illinois, America, belonging to the family of the
Euphoberiidae mentioned further on. It shows a nearer approach to
the Julidae of the present time. The limbs, however, were of very
curious shape, and may possibly have been adapted to locomotion
in water as well as on land, and the small supposed branchiae on
the ventral surface shown in Fig. 46, B, may possibly have been an
arrangement to render respiration in the water possible.

In the secondary period the Myriapods were scantily represented, or,

at any rate, geologists have failed to find their fossils. The class is
represented by a single specimen found in the chalk in Greenland.
This fossil, which has been included in the Julidae under the name of
Julopsis cretacea, may perhaps belong to the Archipolypoda.

Passing on to the Tertiary or Recent period, we find the Myriapods

again numerous, and more nearly resembling those living at the
present time. They belong mostly to the Chilognatha and Chilopoda.
They have been found in the fresh-water gypsum of Provence in
France, the brown coal of Germany, and the green river formations
of America. Several have been found in amber.

Fig. 46.—Acantherpestes major. (After Meek and Worth.) Mazon

Creek, America. A, The whole animal; B, branchiae on the ventral
surface.

Fossil Myriapods have been divided into four Orders, two of which
coincide with the Orders of living Myriapods; the differences between
the fossils and the living Myriapods having been held insufficient to
warrant the establishment of a new Order. These two Orders are the
Chilopoda and the Diplopoda or Chilognatha (Diplopoda is another
name used by some writers for the group which we have hitherto
called Chilognatha). The other two Orders have sufficient differences
from living forms to render it necessary to include them in separate
Orders.

The fossil Myriapods, then, are arranged as follows:—

Order I. Protosyngnatha.
Order II. Chilopoda.
Order III. Archipolypoda.
Order IV. Chilognatha (or Diplopoda).

The following table will show the species that have been discovered
in the different strata:—

Devonian, or
2 species of Archipolypoda
Old Red Sandstone
1 species Protosyngnatha
Carboniferous
31 species Archipolypoda
Permian (Rothliegendes of Germany), 4 specimens belonging to
the
Julidae or Archipolypoda.
Archipolypoda or
Cretaceous 1 species
Chilognatha
17 species Chilopoda
Oligocene Diplopoda
23 species
(Chilognatha)

Diplopoda
Miocene, 1 species
(Chilognatha)
I will now give a short account of the different Orders, and the fossil
forms which are included in them.

Order I. Protosyngnatha.

This Order is represented by a single fossil (Fig. 45), discovered in

the coal at Mazon Creek, Illinois, America, by Meek and Worth. It
differs greatly from any of those in existence at the present day. The
body is cylindrical, and composed of ten segments. The cephalic
appendages (that is, the antennae and mouth parts) are inserted into
a single unsegmented cephalic mass (the head). Each segment
behind the head bears a single dorsal and ventral plate of equal
breadth and length. The limbs are placed in these plates with a wide
space between the base of each leg and that of the opposite one of
the pair. Along the back, bundles of bristles are arranged in
longitudinal rows.

Order II. Chilopoda.

The fossil forms of this Order resemble those of the Chilopoda of the
present day. The oldest of them are found in amber. The following
families have been found:—

Lithobiidae. Several species have been found in amber.

Scolopendridae. One species in amber, several species in later Tertiary

formations.

Geophilidae. Three species in amber.

Two species resembling the Schizotarsia of the present day have

been found in amber.
 Order III. Archipolypoda.

The most numerous of the fossil families. With a few exceptions, all
the Palaeozoic (that is, the oldest) Myriapods belong to this Order.
The Carboniferous Archipolypoda seem to be much more numerous
in the coal of America than in that of England. They resemble for the
most part the Myriapods of the present day, except that all the
segments without exception bear legs.

The families are three in number.

Family 1. Archidesmidae.

Resemble the Polydesmidae of the present day. Two species have been found by
Page in the Old Red Sandstone of Forfarshire. He named them Kampecaris. One
found by Peach in the same formation is called Archidesmus.

Family 2. Euphoberiidae.

They show some resemblance to the Julidae of the present day, but the dorsal
scutes, or plates of the back, are more or less perfectly divided into two divisions
corresponding with the pairs of legs. The following are the principal fossils of this
family:—

Acantherpestes. Found by Meek and Worth in the coal at Mazon Creek in

America (Fig. 46).

Euphoberia. About 12 species found at the same place as the last named.

Amylispes. Found by Scudder, Mazon Creek, America.

Eileticus. Scudder, Mazon Creek, America.

Family 3. Archijulidae.

The dorsal plates nearly consolidated, but the division still apparent. Fossil forms
are—